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1.
Timothy P. Hanrahan   《Geomorphology》2007,86(3-4):529-536
While the importance of river channel morphology to salmon spawning habitat is increasingly recognized, quantitative measures of the relationships between channel morphology and habitat use are lacking. Such quantitative measures are necessary as management and regulatory agencies within the Pacific Northwest region of the USA, and elsewhere, seek to quantify potential spawning habitat and develop recovery goals for declining salmon populations. The objective of this study was to determine if fall Chinook salmon (Oncorhynchus tshawytscha) spawning areas in the Snake River, Idaho, USA, were correlated with specific bedform types at the pool–riffle scale. A bedform differencing technique was used to objectively quantify the longitudinal riverbed profile into four distinct pool–riffle units that were independent of discharge. The vertical location of thalweg points within these units was quantified with a riffle proximity index. Chinook salmon spawning areas were mapped and correlated with the pool–riffle units through the use of cross-tabulation tables. The results indicate that 84% of fall Chinook salmon spawning areas were correlated with riffles (χ2 = 57.5, df = 3, p < 0.001), with 53% of those areas located on the upstream side of riffle crests. The majority of Snake River fall Chinook salmon spawning occurred at elevations greater than 80% of the difference in elevation between the nearest riffle crest and pool bottom. The analyses of bedform morphology will assist regional fish managers in quantifying existing and potential fall Chinook salmon spawning habitat, and will provide a quantitative framework for evaluating general ecological implications of channel morphology in large gravel-bed rivers.  相似文献   

2.
Travertine deposits of calcium carbonate can dominate channel geomorphology in streams where travertine deposition creates a distinct morphology characterized by travertine terraces, steep waterfalls, and large pools. Algae and microorganisms can facilitate travertine deposition, but how travertine affects material and energy flow in stream ecosystems is less well understood. Nearly a century of flow diversion for hydropower production has decimated the natural travertine formations in Fossil Creek, Arizona. The dam will be decommissioned in 2005. Returning carbonate-rich spring water to the natural stream channel should promote travertine deposition. How will the recovery of travertine affect the ecology of the creek? To address this question, we compared primary production, decomposition, and the abundance and diversity of invertebrates and fish in travertine and riffle/run reaches of Fossil Creek, Arizona. We found that travertine supports higher primary productivity, faster rates of leaf litter decomposition, and higher species richness of the native invertebrate assemblage. Observations from snorkeling in the stream indicate that fish density is also higher in the travertine reach. We postulate that restoring travertine to Fossil Creek will increase stream productivity, rates of litter processing, and energy flow up the food web. Higher aquatic productivity could fundamentally shift the nature of the stream from a sink to a source of energy for the surrounding terrestrial landscape.  相似文献   

3.
The Lamar River watershed of northeastern Yellowstone contains some of the most diverse and important habitat in the national park. Broad glacial valley floors feature grassland winter range for ungulates, riparian vegetation that provides food and cover for a variety of species, and alluvial channels that are requisite habitat for native fish. Rapid Neogene uplift and Quaternary climatic change have created a dynamic modern environment in which catastrophic processes exert a major influence on riverine–riparian ecosystems. Uplift and glacial erosion have generated high local relief and extensive cliffs of friable volcaniclastic bedrock. As a result, steep tributary basins produce voluminous runoff and sediment during intense precipitation and rapid snowmelt. Recent major floods on trunk streams deposited extensive overbank gravels that replaced loamy soils on flood plains and allowed conifers to colonize valley-floor meadows. Tree-ring dating identifies major floods in 1918, ca. 1873, and possibly ca. 1790. In 1996 and 1997, discharge during snowmelt runoff on Soda Butte Creek approached the 100-year flood estimated by regional techniques, with substantial local bank erosion and channel widening. Indirect estimates show that peak discharges in 1918 were approximately three times greater than in 1996, with similar duration and much greater flood plain impact. Nonetheless, 1918 peak discharge reconstructions fall well within the range of maximum recorded discharges in relation to basin area in the upper Yellowstone region. The 1873 and 1918 floods produced lasting impacts on the channel form and flood plain of Soda Butte Creek. Channels may still be locally enlarged from flood erosion, and net downcutting has occurred in some reaches, leaving the pre-1790 flood plain abandoned as a terrace. Gravelly overbank deposits raise flood-plain surfaces above levels of frequent inundation and are well drained, therefore flood-plain soils are drier. Noncohesive gravels also reduce bank stability and may have persistent effects on channel form. Overall, floods are part of a suite of catastrophic geomorphic processes that exert a very strong influence on landscape patterns and valley-floor ecosystems in northeastern Yellowstone.  相似文献   

4.
The hydraulic and sedimentary characteristics of the spawning habitat of Atlantic salmon (Salmo salar) in tributary and mainstem locations in a river system in north-east Scotland are described. Salmon used spawning sites with a relatively wide range in sediment characteristics, although measures of central tendency were all in the gravel (2–64 mm) size-class. The dominant factor differentiating the sediment characteristics of study sites was the level of fine sediment, which accounted for significant differences between tributary and mainstem samples. The ranges of depth and velocity in areas used for spawning by salmonids were found to be similar in all tributary study sites. However, due to the interdependence of depth and velocity, major differences were observed between tributary and mainstem study sites in that spawning in larger streams tended to be associated with deeper, faster flowing water. Spawning locations were shown to have similar Froude number, despite different sized streams and species of salmonid. Due to its dimensionless nature and significance in characterising flow hydraulics, the Froude number is proposed as a potentially useful variable for describing the habitat of aquatic organisms.  相似文献   

5.
Sequential aerial photography, sonar bathymetry, ground-penetrating radar (GPR), and sediment sampling and analysis provide the basis for calculating the volumetric and mass rate of progradation of the delta of Fitzsimmons Creek, a steep, high-energy, debris-flow-dominated channel draining about 100 km2 of the southern Coast Mountains of British Columbia. Fitzsimmons Creek is typical of small mountain rivers in the region. GPR imaging is used to define the pre-depositional morphology of the receiving basin, a technique that improves the accuracy of the volumetric survey. The 52-year record (1947–1999) of progradation yielded an average annual volumetric transport rate of 1.00±0.16×104 m3 year−1 for bed load, corresponding to a mass transport rate of 1.60±0.28×104 Mg year−1. Bed load yields are consistent with those obtained in hydrogeomorphically similar basins in the region and elsewhere. Decade-based annual rates, which vary from 0.64±0.11×104 to 2.85±0.38×104 Mg year−1, provide poor estimates of the 52-year average. Indeed, the 52-year record may also not be long enough to fully integrate the significant fluctuations in the sediment efflux from Fitzsimmons Creek. The methodology proposed in this paper can be transferred to other comparable mountain environments worldwide.  相似文献   

6.
At least one large, late Pleistocene flood traveled into the Owyhee River as a result of a rise and subsequent outburst from pluvial Lake Alvord in southeastern Oregon. Lake Alvord breached Big Sand Gap in its eastern rim after reaching an elevation of 1292 m, releasing 11.3 km3 of water into the adjacent Coyote Basin as it eroded the Big Sand Gap outlet channel to an elevation of about 1280 m. The outflow filled and then spilled out of Coyote Basin through two outlets at 1278 m and into Crooked Creek drainage, ultimately flowing into the Owyhee and Snake Rivers. Along Crooked Creek, the resulting flood eroded canyons, stripped bedrock surfaces, and deposited numerous boulder bars containing imbricated clasts up to 4.1 m in diameter, some of which are located over 30 m above the present-day channel.Critical depth calculations at Big Sand Gap show that maximum outflow from a 1292- to 1280-m drop in Lake Alvord was  10,000 m3 s− 1. Flooding became confined to a single channel approximately 40 km downstream of Big Sand Gap, where step-backwater calculations show that a much larger peak discharge of 40,000 m3 s− 1 is required to match the highest geologic evidence of the flood in this channel. This inconsistency can be explained by (1) a single 10,000 m3 s− 1 flood that caused at least 13 m of vertical incision in the channel (hence enlarging the channel cross-section); (2) multiple floods of 10,000 m3 s− 1 or less, each producing some incision of the channel; or (3) an earlier flood of 40,000 m3 s− 1 creating the highest flood deposits and crossed drainage divides observed along Crooked Creek drainage, followed by a later 10,000 m3 s− 1 flood associated with the most recent shorelines in Alvord and Coyote Basins.Well-developed shorelines of Lake Alvord at 1280 m and in Coyote Basin at 1278 m suggest that after the initial flood, postflood overflow persisted for an extended period, connecting Alvord and Coyote Basins with the Owyhee River of the Columbia River drainage. Surficial weathering characteristics and planktonic freshwater diatoms in Lake Alvord sediment stratigraphically below Mt. St. Helens set Sg tephra, suggest deep open-basin conditions at  13–14 ka (14C yr) and that the flood and prominent shorelines date to about this time. But geomorphic and sedimentological evidence also show that Alvord and Coyote Basins held older, higher-elevation lakes that may have released earlier floods down Crooked Creek.  相似文献   

7.
Debris flows are widespread and common in many steeply sloping areas of southern California. The San Bernardino Mountains community of Forest Falls is probably subject to the most frequently documented debris flows in southern California. Debris flows at Forest Falls are generated during short-duration high-intensity rains that mobilize surface material. Except for debris flows on two consecutive days in November 1965, all the documented historic debris flows have occurred during high-intensity summer rainfall, locally referred to as ‘monsoon’ or ‘cloudburst’ rains. Velocities of the moving debris range from about 5 km/h to about 90 km/h. Velocity of a moving flow appears to be essentially a function of the water content of the flow. Low velocity debris flows are characterized by steep snouts that, when stopped, have only small amounts of water draining from the flow. In marked contrast are high-velocity debris flows whose deposits more resemble fluvial deposits. In the Forest Falls area two adjacent drainage basins, Snow Creek and Rattlesnake Creek, have considerably different histories of debris flows. Snow Creek basin, with an area about three times as large as Rattlesnake Creek basin, has a well developed debris flow channel with broad levees. Most of the debris flows in Snow Creek have greater water content and attain higher velocities than those of Rattlesnake Creek. Most debris flows are in relative equilibrium with the geometry of the channel morphology. Exceptionally high-velocity flows, however, overshoot the channel walls at particularly tight channel curves. After overshooting the channel, the flows degrade the adjacent levee surface and remove trees and structures in the immediate path, before spreading out with decreasing velocity. As the velocity decreases the clasts in the debris flows pulverize the up-slope side of the trees and often imbed clasts in them. Debris flows in Rattlesnake Creek are relatively slow moving and commonly stop in the channel. After the channel is blocked, subsequent debris flows cut a new channel upstream from the blockage that results in the deposition of new debris-flow deposits on the lower part of the fan. Shifting the location of debris flows on the Rattlesnake Creek fan tends to prevent trees from becoming mature. Dense growths of conifer seedlings sprout in the spring on the late summer debris flow deposits. This repeated process results in stands of even-aged trees whose age records the age of the debris flows.  相似文献   

8.
Ellen Wohl  David Dust 《Geomorphology》2012,138(1):329-338
Since 1974, flow releases from Long Draw Reservoir have increased annual peak flows on La Poudre Pass Creek, Colorado, from ~ 5.6 m3/s to > 8.4 m3/s. The creek drains 61 km2 and channel morphology varies from step-pool to pool-riffle. Comparison of five channel reaches along the creek to channel reaches along neighboring rivers without flow regulation indicates that channel width has increased by as much as a factor of three along La Poudre Pass Creek. Width-to-depth ratio has also increased, the bed material in step-pool channel reaches has coarsened, and residual pool volumes have increased in pool-riffle channel reaches. Pool-riffle channel reaches have undergone the greatest change in response to flow augmentation. Although discharge has increased consistently for all five channel reaches, morphologic response varies in relation to gradient and channel morphology, making it impractical to precisely predict a priori the magnitude of channel response to flow augmentation.  相似文献   

9.
《Geomorphology》2002,42(3-4):255-278
The Hunter Mountain fault zone strikes northwesterly, is right-lateral strike-slip, and kinematically links the northern Panamint Valley fault zone to the southern Saline Valley fault zone. The most recent displacement of the fault is recorded in the offset of Holocene deposits along the entire length of the fault zone. Right-lateral offsets of drainage channels within Grapevine Canyon reach up to 50 to 60 m. Initial incision of the offset channels is interpreted on the basis of geomorphic and climatic considerations to have occurred approximately 15 ka. The 50 to 60 m of offset during 15 ka corresponds to a right-lateral fault slip rate of 3.3–4.0 mm/year within Grapevine Canyon. Further to the north along the Nelson Range front, the fault is composed of two sub-parallel fault strands and the fault begins to show an increased normal component of motion. A channel margin that is incised into a Holocene surface that is between 10 and 128 ka in age is offset 16–20 m, which yields a broad minimum bound on the lateral slip rate of 0.125–2.0 mm/year. The best preserved single-event displacements recorded in Holocene deposits range from 1.5 to 2.5 m. In addition to faulting within Grapevine Canyon and the main rangefront fault along the southwest edge of Saline Valley, there also exist normal fault strands within the Valley that strike northeasterly and towards Eureka Valley. The northeasterly striking normal faults in the Valley appear to be actively transferring dextral slip from the Hunter Mountain fault zone north and east onto the Furnace Creek fault zone. Separations on northerly trending, normal faults within Saline Valley yield estimates of slip rates in the hundredths of millimeters per year.  相似文献   

10.
This paper presents a hydrodynamic-based investigation of constructed wetland suitability for fish habitat. A constructed wetland adjacent to river is manoeuvred to imitate riparian fish habitat by means of computer modelling. Flow-velocity conditions, water depths and effects of macrophytes to resistance of flow in constructed wetland are modelled and steered towards creating a favourable ground for natural substrate spawning of native fish species. The model suggests combination of two zones, which are a macrophyte zone for typical functions of a wetland and an open water zone with gravel bed to support fish reproductive cycles.  相似文献   

11.
Coarse woody debris affects many streams in forested regions throughout the world. These effects include lateral channel migration, bank slumping, and aggradational or erosional features related to flow redirection. The extent of effect is dependent on the ability of the system to resist the new forces derived from flow redirection. This study on the Pine River, Ontario, looks at how obstructions that are perpendicular to downstream flow modify fluid behavior. Results show that fluctuations in speed and approach azimuth vary considerably depending on the position of the sample relative to the obstruction. The use of time-averaged (1 sec., 30 sec.) recordings of fluid speed and azimuth at selected channel locations shows how flow adjusts to external controls as it moves away from the obstruction zone, giving an indication of the spatial extent of the obstruction influence. These data are represented as a function of the diameter of the obstruction relative to the surrounding flow depth (obstruction ratio), and then are compared to results found in other debris obstructions on the Pine River and Wilmot Creek. Flow obstruction dimensions in the study site equal 26.5 trunk diameters (the average diameter of the tree trunk measured five times along its length), and range between 16.9 and 56.7 trunk diameters on the Pine River (n = 48) and between 7.4 and 63.5 trunk diameters on Wilmot Creek (n = 1066). Knowledge of these spatial relationships may allow for better management of woody debris in streams, primarily from the perspective of aquatic habitat. Multiquadric interpolation formed the basis for plotting fluid vector fields, showing the behavior of flow as it approached and moved through the obstruction zone. This is compared to studies of flow in unobstructed meanders in an attempt to quantify obstruction influence, and is used to provide a depiction of flow under these circumstances. [Key words: flow patterns, woody debris, influence zones, stream management.]  相似文献   

12.
Extensive valley fills have formed at the base of the escarpment in granitic catchments along the south coast of NSW. On the 1865 portion plan, the valley fill surface in the upper part of Wolumla Creek, in the Bega River catchment, was intact, but within a few decades of European settlement of the area the valley fill had been incised. Today the incised channel is up to 10 m deep and 100 m wide. The catchment drains an area of just 18.2 km2. Based on detailed field mapping, with extensive drilling and angering, the volume of the intact valley fill in upper Wolumla Creek in 1865 was approximately 5000 × 103 m3. Between 1865 and the present day, approximately 3500 × 103 m3 of this material has been removed, leaving roughly 1500× 103 m3 of material stored on the valley margins. During an initial period of discontinuous gullying, approximately 230 × 103 m3 of sand accumulated as a floodout. Subsequently, the incised channel became continuous, cutting through the floodout; over 50 per cent of floodout deposits were removed. Flushing of the materials released from upland valley fills has been very efficient in the Wolumla Creek catchment, with a sediment delivery ratio of around 70 per cent. The efficient downstream transfer of deposits reflects bedrock confinement in downstream reaches. Extensive volumes of material have accumulated along the lower reaches of the catchment, exacerbating the transformation to the geomorphic character of the lower Bega River.  相似文献   

13.
The presented paper analyses the variability of grain size distribution parameters of bedload transported by the gravel‐bed Scott River (Svalbard) draining a glacier catchment with an area of 10 km2. The grain size distribution analysis is one of the basic elements of identification of the fluvial transport mechanisms in gravel‐bed rivers. It is used for the determination of threshold values for bedload movement. It is also treated as an important indicator of the origin, routes of distribution, and conditions of transport and deposition of fluvial bedload. The field study in a natural proglacial gravel‐bed channel was carried out at two reaches in the mouth section of Scott River. The study revealed relatively high temporal variability and similar mean parameters of grain size distribution in conditions of low discharges. Bedload transport rates reached a mean of 71.9–76.0 kg d?1 in channel cross‐section. Bedload texture was dominated by gravels with a proportional contribution of the fine‐grained fraction along with very fine‐grained gravels (8‐2 mm) of 38.8%. The medium‐grained fraction (16‐8 mm) constituted 33.7%, with a lower contribution from the coarse‐grained fraction (32‐16 mm) of 23.2%, and the very coarse‐grained fraction (64‐32 mm) of 4.4%. Two periods in the course of bedload transport and distribution of grain size distribution parameters were distinguished based on variation of hydro‐meteorological conditions. The first half of the measurement period was distinguished by significantly higher values of daily loads and increased contribution of the coarse‐grained and very coarse‐grained fraction (28–31% and 6.2–6.6%, respectively). During this time, the river discharged up to 94% of bedload. This resulted in a clear tendency for riverbed scouring. The second half was distinguished by generally low daily bedload transport rates (<10 kg d?1), an increase in contribution of fine‐ and very fine‐grained gravels (42–55.6%), and a change in the tendency to aggradation. Grain size indices were more varied, and grains were usually finer and better sorted. Selective transport processes, often related to redeposition, were dominant in the channel. Along with an increase in flow velocity, conditions for material deposition became more variable. This was manifested in weaker sorting and an increase in grain diameter.  相似文献   

14.
At present, the Ramu River in Papua New Guinea flows directly into the Bismarck Sea, while the adjacent Keram flows into the Sepik River. There is evidence to suggest that the Ramu previously occupied the existing Keram water course, and has been gradually diverting its discharge into its present channel, probably over the last 4,000 years. The diversion process is not complete yet, and during the wet season flood water still travels across these two rivers. The most direct evidence to support this proposal of river diversion is that there are much larger meander bends on which the existing Keram stream channel is superimposed. The diversion of the Ramu is believed to be a result of relative sea‐level rise in the late Quaternary which altered the gradients of the Keram and Ramu rivers, and may have been facilitated through neo‐tectonic movement of the floodplain.  相似文献   

15.
Ronald B. Zelt  Ellen E. Wohl   《Geomorphology》2004,57(3-4):217-233
Large variability in responses of stream sediment and large woody debris (LWD) to severe fire has limited the accurate prediction of the magnitude and duration of fire effects on streams. Conditions in one Absaroka Range stream that was severely burned in 1988 were compared to those in an adjacent, undisturbed stream to improve understanding of fire effects on channel and LWD characteristics beyond the first few years. Ten reaches of each stream were sampled during summer 1999.Average bankfull channel width was greater in burned Jones Creek than in unburned Crow Creek. LWD frequency and overall frequency of LWD accumulations were greater in Crow Creek than Jones Creek. Debris-jam frequency was greater in Jones Creek after accounting for differences in the frequency of pieces with length greater than channel width. Larger piece size and better anchoring contributed to more frequent, small accumulations of LWD in Crow Creek. Differences between streams in LWD frequency are consistent with greater mobility of debris in burned Jones Creek. LWD-associated fine-sediment deposits were thicker but less frequent along Jones Creek than Crow Creek.  相似文献   

16.
This research evaluates the impact of rural-to-urban land use conversion on channel morphology and riparian vegetation for three streams in the Central Redbed Plains geomorphic province (central Great Plains ecoregion) of Oklahoma. The Deep Fork Creek watershed is largely urbanized; the Skeleton Creek watershed is largely rural; and the Stillwater Creek watershed is experiencing a rapid transition from rural to urban land cover. Each channel was divided into reaches based on tributary junctions, sinuosity, and slope. Field surveys were conducted at transects in a total of 90 reaches, including measurements of channel units, channel cross-section at bankfull stage, and riparian vegetation. Historical aerial photographs were available for only Stillwater Creek watershed, which were used to document land cover in this watershed, especially changes in the extent of urban areas (impervious cover).The three streams have very low gradients (< 0.001), width-to-depth ratios < 10, and cohesive channel banks, but have incised into red Permian shales and sandstone. The riparian vegetation is dominated by cottonwoods, ash, and elm trees that provide a dense root mat on stream banks where the riparian vegetation is intact. Channels increased in width and depth in the downstream direction as is normally expected, but the substrate materials and channel units remained unchanged. Statistical analyses demonstrated that urbanization did not explain spatial patterns of changes in any variables. These three channels in the central Redbed Plains are responding as flumes during peak flows, funneling runoff and the wash-load sediment downstream in major runoff events without any effect on channel dimensions. Therefore, local geological conditions (similar bedrock, cohesive substrates and similar riparian vegetation) are mitigating the effects of urbanization.  相似文献   

17.
During February 1992, a series of relatively warm storms passed eastward across southern California, yielding intense precipitation that triggered widespread mass movement, flooding, property damage, and loss of life. These storms were triggered by an intense low pressure system (976 mb) off northern California which deepened as its eastward progress was initially blocked by a high pressure ridge (1040 mb) across western North America. Between February 10 and 13, large areas of Ventura and Los Angeles counties experienced cumulative precipitation of 200–400 mm with intensities reaching 40–50 mm hr-1. Mass movement, mainly as soil slips that transformed downslope into debris flows, occurred where cumulative precipitation exceeded 300 mm and when sustained intensities exceeded 25 mm hr-1. Stream response was rapid, particularly in urban areas where impermeable surfaces and storm drains fed concrete stream channels. The canalized upper Los Angeles River and Arroyo Simi exceeded all previous discharges for over 43 and 36 years of record, respectively. Other streams, from the large Santa Clara River to modest Malibu Creek, yielded recurrence intervals for the peak discharge of between 8 and 24 years, but the rapidity of flooding everywhere was remarkable. Whereas main trunk streams, canalized or not, responded predictably, the storm series emphasized the problems of poorly controlled development of potentially unstable hillsides and floodable lowlands and indicated a need to reassess the assumptions upon which such development is permitted. [Key words: climatology, cyclonic storm, geomorphology, mass movement, flooding, California].  相似文献   

18.
长江历来都是人们所关注的重要水资源之一。它的丰沛的水量除了满足本流域各种用水之外,能否从其中提供一部分水量向我国北方地区跨流域供水,这是探讨南水北调可能性的关键问题之一。  相似文献   

19.
《自然地理学》2013,34(4):269-290
Historical information on stream stage/discharge relations from eight U.S. Geological Survey (USGS) streamflow-gaging stations was used to analyze channel-bed elevation change along Soldier Creek, a stream affected by multiple disturbances in northeast Kansas. The analysis provided information on the spatial (location, type, magnitude) and temporal (timing, duration, trend, rate) dimensions of channel change. Channel changes determined for Soldier Creek included extensive changes resulting from channelization and changes of relatively limited extent following a flood. The results were used to document channel changes, partly reconstruct historical channel conditions, infer the causes of channel change, and estimate the occurrence of future channel changes.  相似文献   

20.
Artificial water sources have been used for decades to enhance and restore wildlife habitat but the benefits of their use have been subject to debate. During the past century, the number of natural springs in Joshua Tree National Park, California, USA, has declined. In response to concerns about the viability of the bighorn sheep (Ovis canadensis nelsoni) population, a number of water developments were constructed throughout the park. We modeled potential historical and present-day summer habitat of female bighorn sheep to evaluate the effectiveness of the artificial and remaining natural water sources in maintaining habitat and to determine how loss of artificial sources might affect future habitat availability. Prior to 1950, 583.5 km2 of summer habitat was potentially available. Presently, only 170.6 km2 of habitat is available around natural water sources and 153.5 km2 is available around guzzlers. When all perennial water sources are included in the habitat model (minus overlap), 302.3 km2 of summer habitat is potentially available. This represents only 51.7% of summer habitat available prior to 1950. Without artificial water developments, 47.7% of present-day summer habitat would be lost, which raises important management questions regarding the debate about what is natural or artificial within otherwise protected areas.  相似文献   

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