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The increasing number of fossil pterosaur eggs sheds light on nesting environments and breeding be- haviors of these extinct flying reptiles. Here we report the first partial three-dimensional egg of the pterosaur, Pteroduustro guinazui, from central Argentina. The specimen was discovered from the same Albian deposits as the exceptional R guinazui embryo described in 2004. Microscopic characterizations indicate a pristine preservation of the 50 Bm thick calcium carbonate, which differs significantly from the soft shell of Chinese pterosaur eggs. Estimate of the eggshell conductance implies that the nest had a minimum moisture content of 75%. This moisture estimate, combined with geological and taphonomical data, suggests that P. guinazui may have adopted a nesting strategy similar to those of grebes and flamingos rather than being buried on land, as previously hypothesized. Moreover, our results demon- strate that the nesting paleoenvironment of this pterosaur species was closely linked to a mesohaline lacustrine ecosystem in a basin governed by regional tectonic subsidence, a setting characteristic for the feeding and reproduction of modern flamingos.  相似文献   

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Phylogenetic analyses of early pterosaur species are relatively new and contradictory. However, they imply a similar evolutionary history for early pterosaurs: a large divergence of all major early groups of pterosaurs before the first named pterosaur species appeared in the fossil record. This large, unsampled divergence may be a record of the initial adaptive radiation of the pterosaurs, but a recent comprehensive phylogenetic analysis reconstructs named Triassic pterosaur species grouped together in a monophyletic clade with all other major groups diverging later. Ten patterns that characterize adaptive radiations put forward by Gavrilets and Losos (2009) are assessed for congruence with the comprehensive phylogeny and two previous phylogenies to determine which are consistent with our notions of an adaptive radiation. Of the eight patterns that can be applied to pterosaurs, only six vary between pterosaur phylogenies. Of these six patterns, all apply to the three phylogenies with varying amounts of support. For many patterns, however, the comprehensive phylogeny has a greater magnitude, shorter time span, or is the only topology within which they can be assessed. Pterosaurs are characterized by an adaptive radiation no matter which phylogeny is considered, but congruence with the fossil record increases the support and information content of a phylogeny by reducing the time and area within which to reconstruct evolutionary history.  相似文献   

4.
The cross-section and surface structures of wing membranes from the ctenochasmatid pterosaur Beipiaopterus chenianus were observed through a scanning electron microscope (SEM). The results show that the wing membrane contains a high density of blood vessels, implying strong thermoregulatory function, similar to that of a bat wing membrane. This is the first comparison of the microstmctures of pterosaur wing membranes with those of the modem bat. It is inferred that a bat-like physiology exists, at least in relatively small pterosaurs suggesting that these pterosaurs were warm-blooded, active fliers.  相似文献   

5.
New Azhdarchid Pterosaur from the Early Cretaceous of Western Liaoning   总被引:5,自引:0,他引:5  
A nearly complete skeleton with a lower jaw of pterosaur from the Early Cretaceous of western Liaoning is described and assigned to a new genus, Eoazhdarcho gen. nov.. The new genus is characterized by a relatively small size,the ratio of the length to width of the middle series cervical vertebrae approximately 3.5 and the ratio of humeral length to femoral length approximately 0.96. The humerus of Eoazhdarcho shows great resemblances to that of previously described Azhdarchidae, so it is assigned to the family Azhdarchidae.  相似文献   

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<正>A new boreopterid pterosaur:Zhenyuanopterus longirostris gen.et sp.nov.from the Yixian Formation of western Liaoning is erected,based on the complete skeleton with a skull and lower jaws preserved.It is characterized by:a large boreopterid pterosaur with a high number of teeth,where the anterior teeth are much larger than posterior ones;the length of the dorsal + sacral vertebrae is nearly half the length of the skull;ratio of the length of the humerus to metacarpal IV is approximately 91%and the,humerus,femur and third wing phalanx are all equal in length and the feet are specially small.It represents the largest boroepterid pterosaur discovered from western Liaoning and its surrounding areas so far  相似文献   

7.
A new Sordes-like pterosaur Jianchangopterus zhaoianus gen. et sp. nov. is erected based on a almost complete skeleton with skull preserved. It is characterized by the following characters: seven and six pairs of upper jaw and lower jaw teeth respectively; the development of a recess on maxilla; the evenness of the dental margin in lateral view; a distinct central ridge along dorsal surface of the mandibular symphysis; wing phalanx 4 with strongly curved shaft approximately 96% the length of the wing phalanx 1. The discovery of a Sordes-like pterosaur Jianchangopterus, and other taxa from the same formation indicates that pterosaurs reached great diversities during the Middle to Upper Jurassic period.  相似文献   

8.
A new baby pterodactyloid pterosaur with soft tissue preserved, Ningchengopterus liuae gen. et sp. nov., is erected based on a nearly complete skeleton with a skull. It was characterized by the skull, slightly longer than the combined length of the dorsal and sacral vertebrae; 50 teeth (including upper and lower jaws); short mid-cervical vertebrae; the humerus and the scapula, equal to that of the wing metacarpal in length; the ulna and the femur, equal to that of the first and third wing phalanx in length, respectively. The similar ratio of the wing phalanx 2 to wing phalanx 1 of Ningchengopterus and Eosipterus implies that Ningchengopterus may be close to the ctenochasmatid pterosaur. However, it may also imply that the isometrically growing of the first two wing phalanges exists among some pterosaurs.  相似文献   

9.
The anatomical term tuba vertebralis was introduced by Martill et al. (1998) to describe a bony tube which enclosed the spinal cord in the type cervical vertebra of the giant azhdarchid pterosaur Arambourgiania philadelphiae, from the Late Cretaceous of Jordan.  相似文献   

10.
A new pterosaur Moganopterus zhuiana gen. et sp. nov. is erected based on a complete skull with lower jaws and anterior cervical vertebrae. It is characterized by much elongated upper and lower jaws with at least 62 total, long, curved teeth with sharp tips, a well developed parietal crest extending posterodorsally, forming an angle of 15 degrees with the ventral margin of the skull, the ratio of length to width of cervical vertebrae greater than 5:1. The skull length is 750 mm, and it is the largest toothed pterosaur found so far in the world. Based on this new pterosaur, the Boreopteridae can be divided into two subgroups: Boreopterinae sub-fam. nov. and Moganopterinae sub-fam. nov., which is also confirmed by the phylogenetic analysis.  相似文献   

11.
甘肃永靖盐锅峡早白垩世翼龙足迹的发现及意义   总被引:3,自引:0,他引:3  
甘肃永靖盐锅峡恐龙足迹1号化石点中保存有翼龙足迹, 该足迹是我国翼龙足迹的首次发现.翼龙足迹为四足行走, 行迹宽.前脚为不对称三趾型, 趾行式, 其中Ⅱ趾最短, Ⅳ趾最长, 足迹外偏, 偏角大, Ⅳ趾与行迹交角为160°~ 180°.后脚外形为近长方形, 四趾型, Ⅱ、Ⅲ趾较Ⅰ、Ⅳ趾长, 行式, 脚长为宽的3倍, 足迹外偏, 偏角30°.足迹特征表明该翼龙足迹属于一新种.沉积特征表明翼龙足迹产于湖岸环境.   相似文献   

12.
A new pterosaur, Microtuban altivolans gen. et sp. nov., is described from the Sannine Formation of northern Lebanon. The specimen is the first pterosaur from the Early Cenomanian (Late Cretaceous) locality of Hjoûla and is regarded as the most complete pterosaur fossil discovered from Africa. While postcranial characters indicate a possible relationship with members of the Thalassodromidae or Chaoyangopteridae, the specimen possesses an exceptionally short wing-finger phalanx 4, forming only 1.1% of the total length of the wing-finger. Its appearance along with an unnamed ornithocheiroid from the slightly younger locality of Hâqel suggests that a number of pterosaur taxa existed within the local area, perhaps living on exposed carbonate platforms.  相似文献   

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姬书安  张立军  路芳 《地质学报》2023,97(6):1723-1740
中国辽宁西部早白垩世翼龙类化石非常丰富,其上部的九佛堂组翼龙类组合以含有丰富的无齿的古神翼龙科、朝阳翼龙科等进步类型而与下部的义县组翼龙类组合相区别。本文描述了辽宁西部建昌盆地九佛堂组一新的大型翼龙类化石,其以上下颌无齿、吻端指数为3.7、前颌骨背支细长且封闭鼻眶前孔背缘、很大的鼻眶前孔后端超过上下颌关节位置、轭骨的上颌骨突基部宽大而应被归入朝阳翼龙科(Chaoyangopteridae)神州翼龙属(Shenzhoupterus)。其以较大的体形(翼展2.05 m)、平直的上下颌咬合面、轭骨的眶后骨突较泪骨突长且基部略宽、第4~7颈椎长度依次减小等特征,区别于朝阳神州翼龙(Shenzhoupterus chaoyangensis),而被命名为三亚神州翼龙(新种)(Shenzhoupterus sanyainus sp. nov.)。这是辽宁西部早白垩世翼展最大且不具牙齿的翼龙类属种,它的发现丰富了九佛堂组翼龙类组合内容,对认识朝阳翼龙科的骨骼形态与生态习性亦具有积极意义。  相似文献   

15.
姬书安  张笠夫 《地学前缘》2020,27(6):365-370
发现于内蒙古鄂托克旗召稍早白垩世罗汉洞组的一件不完整翼龙类下颌标本,以下颌愈合部长且平直、齿骨侧嵴发育、近圆形齿窝沿侧嵴之上的齿骨侧面上半部分自前向后呈直线排列、齿窝直径(2~2.5 mm)前后变化较小、相邻齿窝的间距约为齿窝直径的一半、下颌牙齿密度为3枚/cm等特征,可归入梳颌翼龙科(Ctenochasmatidae),且代表一新属种——郝氏鄂托克翼龙Otogopterus haoae gen.et sp.nov.。该化石是继鄂托克旗新召准噶尔翼龙科的平颌鄂尔多斯翼龙Ordosipterus planignathus之后在内蒙古鄂尔多斯地区发现的第2件确切翼龙类材料,丰富了这一地区早白垩世脊椎动物群的组成。该翼龙也是继甘肃庆阳环河翼龙Huanhepterus quingyangensis之后在鄂尔多斯盆地发现的第2种梳颌翼龙科化石,进一步扩大了鄂尔多斯盆地梳颌翼龙科的地理分布范围,同时表明鄂尔多斯盆地是继辽宁西部之后梳颌翼龙科在中国的又一重要分布区。  相似文献   

16.
Fragmentary remains, including cervical vertebrae and limb bones, of a large pterosaur from the Upper Cretaceous of Tous, province of Valencia (Spain), are described. The material was recovered from lacustrine beds in the upper part of the Calizas y Margas de Sierra Perenchiza Formation, which is probably Maastrichtian in age. Six fragments of vertebrae allow a reconstruction of the anatomy of the mid-series cervicals of the animal. The general morphology of the cervical vertebrae is closely similar to that of the long-necked Azhdarchidae. Compared to other azhdarchids, the Valencia pterosaur shows minor differences from the genera Azhdarcho and Quetzalcoatlus, and is here provisionally referred to as Azhdarchidae indet. A wingspan of about 5.5 m is calculated by comparison with other known azhdarchids. This is the second azhdarchid pterosaur described from the Iberian peninsula. It confirms the wide distribution of this group of large pterosaurs at the end of the Cretaceous.  相似文献   

17.
本文报道了浙江东阳发现的中国第2个翼龙足迹化石点, 也是亚洲发现的第8个翼龙足迹化石点。化石足迹产于晚白垩世早期方岩组的紫红色泥质粉砂岩中。发现有3个手的印迹和1个右足印。手的印迹的长宽分别为6.5 cm 和4cm. 非对称具有3个指的印迹。足迹9 cm长和1.5 cm宽。该足迹不同于以前发现的, 可能代表一新的类型。除了翼龙足迹外, 还发现鸟类、小型兽脚类、鸟脚类及蜥脚类脚印, 形成一个丰富的动物群, 它的发现为研究该地区的古生态环境及古地理具有重要意义。同时为以后在该地区发现这些造迹动物的骨骼化石提供了重要依据。  相似文献   

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