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1.
The Eyam Limestone Formation of Steeplehouse Quarry, Wirksworth, Derbyshire, UK yields a diverse assemblage of Lower Carboniferous vertebrate remains. The assemblage is dominated by dermal denticles of the enigmatic selachian Petrodus patelliformis M’Coy, 1848, but also contains teeth of petalodonts, hybodonts and neoselachians. Actinopterygian remains also occur. The assemblage has yielded the earliest Neoselachian, Cooleyella fordi (Duffin and Ward, 1983) and the earliest British lonchidiid, Reesodus wirksworthensis (Duffin 1985). The first occurrence of the enigmatic spiny shark Acanthorhachis (Listracanthidae) is reported from the Viséan, extending its range back some 10 million years. Associated invertebrate remains and sedimentological data indicates a thriving fore-reef environment, deposited in a low energy off-reef setting. The vertebrate remains are well preserved with little abrasion, indicating short transport distances. Conodont elements indicating a late Brigantian age (Early Carboniferous, Viséan) have unusual and extensive euhedral apatite overgrowths.  相似文献   

2.
In the first ever systematic study of trace fossils from the Badhaura Formation, the authors described a nesting burrow, which they ascribed to a stomatopod. The purpose of this paper is two-fold: primarily, to document ichnofauna from (post-glacial marine late Palaeozoic rocks of peninsular India) the Badhaura Formation (Sterlitmakian) representing marine rocks deposited following the Late Palaeozoic glaciation and secondly to contribute to the data on post-glacial ichnofauna from constituent continents of the Gondwanaland. Trace fossils described here are from the Harbans Bed, the topmost lithounit of the Badhaura Formation. The ichnofauna includes Arenicolites tenuis, Beaconites isp., Curvolithus isp., Cylindrichnus concentricus, Didymaulichnus lyelli, Ophiomorpha isp., Palaeophycus tubularis, Planolites beverleyensis, P. montanus, Rosselia chonoides, R. socialis, Skolithos linearis, Taenidium cameronensis, Thalassinoides paradoxicus, Thalassinoides isp. and a flask-shaped brood chamber assigned to a stomatopod crustacean. This mixed assemblage is assigned to distal Skolithos ichnofacies and is suggestive of a period of relatively quiet, shallow water conditions of deposition. The ichnofauna, when viewed in context of peri-gondwanic ichnofaunas, mainly consisting of simple tracks and trails, from late Palaeozoic post-glacial deposits of other Gondwanan continents, is interesting due to dominance of domichnia. Profusion of brood chambers along with Thalassinoides in the Badhaura Formation validates the concept of pre-Mesozoic Thalassinoides being non-decapod in origin and suggestive of adaptive convergence.  相似文献   

3.
The fossil record of mid to late Permian terrestrial vertebrates in the South African Karoo Basin is regarded as the most abundant and diverse in the world. Despite the extensive research on body fossils, to-date the vertebrate footprint sites have not been subjected to an anatomy-consistent ichnotaxonomic investigation. Here we present a comprehensive ichnotaxonomic revision of Permian-Early Triassic tracksites in the main Karoo Basin of South Africa. Furthermore, a track-trackmaker correlation for all Permian synapsid groups is provided for the first time, based on the analysis of the functional morphology of potential producers. The following ichnotaxa and their proposed trackmakers are recognized: Brontopus giganteus (dinocephalians), cf. Capitosauroides isp. (therocephalians), cf. Dicynodontipus isp. (cynodonts), Dolomitipes accordii (small dicynodonts), Dolomitipes icelsi n. comb. (large dicynodonts), Karoopes gansfonteinensis n. igen. n. isp. (gorgonopsids), Procolophonichnium nopcsai (procolophonids) and Rhynchosauroides isp. (non-archosauriform diapsids). Three different footprint assemblages (FA I–III) are proposed for footprint biostratigraphy: FA I (lower Tapinocephalus AZ), a Guadalupian assemblage dominated by dinocephalian tracks; FA II (topmost Tapinocephalus-Cistecephalus AZ), a latest Guadalupian-Wuchiapingian assemblage dominated by gorgonopsid and dicynodont tracks in association with subordinate therocephalian tracks and FA III (lower Lystrosaurus AZ), an Induan assemblage with dicynodont, cynodont, procolophonid and diapsid tracks. The lower FA II includes the earliest ichnofauna with Lopingian affinity all over the world (topmost Tapinocephalus-Pristerognathus AZ, ~260–259 Ma) and could indicate an early recovery phase after the end-Guadalupian mass extinction, because of the high abundance of large gorgonopsid tracks and absence of dinocephalian tracks. This footprint record may also predate the body fossil record, suggesting an earlier gorgonopsid radiation. FA III represents the earliest and most complete post end-Permian extinction ichnofauna, which includes an early phase of abundant small dicynodont tracks, potentially indicating a stressed post-event community. Nevertheless, this ichnofauna looks very similar to pre-extinction ichnofaunas from Europe, in agreement with the skeletal record at the Daptocephalus-Lystrosaurus AZ transition.  相似文献   

4.
Trace fossils from the Middle Devonian Caherbla Group of the Dingle Peninsula, southwest Ireland, record a diverse arthropod fauna inhabiting a hot‐arid intracontinental rift setting. Aeolian dunefield and coeval fluvial interdune deposits interfinger spatially and temporally with alluvial fan sedimentary rocks. Three distinct trace fossil assemblages are recognized. The Taenidium‐Scoyenia ichnocoenosis occurs in alluvial fan and fluvial channel deposits, and includes the large backfilled burrow Taenidium, interpreted as eoarthropleurid aestivation chambers. The Rusophycus‐Protichnites ichnocoenosis, composed of arthropod trackways and surface pits, occurs in an interdune ponded area that was susceptible to ephemeral fluvial flow, with Rusophycus showing preferred orientation into the oncoming palaeocurrent. Both the Taenidium‐Scoyenia and Rusophycus‐Protichnites ichnocoenoses are assignable to the globally recurring continental Scoyenia ichnofacies. They are clearly substrate‐controlled and moisture‐related due to the ephemeral nature of the fluvial system. The Palmichnium‐Entradichnus ichnocoenosis occurs in aeolian dune deposits, and includes Palmichnium, attributed to large stylonurid eurypterids, and Diplichnites, attributed to eoarthropleurids. These trackways represent the activities of dune pioneers that left their fluvial habitat to forage for detritus. Interface burrows (Entradichnus, Palaeophycus) were also constructed by arthropods moving just under the sand surface and vertical burrows (Cylindricum, Pustulichnus) were made by arthropods digging downward. Trace preservation in the aeolian environment was probably enhanced by heavy nocturnal dew‐fall or light rain. The Palmichnium‐Entradichnus ichnocoenosis is assigned to the globally recurring aeolian Octopodichnus‐Entradichnus ichnofacies. This aeolian facies, and associated ichnofauna described herein, represents the oldest development of a unique erg system in the Old Red Sandstone (Devonian) of the southern British Isles, and one of the oldest and most diverse aeolian ichnofaunas to be reported worldwide. Copyright © 2011 John Wiley & Sons, Ltd.  相似文献   

5.
The Marwar Supergroup of the Bikaner-Nagaur Basin is composed of sediments deposited from the late Neoproterozoic (Ediacaran) to Upper Cambrian. The Nagaur Sandstone Formation of the Nagaur Group (uppermost division of the Marwar Supergroup) preserves trace fossils significant for establishing Early Cambrian biostratigraphic zones and depositional facies. Fifteen ichnospecies (and eight ichnogenera) identified in the Nagaur Sandstone Formation include “Treptichnus” pedum, Cruziana cf. tenella, Cruziana isp., Diplichnites ispp. A, B, and C, Gyrophyllites isp., Lockeia isp., Merostomichnites isp., Monomorphichnus gregarius isp. nov., Monomorphichnus isp., Planolites isp., Psammichnites isp., Rusophycus bikanerus isp. nov., Rusophycus cf. carbonarius, Rusophycus isp. and radial trace fossils.These trace fossils belong to ethological categories pascichnia, repichnia, cubichnia, and fodinichnia and represent arthropod and worm-like burrowing biota. The assemblage and a regional comparison with contemporaneous trace fossils in the eastern Gondwanan realm suggest that the sequence in the study area belongs to the Cruziana tenella Ichnozone and to Stage 2 (upper part of Terreneuvian), however the Middle Cambrian is not excluded. The trace fossil assemblage belongs to the archetypal Cruziana ichnofacies. Cross bedded sandstone, mud cracks and rainprints in the ichniferous strata of the Nagaur Sandstone Formation indicate deposition in an intertidal sand flat with channels that was exposed episodically.  相似文献   

6.
A diverse fossil vertebrate record in the Devonian of coastal southeastern Australia includes at least 30 genera and species representing all three major groups of extant jawed vertebrates (bony and cartilaginous fishes, and tetrapods), and both extinct groups (placoderm and acanthodian fishes). A bone recorded by W. B. Clarke in 1860 from Twofold Bay is the first published record of a Devonian vertebrate from the Southern Hemisphere. Abundant plant remains at some localities include large impressions of arborescent lycopsids, indicating one of the earliest forest environments for the Gondwana supercontinent. An early terrestrial invertebrate fauna is evidenced by fossil millipede remains. A review of the evidence for age control using paleontology and isotopic dating suggests that the lower part of the Devonian sequence (Bunga beds, Boyd Volcanic Complex) may be somewhat older (Emsian – Eifelian) than recently proposed; the tetrapod trackways at Genoa River in Victoria (lower part of the Combyingbar Formation) may correlate with the lowermost Twofold Bay Formation of the Merrimbula Group, which is overlain by the marine Bellbird Creek Formation, a manifestation of the global late Frasnian marine transgression and extinction event. Abundant placoderm remains at all higher fossil fish horizons in the overlying Worange Point Formation indicate a latest Devonian rather than Carboniferous age for the upper Merrimbula Group.  相似文献   

7.
Sediments of Lower Carboniferous age in eastern Menorca, Balearic Islands contain a diverse and exceptionally well preserved ichnofauna, including Neonereites biserialis, Nereites isp., Arthrophycus isp., Dictyodora liebeana, two ichnospecies of Chondrites, several ichnospecies of Lophoctenium, two ichnospecies of Phycosiphon, Syncoprulus pharmaceus, annulated burrows and a vertical burrow. The host lithologies are conglomerates, sandstones, siltstones and mudstones; most are the product of debris flows, and of high concentration [Ta(bc) intervals] to relatively dilute (Tcd/Tde intervals) turbidity currents. The rocks are interpreted as deposits of an inner- to mid-fan palaeoenvironment. Channelized deposits, sequences of overbank deposition and interchannel deposits interspersed with the deposits of unconfined debris flows and high concentration turbidity currents are present. The ichnofauna is most frequently, and best, preserved within the Td interval of turbidities, which are interpreted as interchannel deposits, produced by low concentration turbidity flows. The distribution of the ichnofauna is partly controlled by the lithologies in which they are preserved. The rarity and poor preservation of trace fossils in the coarser grained facies contrasts with the detailed preservation of very delicate traces in the finer grained lithologies. However, the ichnofauna is also partitioned between different subenvironments of the mid-fan to produce a series of palaeoichnocoenoses. Thin intervals of interchannel deposits, separated by deposits of high concentration turbidity currents, repeatedly contain only Phycosiphon incertum and small (?juvenile)Dictyodora liebeana. These traces are interpreted as the products of opportunistic colonization of near-channel environments during episodes of quiescent deposition. Thicker intervals of interchannel deposits contain diverse assemblages of trace fossils characteristic of more stable environments, in which widespread colonization occurred. Overbank deposits at Cabo de Favaritz are medium- to thick-bedded, fine-grained beds. In these, the ichnofauna occurs in a simple, two-tier profile. The upper tier is dominated by Nereites isp.; this is underlain by a partially bioturbated layer characterized by large Dictyodora liebeana and Arthrophycus isp.  相似文献   

8.
Two natural casts of two-toed (didactyl) tracks from the Cretaceous (Albian) Plainview Sandstone (Plainview Member) of the South Platte Formation (Dakota Group) at Dinosaur Ridge, Colorado are attributed to deinonychosaurian theropod dinosaurs and placed in the ichnogenus Dromaeosauripus. This is both the first report of tracks from this unit in the Dinosaur Ridge area and the first report of deinonychosaurian tracks from Colorado. It is also only the third report of this track type from North America. The rarity of tracks from the Albian-aged, Plainview Sandstone (Dakota Group Sequence 2) contrasts with their abundance in the upper (Cenomanian) part of the overlying South Platte Formation (Dakota Group Sequence 3), which has yielded more than 120 sites mostly in Colorado, giving rise to the “Dinosaur Freeway” concept. As no deinonychosaurid tracks are known from the sequence 3 part of the South Platte Formation, despite the large vertebrate and invertebrate ichnological database available, it is evident that the sparse vertebrate ichnofauna from the Plainview Member (Sequence 2) is inherently different. This striking difference in both track abundance and track type reflects differences in both age and depositional environment. Based on the Albian age, and track type, the Plainview tracks invite comparison with the ichnofaunas of the Cedar Mountain Formation and not with those well-known from the upper part of the South Platte Formation known as the Dinosaur Freeway.  相似文献   

9.
The West Junggar lies in the southwest part of the Central Asian Orogenic Belt (CAOB) and consists of Palaeozoic ophiolitic mélanges, island arcs, and accretionary complexes. The Barleik ophiolitic mélange comprises several serpentinite-matrix strips along a NE-striking fault at Barleik Mountain in the southern West Junggar. Several small late Cambrian (509–503 Ma) diorite-trondhjemite plutons cross-cut the ophiolitic mélange. These igneous bodies are deformed and display island arc calc-alkaline affinities. Both the mélange and island arc plutons are uncomfortably covered by Devonian shallow-marine and terrestrial volcano-sedimentary rocks and Carboniferous volcano-sedimentary rocks. Detrital zircons (n = 104) from the Devonian sandstone yield a single age population of 452–517 million years, with a peak age of 474 million years. The Devonian–Carboniferous strata are invaded by an early Carboniferous (327 Ma) granodiorite, late Carboniferous (315–311 Ma) granodiorites, and an early Permian (277 Ma) K-feldspar granite. The early Carboniferous pluton is coeval with subduction-related volcano-sedimentary strata in the central West Junggar, whereas the late Carboniferous–early Permian intrusives are contemporary with widespread post-collisional magmatism in the West Junggar and adjacent regions. They are typically undeformed or only slightly deformed.

Our data reveal that island arc calc-alkaline magmatism occurred at least from middle Cambrian to Late Ordovician time as constrained by igneous and detrital zircon ages. After accretion to another tectonic unit to the south, the ophiolitic mélange and island arc were exposed, eroded, and uncomfortably overlain by the Devonian shallow-marine and terrestrial volcano-sedimentary strata. The early Carboniferous arc-related magmatism might reflect subduction of the Junggar Ocean in the central Junggar. Before the late Carboniferous, the oceanic basins apparently closed in this area. These different tectonic units were stitched together by widespread post-collisional plutons in the West Junggar during the late Carboniferous–Permian. Our data from the southern West Junggar and those from the central and northern West Junggar and surroundings consistently indicate that the southwest part of the CAOB was finally amalgamated before the Permian.  相似文献   

10.
The Bajiu tracksite exposed on a fine sandstone surface at the top of the Feitianshan Formation, close the contact with the overlying Xiaoba Formation, reveals a saurischian dominated ichnofauna consisting of four sauropod and at least eight theropod trackways. The sauropod trackways display a medium-wide gauge pattern characteristic of titanosauriforms, while the theropod trackways can be subdivided into two distinct morphotypes. One is similar to the ichnogenera Eubrontes and Megalosauripus, the other (cf. Dromaeopodus) appears to represent a functionally didactyl dromaeosaur where pedal digit II is represented only by an oval basal pad. A single swim track is possibly attributable to a turtle or crocodylian. Such sauropod-theropod dominated ichnofaunas are consistently typical of the red-bed tracksites in the region and useful for characterizing a fauna that is otherwise poorly known from body fossils. The stratigraphic position and late occurrence of the (typical Jurassic) Eubrontes-Megalosauripus morphotype is well known from other Cretaceous localities in China and seems to reflect a peculiarity in the theropod faunas of East Asia.  相似文献   

11.
An analysis of lower Barremian invertebrate trace fossils at the El Hoyo dinosaur tracksite (Teruel, Spain) was conducted in order to interpret the macrobenthic tracemaker community and the palaeoenvironmental conditions during bioturbation. Abundant bioturbation structures are characterized by meniscate, cylindrical, straight to sinuous, unbranched epichnia, showing the absence of lining, mantle and longitudinal striations. These structures are included in the ichnogenus Taenidium. Burrow width, distance between menisci, type of their fill, and general shape, allow for identification of the ichnospecies Taenidium serpentinum and T. barretti. Dominance to near exclusiveness of Taenidium in the invertebrate trace fossil assemblage suggests assignation to the Scoyenia ichnofacies. This agrees with the presence of vertebrate tracks. Accordingly, a transitional zone between terrestrial and nonmarine aquatic environments, i.e., floodplain areas adjacent to rivers and affected by low energy conditions, could correspond to the El Hoyo site. The pervasive ichnofabric of Taenidium reveals an opportunistic behaviour of the tracemaker, bioturbating rapidly after deposition; bioturbation took place in moist to wet substrates—softgrounds—in shallow tiers, during favourable yet episodic conditions. The opportunistic strategy might owe to a rapid accumulation of nutrients buried within the sediment during alluvial discharges. After bioturbation, minor erosions during fluvial discharges or subaerial exposure may have caused differences in the preservation of the Taenidium.  相似文献   

12.
There are a growing number of Early Cretaceous avian tracks and trackways from around the world, with Asia (China and Korea) having the largest reported number and diversity of Mesozoic avian traces to date, and these new discoveries are increasing the Early Cretaceous avian ichnodivesrity of Laurasia. Here we report on a new Lower Cretaceous avian track locality in the Guanshan area, Yongjing County, Gansu Province, northwest China, and on a novel ichnospecies of Koreanaornis, Koreanaornis lii ichnosp. nov. Koreananornis lii is distinct from other Koreanaornipodidae in that it possesses a consistently wider digit divarication than previously described tridactyl tracks, and possess a short, small, posteromedially oriented hallux that displays a different orientation than that seen in Koreanaornis hamanensis. The lack of linear and angular data reported for digit I traces of many avian ichnotaxa has the potential to give misleading results in multivariate statistical analyses. Also, the wide divarication of Koreanaornis lii causes the ichnotaxon to not group with other Koreanornipodidae in multivariate analyses, but with Ignotornidae. Despite the results of the analyses, K. lii is morphologically distinct from these ichnotaxa. The results demonstrate that relying solely on multivariate statistical analyses without careful examination of footprint morphology will result in erroneous ichnospecies groupings. While new vertebrate ichnotaxa discoveries from Asia may support the hypotheses of the presence of a unique and endemic Asian vertebrate ichnofauna during the Cretaceous, the recent discovery of skeletal remains interpreted to be of a volant wading bird from the Early Cretaceous, and recent reports of tracks from volant avians, could suggest that flighted avians of the shore- and wading bird ecotypes could have had a Laurasian-wide distribution during the Early Cretaceous. However, strong convergence in foot morphology of shore- and wading birds suggests that avian ichnotaxa found in both present-day Asia and North America may have been made by birds endemic to eastern and western Laurasia during the Early Cretaceous.  相似文献   

13.
The Late Triassic fissure fills from the region of Bristol, SW England and S Wales, preserve unique assemblages of small vertebrates derived from an archipelago of palaeo-islands that document aspects of a critical transition in the history of terrestrial ecosystems. Tytherington Quarry, in south Gloucestershire, is the site of several fossiliferous fissures, all dated as Rhaetian (terminal Triassic), and source of abundant remains of the ‘Bristol dinosaur’, Thecodontosaurus antiquus. In addition, the fissure sediments have yielded previously unreported microvertebrate assemblages, including over 400 jaw remains from three genera of sphenodontians and 100 archosaur teeth assigned to 15 morphotypes. The land fauna is dominated by sphenodontians, with Diphydontosaurus by far the most common form, followed by Clevosaurus, then the sauropodomorph dinosaur Thecodontosaurus, and then the sphenodontian Planocephalosaurus. There are, in addition, rare remains of contemporaneous bony fishes, as well as fossils apparently reworked from the Carboniferous limestones, namely conodonts, holocephalian (chimaeroid) teeth, and a shark tooth. Many typical latest Triassic animals, such as temnospondyls, phytosaurs, aetosaurs, rauisuchians, plateosaurids and dicynodonts are not represented at Tytherington, perhaps because these generally larger animals did not live on the palaeo-island, or because their carcasses could not fit into the fissures. The absence of tritylodonts and early mammals is, however, less easy to explain on the basis of size, although it is known that these forms were abundant here by the Early Jurassic.  相似文献   

14.
《Gondwana Research》2013,23(3-4):1009-1029
The Carboniferous tectonic setting of the Junggar terrane, northern Xinjiang, NW China, has long been a matter of debate. Voluminous Carboniferous volcanic rocks are widely distributed in the Karamaili area, the southern part of the eastern Junggar terrane. Early Carboniferous rocks comprise basalts and basaltic andesites, with enrichment of LREE and LILE and depletion of HFSE, and uniformly high εNd(t) (+ 3.7 to + 4.0). Late Carboniferous rocks consist of basalts, basaltic andesites, rhyolites and minor dacites, and can be subdivided into basic and felsic groups. The basic rocks are depleted in HFSE, and show variable high εNd(t) (+ 4.8 to + 6.9). They have higher Cr and Ni and lower Na2O, U and Th contents than early Carboniferous basic rocks. The felsic rocks show A-type affinity, with typical enrichment of alkalis, LREE and HFSE and strong depletion in Ba, Sr, Eu and Ti. They have high values of εNd(t) and zircon εHf(t) (+ 11.6 to + 17.9). New LA-ICPMS zircon U–Pb analyses constrain their emplacement to late Carboniferous time (306.5–314.3 Ma).The Carboniferous basic rocks show negative Zr-Hf anomalies and low Th/Ce (< 0.07) and Th/La (0.06–0.16), excluding significant crustal contamination during magma evolution. They have low La/Ba (0.03–0.12), Ce/Y (< 3) and (Tb/Yb)N (< 2) and variable Ba/Th (28–318) and Ba/La (3.1–34), suggesting that they were derived from a main spinel with minor garnet lherzolite mantle source metasomatized by slab-derived fluids. The late Carboniferous felsic rocks were produced when upwelling asthenosphere triggered partial melting of juvenile lower crust. The early Carboniferous volcanism occurred in an island-arc setting related to the southward subduction of the Paleo-Junggar Ocean plate, whereas the late Carboniferous rocks erupted in a post-collisional extensional setting. Thus, a rapid tectonic transition from arc to post-collisional extension may have occurred between early and late Carboniferous, and probably resulted from slab break-off or lithospheric delamination.  相似文献   

15.
Lakes are particularly sensitive to environmental fluctuations, which are recorded in their facies and stratigraphy. Ephemeral lakes reveal their sensitivity to palaeoenvironmental changes in the overprinting of the sedimentary features in every single bed. Tetrapod-track taphonomic-modes and ichnological taphonomic-pathways can be used as sensitive indicators of environmental conditions of the track-bearing beds during deposition and imprinting. The Middle Triassic Cerro de las Cabras succession (Cuyana Basin, Argentina) provides an excellent opportunity to these environmental indicators in an underfilled palaeolake. A model of ichnological preservation for underfilled lake systems is proposed and the role of the ichnology record in the sequence stratigraphy analysis is evaluated, based on the integration of tetrapod-track modes, taphonomic-pathways of playa-lake ichnofauna, mineralogy and physical data. Soft-ground suites include those dominated by invertebrate grazing traces and arthropod locomotion traces (Suite 1), and those overprinted by horizontal-vertical dwelling burrows with tetrapod tracks preserved in taphonomic modes B and C (Suite 2). The firm-ground suite (Suite 3) comprises tetrapod-tracks with the best preservation styles (modes A and B) along less abundant invertebrate dwelling and feeding traces as found in Suite 2. Clay mineralogy (dominated by illite with subordinate smectite) suggests low plasticity of the layers, in agreement with low-relief deformation structures observed in tetrapod-track taphonomic-modes. The well-preserved track tetrapod features documented in the Cerro de las Cabras succession, together with the absence of pedogenic disturbance, trampling obliterating the footprints, and/or evidence of strong disturbance by wind, desiccation and/or precipitation, supports short periods of exposure of the imprinted surface particular to this succession. An integrated multiproxy approach is proposed to evaluate the evolutionary interpretation and identification of autogenic versus allogenic controls in underfilled lake-basin histories. The observed aggradational-trend suggests an equilibrium between rates of accommodation change and sediment supply, and that the basin-centre did not experience prolonged sediment-starved conditions.  相似文献   

16.
Ashdown Brickworks, near Bexhill, East Sussex, has produced a large number of vertebrate fossils from the Wadhurst Clay Formation, part of the Wealden Supergroup (Hastings Group; Valanginian; Lower Cretaceous). Here we describe the microvertebrate fauna of the ‘conglomerate bed’, representing a rich sample of taxa. While most of the recovered teeth and bones are abraded, some heavily, most can be identified to species level. The taxa include four species of hybodont sharks (Egertonodus basanus, Planohybodus ensis, Polyacrodus parvidens, P. brevicostatus), three taxa of bony fishes (an unidentified Lepidotes-like semionotiform, the pycnodontiform Ocloedus, and an albuliform), three taxa of crocodyliforms (the goniopholid Hulkepholis, a bernissartiid, and the atoposaurid Theriosuchus), and the theropod dinosaurs Baryonyx and an allosauroid. Sediments of the Wadhurst Clay Formation as a whole indicate freshwater to very slightly brackish-water environments of deposition, and the mainly aquatic time-averaged mixture of fishes and tetrapods recovered from the ‘conglomerate bed’, together with isolated terrestrial species, confirms this interpretation.  相似文献   

17.
The terrestrial carbon cycle and the role of atmospheric CO2 concentrations in controlling global temperatures can be inferred from the study of ancient soils (paleosols). Soil-formed goethite and calcite have been the primary minerals used as a geochemical proxy for reconstructing atmospheric pCO2 from ancient terrestrial records. In the case of goethite, optimum sampling strategies for reconstructing pCO2 focus on the portion of the soil profile that displays steep gradients in both soil CO2 concentration and δ13C values of soil CO2 such that a keeling plot can be developed for a given soil and atmospheric pCO2 can be calculated from it. We report data from a Carboniferous paleosol that depart from the expected linear trends. The results indicate that pedogenic goethite is sensitive to variations in the isotopic composition of soil CO2, over a range of timescales, and can record these variations in the carbon isotope composition and mole fraction of Fe(CO3)OH in solid solution with goethite. We explore possible environmental conditions that can drive these changes as a function of either moisture controlled variations in soil respired CO2 or in the residence time of carbon in soils. The implications of this result are overestimation of paleoatmospheric pCO2 from pedogenic goethite.  相似文献   

18.
The Cambrian succession in the Zanskar Basin of Tethys Himalaya contains an abundant ichnofossils like in the other Tethyan Himalayan successions. The ichnofossils are stratigraphically important as they occur below the trilobite body fossils and are useful to define the basal part of the Cambrian. The ichnofossil assemblage reported from the Zanskar Basin of Ladakh Himalaya is significant to demarcate the Early Cambrian age due to lack of other faunal elements so far. The body fossils of trilobites recorded from the overlying beds indicates the earliest part of the Middle Cambrian age. Sixteen ichnogenera identified include: Bifungites, Cruziana, Diplichnites, Dimorphichnus, Isopodichnus, Lockeia, Merostomichnites; Monomorphichnus, Psammichnites, Palaeophycus, Planolites, Rhizocorallium, Skolithos, Taphrhelminthopsis, Teichichnus, Trepitichnus and trilobite scratch marks etc. The ichnogenera reported so far from this part of the Tethyan Himalayan region belongs mostly to the traces of arthropod origin. The ichnofauna ranges in age from Lower Cambrian to late part of the Middle Cambrian. The ichnofaunal assemblage can be assigned to repichnial, cubichinial, pascichnial, to fodinichnial behaviour. The distribution of ichnofossils in the studied sections shows that the ichnofossils are predominately less in occurrence in the sections were trilobites dominates and higher in the successions the abundance of ichnofossils decreases.  相似文献   

19.
The forests of the Siskiyou Mountains are among the most diverse in North America, yet the long-term relationship among climate, diversity, and natural disturbance is not well known. Pollen, plant macrofossils, and high-resolution charcoal data from Bolan Lake, Oregon, were analyzed to reconstruct a 17,000-yr-long environmental history of high-elevation forests in the region. In the late-glacial period, the presence of a subalpine parkland of Artemisia, Poaceae, Pinus, and Tsuga with infrequent fires suggests cool dry conditions. After 14,500 cal yr B.P., a closed forest of Abies, Pseudotsuga, Tsuga, and Alnus rubra with more frequent fires developed which indicates more mesic conditions than before. An open woodland of Pinus, Quercus, and Cupressaceae, with higher fire activity than before, characterized the early Holocene and implies warmer and drier conditions than at present. In the late Holocene, Abies and Picea were more prevalent in the forest, suggesting a return to cool wet conditions, although fire-episode frequency remained relatively high. The modern forest of Abies and Pseudotsuga and the present-day fire regime developed ca. 2100 cal yr B.P. and indicates that conditions had become slightly drier than before. Sub-millennial-scale fluctuations in vegetation and fire activity suggest climatic variations during the Younger Dryas interval and within the early Holocene period. The timing of vegetation changes in the Bolan Lake record is similar to that of other sites in the Pacific Northwest and Klamath region, and indicates that local vegetation communities were responding to regional-scale climate changes. The record implies that climate-driven millennial- to centennial-scale vegetation and fire change should be considered when explaining the high floristic diversity observed at present in the Siskiyou Mountains.  相似文献   

20.
Measured lithostratigraphic sections of the classic Permian–Triassic non-marine transitional sequences covering the upper Quanzijie, Wutonggou, Guodikeng and lower Jiucaiyuan Formations at Dalongkou and Lucaogou, Xinjiang Province, China are presented. These measured sections form the framework and reference sections for a range of multi-disciplinary studies of the P–T transition in this large ancient lake basin, including palynostratigraphy, vertebrate biostratigraphy, chemostratigraphy and magnetostratigraphy. The 121 m thick Wutonggou Formation at Dalongkou includes 12 sandstone units ranging in thickness from 0.5 to 10.5 m that represent cyclical coarse terrigenous input to the lake basin during the Late Permian. The rhythmically-bedded, mudstone-dominated Guodikeng Formation is 197 m and 209 m thick on the north and south limbs of the Dalongkou anticline, respectively, and 129 m thick at Lucaogou. Based on limited palynological data, the Permian–Triassic boundary was previously placed approximately 50 m below the top of this formation at Dalongkou. This boundary does not coincide with any mappable lithologic unit, such as the basal sandstones of the overlying Jiucaiyuan Formation, assigned to the Early Triassic. The presence of multiple organic δ13C-isotope excursions, mutant pollen, and multiple algal and conchostracan blooms in this formation, together with Late Permian palynomorphs, suggests that the Guodikeng Formation records multiple climatic perturbation signals representing environmental stress during the late Permian mass extinction interval. The overlap between the vertebrates Dicynodon and Lystrosaurus in the upper part of this formation, and the occurrence of late Permian spores and the latest Permian to earliest Triassic megaspore Otynisporites eotriassicus is consistent with a latest Permian age for at least part of the Guodikeng Formation. Palynostratigrahic placement of the Permian–Triassic boundary in the Junggar Basin remains problematic because key miospore taxa, such as Aratrisporites spp. are not present. Palynomorphs from the Guodikeng are assigned to two assemblages; the youngest, from the upper 100 m of the formation (and the overlying Jiucaiyuan Formation), contains both typical Permian elements and distinctive taxa that elsewhere are known from the Early Triassic of Canada, Greenland, Norway, and Russia. The latter include spores assigned to Pechorosporites disertus, Lundbladispora foveota, Naumovaspora striata, Decussatisporites mulstrigatus and Leptolepidites jonkerii. While the presence of Devonian and Carboniferous spores and Early Permian pollen demonstrate reworking is occurring in the Guodikeng assemblages, the sometimes common occurrence of Scutasporites sp. cf. Scutasporites unicus, and other pollen, suggests that the Late Permian elements are in place, and that the upper assemblage derives from a genuine transitional flora of Early Triassic aspect. In the Junggar Basin, biostratigraphic data and magnetostratigraphic data indicate that the Permian–Triassic boundary (GSSP Level) is in the middle to upper Guodikeng Formation and perhaps as high as the formational contact with the overlying Jiucaiyuan Formation.  相似文献   

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