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1.
最早的栉水母动物化石——华南伊迪
卡拉纪的“八臂仙母虫” 总被引：2，自引：1，他引：1
华南伊迪卡拉系陡山沱组保存的磷酸盐化球状化石中有一部分被认为是最早的后生动物胚胎化石, 但由于缺乏成年或幼年过渡期的化石证据, 对这些最早微化石的动物属性还一直存在争议。最近在贵州江口县翁会地区伊迪卡拉(震旦)系陡山沱组的页岩中发现一个与“庙河生物群”相似的宏体碳质压膜化石群, 其中包含一个盘状的宏体化石新类型——八臂仙母虫(Eoandromeda octobrachiata, Tang et al. 2008), 具有八条螺旋状辐射的旋臂, 封闭在一个包膜内。这种八辐射结构在八射珊瑚和栉水母动物中都可见到, 但同时具有的螺旋特性与现生的所有类群都无法对比。在贵州陡山沱组的“瓮安生物群”中发现的球状胚胎化石同样具有螺旋性状, 尽管还没有直接的个体发育证据, 但我们认为新属种可能代表这些螺旋状胚胎化石的成年实体化石。最新报道的南澳大利亚Rawnsley石英岩层中也发现相似的印痕化石, 被归为同一类别, 表明华南的“庙河生物群”与澳大利亚的“伊迪卡拉生物群”时代相当。翁会化石库中少量出露的三辐射化石与俄罗斯“白海生物群”的典型分子—Albumares和Anfesta可以对比。新发现的倾斜保存的八臂仙母虫化石和碳质条带呈子午线状排列的核桃状化石, 可能是该类化石归属栉水母类的有力证据, 这一发现推测可将栉水母类的最早化石记录从“澄江生物群”的早寒武世推前大约3千万年。 相似文献
2.
独特的八辐射动物化石——八臂仙母虫(Eoandromeda octobrachiata)新记录被推断为原始的栉水母动物,在华南震旦系陡山沱组和南澳大利亚伊迪卡拉生物群产地均有发现,不仅为相关地层的国际对比架起了桥梁,而且将栉水母动物的最早化石记录推前了大约30Ma,并可能重塑动物系统树,原始的性状特征(如:栉板、栉板带、管状辐射臂、中央环带和八辐射螺旋对称性等)可以把栉水母动物定位在动物系统树更为原始的位置上,使我国后生动物起源和早期演化研究再次获得突破性进展,为破解早期动物演化之谜提供了化石方面的新证据,也与分子生物学研究的最新成果基本吻合。 相似文献
3.
最近,在世界上开始发现了有栉动物(栉水母类)化石(G·D·stanley,Jr,和W·sturmer,1983)据发现,化石在西德产出于早期泥盆纪的粘板岩中.在这种岩石中迄今已发现了将近370种无脊椎动物化石,这个栉水母也和那些化石一样.它的有机硫黄成分为黄铁矿所置换,因此软体部分形态被保存下来,这已通过X射线法查出.此栉水母化石现已命名为Paleoctenopborarasseli,呈放射对称的卵形,体长13mm,直径 相似文献
4.
作为观察寒武纪大爆发主幕奥秘的最佳科学窗口,澄江化石库展示了三胚层动物(即两侧对称动物)首次全面大辐射的历史产物,让学术界终于看到了已知最古老的同时包括原口类和后口类两大支系的动物谱系庞大“树冠”的主体轮廓,从而给许多较高级动物类群的起源和早期演化研究提供了十分重要的信息。然而,该化石宝库中却很少见到作为“树干”的双胚层动物(即辐射动物)的化石记录,这给“真动物”的源头探秘蒙上了阴影。尽管广布于前寒武纪末期全球浅海域的“文德生物群”一直被认为是动物界早期一次“失败的演化试验”的牺牲品,但其中一支成功延续到早寒武世的春光虫(Stromatoveris)具有叠覆状梳齿构造,很可能代表着栉水母类的一类原始祖先。文中描述新的羽毛状动物王氏澄江海笔(Chengjiangopennawangii)不同于春光虫,前者具有彼此分离的羽枝,且羽枝上保存了排列规整的珊瑚个体虫室,因而与现代八射珊瑚中海笔类的形态学特征十分一致;新的谱系分析显示,它应该代表刺胞动物门中的一个原始类群。 相似文献
5.
埃迪卡拉纪陡山沱期瓮安生物群中磷酸盐化
球状化石新材料及其问题讨论 总被引：2，自引：0，他引：2
磷酸盐化球状化石是埃迪卡拉纪瓮安生物群中的主要化石类型。研究初期其生物属性被归于绿藻; 随后, 由于发现一些球状化石标本保存有类似后生动物卵裂的二分、四分、八分、十六分、三十二分等细胞分裂特征及分裂球规则有序的排列方式, 并且不同分裂阶段的球体大小基本保持一致等特征, 一些学者将其解释为动物的胚胎化石, 并由此引起了国内外众多学者的关注和热烈的讨论。近年来, 又有学者将其解释为巨大的氧化硫细菌。本文在系统介绍这些磷酸盐化球状化石研究现状的基础上, 报道了一些新的球状化 石材料, 包括一些具有出芽现象的球状化石以及Parapandorina和Megaclonophycus之间的过渡分子。新材料的发现显示, 尽管休眠卵和胚胎假说是迄今为止对瓮安生物群中磷酸盐化球状化石的最佳解释, 但仍 有无法解释的一些问题, 这些球状化石确切亲缘关系的确定还有待于新材料的不断发现和研究程度的不断深入。 相似文献
6.
中国恐龙蛋化石及其生态地层浅析 总被引：2，自引：0，他引：2
文章系统地对中国恐龙蛋化石的地理分布作了总结,并将恐龙蛋化石类型归纳为９科,１３属,２１种。在此基础上,着重分析了恐龙蛋化石古生态特征,分别从蛋化石的排列方式,恐龙产蛋的方式,不同类型蛋化石间及蛋化石与骨骼化石共生几方面作了探讨,并进一步阐述恐龙蛋化石在晚白垩世地层划分对比所具有的直接指示意义 相似文献
7.
著名的埃迪卡拉纪化石Dickinsonia是最有争议的一种生物,曾被认为是刺胞动物、扁行动物的、环节动物,甚至被归入非后生动物文德生物界或地衣类。从俄罗斯白海地区冬季海岸5亿5千5百万年前的细砂岩中新发现的一枚化石标本保存了可靠的内部器官解剖构造印痕,从而肯定了它是后生动物的属性。这些构造可与现生的栉水母动物的胃水管系统类比。另外,我们观察发现Dickinsonia显示了两辐射对称的特性,并非以前认为的两侧对称生物。而栉水母动物门是现生36个后生动物门类中唯一具两辐射对称的动物类别。因此,我们认为Dickinsonia无疑属于后生动物,… 相似文献
8.
龙门山地区泥盆纪地层中含有种类丰富、数量众多的海相遗迹化石。以野外剖面实测为基础,对北川甘溪石沟里剖面中泥盆统养马坝组风暴沉积中的遗迹化石进行了系统描述,共识别出5种类型的遗迹化石属种,包括Chondrite、Changchengia、Rhizocorallium、Skolithos和Zoophycos。根据遗迹化石的组合特征,将其划分为Chondrites-Zoophycos遗迹化石组合类型,反映了受风暴影响的浅水沉积环境。通过遗迹化石的分布位置、相互间的切割关系,建立了龙门山地区石沟里剖面养马坝组风暴岩中各种遗迹化石形成的先后顺序,依次为Chondrites、Zoophycos、Rhizocorallium、Changchengia和Skolithos。发育于风暴岩中的Chondrites和Zoophycos遗迹化石,具有鲜明的机会种特征,对沉积环境有重要的指示意义。保存于龙门山地区泥盆系养马坝组风暴岩中丰富的具细小分支直径的Chondrites遗迹化石,反应水体氧含量并非控制Chondrites潜穴直径大小的唯一控制因素,可能与造迹生物生活环境的恶劣与否间存在着一定关系;风暴沉积中发育的Zoophycos遗迹化石,指示其在泥盆纪可以沉积于浅海环境。 相似文献
9.
我国贵州瓮安陡山沱期磷块岩中的磷酸盐化球状化石是近年来受到广泛关注的一类微体化石。对于它们的生物亲缘认识，长期以来存在着动植物之争，特别是近年来一些学者发现，部分磷酸盐化球状化石内部保存了2、4、8、16等几何级数增长的细胞特征，根据化石的大小、细胞排列方式和表面纹饰可与现生甲壳类动物的胚胎比较，认为它们很可能是运动胚胎化石，但另一些学者却持截然不同的观点，认为它们不是动物胚胎化石的观察与研究，同时结合相同层位岩石薄片所发现的大量同类化石微细结构的对比。特别是把磷块岩中的化石与燧石中发现的化石详细比较，对一些磷酸盐化球状化石的归属和亲缘关系进行探讨，研究表明贵州瓮安陡山沱期燧石中发现的化石详细比较，对一些磷酸盐化球状化石的归属和亲缘关系进行探讨。研究表明贵州瓮安陡山沱期磷酸盐化微体化石群具有多样性特征，包含不同门类，是迄今为止全球保存最好的晚新元古代磷酸盐化的化石库。 相似文献
10.
通过近年的野外工作,证明巨型山东龙产于王氏群下部,从而修正了以前认为是王氏群上部的说法。晚白垩世早期的鸭嘴龙化石在世界上还未发现过,巨型山东龙填补了这一空白。在演化上提出平头型鸭嘴龙亚科为主支,栉龙亚科和兰氏龙亚科均为其侧支。原角龙科骨骼和蛋化石在王氏群上部首次发现,同时在亚洲也是首次具有确切层位和共生动物群及其地质时代的发现。在同层中还发现了大型的长形蛋化石,证明在层位上已接近于上白垩统顶部。 相似文献
11.
R. J. Larson 《Estuaries and Coasts》1986,9(4):376-379
The sea nettle,Chrysaora quinquecirrha, can capture ctenophore prey on its exumbrella, tentacles, and oral arms. Once attached to the medusa, the prey is transferred to the oral arms, where it becomes enveloped by the lip-like folds of the oral arms. The oral arms move the prey as far proximally as possible by ciliary creeping and muscular contractions. If the prey is too large to enter the stomach, it is digested within the oral arms by extracellular, proteolytic enzymes that probably originate from the gastrodermis of the oral arms and/or from the gastric cirri. Several large ctenophores can be digested simultaneously, owing to the extensive surface areas of the oral arms. Of the threeC. quinquecirrha medusae maintained in the laboratory for up to 60 d, the smaller two specimens (25–150 g wet weight) showed a biomass doubling in ～14 d; the largest specimen (300 g), a sexually mature female, showed little growth but did release >100 ova 24 h?1. Growth efficiencies were estimated to be ～2% for the largest specimen (not including biomass of gametes) and ～10% for the smaller two specimens. These growth efficiencies are comparable to those of other gelatinous predators. 相似文献
12.
The ctenophore Mnemiopsis leidyi is an ecologically important predator in temperate coastal environments. Their populations fluctuate seasonally, serving as sinks of nutrients during periodic blooms, but as sources via excretion and during population collapse. Ctenophores were analyzed for elemental composition (C, N, and P) during 2008 and 2009 in Great South Bay, NY, USA. Salt-free weight percent C, N, and P correlated positively with ctenophore sizes and zooplankton prey abundances. Nitrogen and P were higher at the onset of blooms than during collapse when prey were substantially fewer. Ctenophores collected during average to high zooplankton densities had atomic ratios averaging C/N ~6:1 and C/P ~66:1, but became C- and P-depleted (C/N ~5:1, C/P ~128:1) with decreasing zooplankton. Incubations demonstrated rapid remineralization of ctenophore biomass (as NH4 +, HPO4 2?), following first order kinetics (e.g., k ~0.1–0.4 day?1) with enriched stoichiometric N and P fractionation relative to biomass under both oxic and anoxic conditions. Based on reported excretion rates, nutrient regeneration from excretion by active populations greatly exceeds nutrients remineralized during population crashes. To our knowledge, this is the first study documenting natural seasonal patterns in ctenophore elemental stoichiometry as a function of ctenophore size and prey availability. 相似文献
13.
Thermobarometry and Geotectonic Significance of High-Pressure Granulites: Examples from the Moldanubian Zone of the Bohemian Massif in Lower Austria 总被引：14，自引：1，他引：14
Petrographic details together with mineral and whole-rock compositiondata are provided for acid-intermediate garnet granulites fromexposed granulite complexes in Lower Austria. Thermobarometricevaluation integrated with available isotopic age data indicatesthe initial equilibration of early Variscan ({small tilde}370Ma) high-pressure granulite assemblages at {small tilde}16 kbarand 1000C and their partial overprinting by retrograde assemblageswhich reflect the blocking of mineral exchange reaction equilibriaat {small tilde}6•5 kbar and 725C during subsequent Variscanuplift and cooling. These calculated P-T estimates, togetherwith general phase equilibria constraints and evidence frompreserved prograde coronitic reaction textures and garnet compositionalzoning profiles, indicate a clockwise P-T-t evolutionary pathof the type expected during crustal thickening in a major platecollision orogen and characterized by near-isothermal decompressionduring initial uplift. Geochemical characterization of the rockprotoliths as calc-alkaline igneous rocks and the high metamorphictemperatures suggest that garnet granulite formation involvedthe subduction of a magmatic arc at a continental plate margin. Reviewed evidence from granulites in the Central European Variscidesruns counter to suggestions by Bohlen (1987, 1991) that high-pressuregranulites are of little regional geotectonic significance incomparison with low- to medium-pressure granulites. The differentevolutionary P-T paths for these two important groups of granulitespoint to formation in contrasting plate settings. However, questionsare raised regarding petrogenetic models for low- to medium-pressuregranulites which have emphasized the importance of magmatic,rather than tectonic, crustal thickening and the recognitionof stabilization along deduced anti-clockwise P-T-t paths characterizedby post-peak near-isobaric cooling. It is suggested here that the reality of stabilization of atleast some low- to medium-pressure granulites in a collisionaltectonic regime may have been concealed either because lower-pressureassemblages have overprinted mineralogical evidence for an earlierhigh-pressure history at deeper crustal levels or through invaliddeduction of near-isobaric cooling trajectories as a resultof the different closure temperatures for the mineral reactionsused to monitor the equilibration temperatures and pressuresin granulites. However, the sequential underthrusting modelfavoured for the tectonometamorphic evolution of the Variscannappe pile in the Bohemian Massif renders it unlikely that alllate Variscan low- to medium-pressure granulites have experiencedthe early Variscan high-pressure metamorphism. 相似文献
14.
Documentation of non- or weakly biomineralizing animals that lived during the Furongian is essential for a comprehensive understanding of the diversification dynamics of metazoans during the early Palaeozoic. However, the fossil record of ‘soft’-bodied metazoans is particularly scarce for this critical time interval, consisting of rare fossils found at a dozen or so localities worldwide. Here we report new occurrences of exceptional preservation in Furongian (Jiangshanian) strata of the McKay Group near Cranbrook, British Columbia, Canada. This locality had already yielded trilobites with phosphatised guts, with all specimens representing the same species and occurring within a 10-m-thick interval. Two stratigraphically higher horizons with soft-tissue preservation are reported herein; one has yielded a ctenophore and an aglaspidid arthropod, the other a trilobite with a phosphatised gut belonging to a different species than the previously described specimens. The ctenophore represents the first Furongian record of the phylum and the first reported occurrence of Burgess Shale-type preservation in the upper Cambrian of Laurentia. The aglaspidid belongs to a new species of Glypharthrus, and is atypical in having twelve trunk tergites and an anteriorly narrow ‘tailspine’. These features suggest that the tailspine of aglaspidids evolved from the fusion of a twelfth trunk segment with the telson. They also confirm the vicissicaudatan affinities of these extinct arthropods. Compositional analyses suggest that aglaspidid cuticle was essentially organic with a thin biomineralised (apatitic) outer layer. The trilobite reveals previously unknown gut features, such as medial fusion of digestive glands, possibly related to enhanced capabilities for digestion, storage, or the assimilation of food. The new fossils demonstrate that conditions conducive to soft-tissue preservation repeatedly developed in the outer shelf environment represented by the Furongian strata near Cranbrook. Future exploration of the c. 600-m-thick, mudstone-dominated upper part of the section may result in more abundant discoveries of exceptional fossils. 相似文献
15.
Diversity of palmately lobed leaves of angiosperms of the early–middle Albian floras of the Kolyma River Basin, the Omsukchan Coal Basin, Khabarovsk and Primorye Regions was studied. Leaf fossils, previously compared with those of genera Aralia, Sassafras and Lindera, now reassigned to the fossil genus Araliaephyllum. Four new combinations and one new species are published: A. kolymense (Kryshtofovich) Golovneva, comb. nov., A. luciferum (Kryshtofovich) Golovneva, comb. nov., A. ussuriense (Krassilov) Golovneva, comb. nov., A. ievlevii (Samylina) Golovneva, comb. nov., and A. popovii Golovneva, sp. nov. The type material has been restudied in detail, and lectotypes have been selected to all newly typified species. These species share many lauralean morphological and venation features. They represent the most likely early members of this group. This relationship is based on detailed study of the leaf architecture and comparison with other fossils with studied epidermal features. This study expands our knowledge of radiation and biogeography patterns of early angiosperms in northeastern Asia. 相似文献
16.
Oliver W.M. Rauhut 《Proceedings of the Geologists' Association. Geologists' Association》2012,123(5):779-786
Abelisauroidea are a recently recognized clade of theropod dinosaurs that have a predominantly Gondwanan distribution. Recently, a distal theropod tibia from the Middle Jurassic of England was identified as an abelisauroid, representing one of the oldest records of the group in general and the only Jurassic occurrence in Europe. On this basis, rapid radiation of abelisauroid and a global distribution of this clade in the Jurassic were suggested. Here, the specimen in question is re-examined and the characters used for referral to the Abelisauroidea are re-evaluated. None of the proposed characters can be demonstrated to represent abelisauroid synapomorphies and all have a wider distribution; especially coelurosaurian theropods, which are known from contemporaneous beds in England, frequently show the same character combination. Thus, there is currently no secure evidence for the occurrence of abelisauroids in the Jurassic of the northern Hemisphere, and the early evolution of this clade remains poorly known. Furthermore, other fragmentarily known taxa previously referred to Abelisauroidea based on putative synapomorphies of the distal tibia, such as Ozraptor and Austrocheirus, should be considered as Theropoda indet. 相似文献
17.
WANG Xuri Jingmai K. O’Connor ZHAO Bo Luis M. Chiappe Gao Chunling Cheng Xiaodong 《《地质学报》英文版》2010,84(2):247-256
<正>We report on a new species of enantiornithine bird from the Lower Cretaceous Qiaotou Formation of northern Hebei,China.The new taxon,Shenqiornis mengi gen.et sp.nov.,possesses several enantiornithine synapomorphies but is unique from other known species.The specimen has a well-preserved skull that reveals new information about enantiornithine cranial morphology.The new taxon possesses a large postorbital with a long tapering jugal process indicating that some enantiornithines may have had a fully diapsid skull,as in Confuciusornis.The tooth morphology of the specimen is unique and likely represents a previously unknown trophic specialization within Enantiornithes. 相似文献
18.
Patricia Kremer 《Estuaries and Coasts》1979,2(2):97-105
The impact ofMnemiopsis leidyi as a predator was studied quantitatively to determine their feeding rates on mixed natural zooplankton. These studies showed that feeding rate was independent of food concentration, but a function of both temperature and size of ctenophore. The feeding rate (liters cleared per mg dry weight per day) ranged from about 0.01 for larger ctenophores at lower temperatures (10–15°C) to about 0.1 for smaller ctenophores at higher temperatures (20–25°C). Combining these results with ctenophore biomass estimates from three years of sampling, numerical estimates were made of zooplankton mortality due to ctenophore predation. The maximum summer cropping byM. leidyi was calculated to be an average of 5–10% per day for the bay as a whole. Although substantial, this predation pressure alone does not account for the observed summer decline of zooplankton. 相似文献
19.
K-feldspar megacrysts (Kfm) are used to investigate the magmaticevolution of the 7 Ma Monte Capanne (MC) monzogranite (Elba,Italy). Dissolution and regrowth of Kfm during magma mixingor mingling events produce indented resorption surfaces associatedwith high Ba contents. Diffusion calculations demonstrate thatKfm chemical zoning is primary. Core-to-rim variations in Ba,Rb, Sr, Li and P support magma mixing (i.e. high Ba and P andlow Rb/Sr at rims), but more complex variations require othermechanisms. In particular, we show that disequilibrium growth(related to variations in diffusion rates in the melt) may haveoccurred as a result of thermal disturbance following influxof mafic magma in the magma chamber. Initial 87Sr/86Sr ratios(ISr) (obtained by microdrilling) decrease from core to rim.Inner core analyses define a mixing trend extending towardsa high ISr–Rb/Sr melt component, whereas the outer coresand rims display a more restricted range of ISr, but a largerrange of Rb/Sr. Lower ISr at the rim of one megacryst suggestsmixing with high-K calc-alkaline mantle-derived volcanics ofsimilar age on Capraia. Trace element and isotopic profilessuggest (1) early megacryst growth in magmas contaminated bycrust and refreshed by high ISr silicic melts (as seen in theinner cores) and (2) later recharge with mafic magmas (as seenin the outer cores) followed by (3) crystal fractionation, withpossible interaction with hydrothermal fluids (as seen in therim). The model is compatible with the field occurrence of maficenclaves and xenoliths. KEY WORDS: Elba; monzogranite; K-feldspar megacrysts; zoning; magma mixing; trace element; Sr isotopes; petrogenesis 相似文献