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1.
Benjamin P. Kear Thomas H. Rich Mohammed A. Ali Yahya A. Al-Mufarrih Adel H. Matiri Abdu M. Al-Masary Yousry Attia 《Cretaceous Research》2009,30(5):1164-1168
Actinopterygian remains have been recovered from Upper Cretaceous (lower Campanian to lower Maastrichtian) marginal marine deposits of the Adaffa Formation in northwestern Saudi Arabia. The fossils comprise gars (Lepisosteidae), pachycormids (cf. Protosphyraena sp.), indeterminate pycnodontiforms, enchodontid teleosts (cf. Enchodus sp.) and other indeterminate Teleostei. This assemblage is significant because it includes a novel occurrence for the Middle East (Pachycormidae) together with taxa (Lepisosteidae, Pycnodontiformes, Enchodontidae) that have been previously recorded from Late Cretaceous faunas elsewhere in the Mediterranean Tethyan region. 相似文献
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This paper discusses a well-represented fossil record of cartilaginous fishes (Chondrichthyes) from southern South America. The recovered samples allow the recognition of three assemblages with chronostratigraphic and paleogeographic value: i) typical Maastrichtian sharks and rays with affinities to eastern Pacific fauna, including the taxa Ischyrhiza chilensis, Serratolamna serrata, Centrophoroides sp. associated to Carcharias sp., and Dasyatidae indet.; ii) a scarce reworked assemblage of Paleocene–Early Eocene age including the taxa Otodus obliquus and Megascyliorhinus cooperi; iii) a rich assemblage with reworked taxa of Early to Middle Eocene age, together with autochthonous deposited Middle to Late Eocene taxa with close affinities to paleoichthyofaunas recovered from the North Atlantic, represented by Carcharias ‘hopei’, Odontaspis winkleri, Carcharoides catticus, Macrorhizodus praecursor, Carcharocles auriculatus, Striatolamia sp., Striatolamia macrota, Hexanchus agassizi, Notorhynchus sp., Myliobatis sp., Abdounia sp., Pristiophorus sp., Squatina sp., cf. Rhizoprionodon sp., Ischyodus sp., and one new species, Jaekelotodus bagualensis sp. nov. The studied samples include for the first time taxa with well established chronostratigraphic resolutions as well as taphonomic information that help clarifying the age of the fossil-bearing units. In addition, they provide relevant information about the evolution of the Magallanes (=Austral) Basin from the Upper Cretaceous to the Paleogene, suggesting a probable connection with the Quiriquina Basin of south-central Chile during the latest Cretaceous. Finally, the studied assemblages indicate a latitudinal pattern of distribution that provides valuable data on the environmental evolution and temperature of southern South America during the Paleogene. 相似文献
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The North American fossil record of dinosaur eggshells for the Cretaceous is primarily restricted to formations of the middle (Albian–Cenomanian) and uppermost (Campanian–Maastrichtian) stages, with a large gap in the record for intermediate stages. Here we describe a dinosaur eggshell assemblage from a formation that represents an intermediate and poorly fossiliferous stage of the Upper Cretaceous, the Santonian Milk River Formation of southern Alberta, Canada. The Milk River eggshell assemblage contains five eggshell taxa: Continuoolithus, Porituberoolithus, Prismatoolithus, Spheroolithus, and Triprismatoolithus. These ootaxa are most similar to those reported from younger Campanian–Maastrichtian formations of the northern Western Interior than they are to ootaxa reported from older middle Cretaceous formations (i.e., predominantly Macroelongatoolithus). Characteristics of the Milk River ootaxa indicate that they are ascribable to at least one ornithopod and four small theropod species. The taxonomic affinity of the eggshell assemblage is consistent with the dinosaur fauna known based on isolated teeth and fragmentary skeletal remains from the formation, although most ornithischians and large theropods are not represented by eggshell. Relative to the Milk River Formation eggshell, similar oospecies occurring in younger Cretaceous deposits tend to be somewhat thicker, which may reflect an increase in body size of various dinosaur lineages during the Late Cretaceous. 相似文献
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Exposures of the Menuha Formation (Santonian–Early Campanian, Mount Scopus Group) in the Makhtesh Ramon region of the southern Negev have produced numerous chondrichthyan teeth. The isolated teeth represent at least ten different species: Cretalamna appendiculata, Cretoxyrhina mantelli, Squalicorax falcatus?, S. kaupi, Scapanorhynchus rapax, S. raphiodon?, Carcharias samhammeri, Carcharias cf. C. holmdelensis?, and two other fish (Hadrodus priscus and a pycnodont). This assemblage has important implications for Late Cretaceous chondrichthyan palaeobiogeography. The majority of teeth were contained within a glauconite-rich, yellow-brown, soft chalk that included oysters (Pycnodonte vesicularis?), trace fossils (Planolites, Thalassinoides, and Chondrites), phosphatic peloids, and foraminiferans (globigerinids). The teeth were collected mainly through surface-sampling and sieving. The Menuha Formation probably represents a temperate to subtropical, shallow, open-shelf environment deposited during the formation of the Ramon anticline. Reworked conglomeratic chalks in the western section represent marginal facies derived from this structural uplift. With little to no published material describing the chondrichthyan fauna of the Menuha Formation, these data improve interpretations of its palaeoenvironment. Interpretation of the palaeoenvironment of the formation is important for understanding the larger stratigraphic/tectonic framework of the Ramon monocline region of southern Israel. 相似文献
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The Coll de Nargó area (Lleida Province, south-central Pyrenees) has yielded thousands of dinosaur eggs distributed in more than 30 levels across 370 m of Upper Cretaceous Tremp Formation deposits. Two stratigraphic units yielding dinosaur eggshells, eggs and clutches, can be distinguished. Multiple eggshells were collected from up to 75 different in situ clutches, located in consecutive stratigraphical levels throughout the Mas de Pinyes section. Four different oospecies have been recognized: Megaloolithus aureliensis, Megaloolithus siruguei, Megaloolithus cf. baghensis and Cairanoolithus cf. roussetensis. In absence of typical biochronological markers, fossil eggshells may have potential as a tool for dating. Based on these ootaxa, three oozones can be established along the stratigraphic section. The lower assemblage is composed by M. aureliensis, M. siruguei and Cairanoolithus cf. roussetensis; the middle one consists only of M. siruguei, while the upper oozone is characterized by M. cf. baghensis. Results suggest that the age of Coll de Nargó rocks ranges from the latest Campanian to the beginning of the Late Maastrichtian. Also reported is the first evidence of the oogenus Cairanoolithus outside of France, where it was regarded as an endemic ootaxon. The Coll de Nargó area is considered to be one of the most important dinosaur nesting areas in Europe recurrently used by several dinosaurs during a long time span. Finally, the presence of fossilized eggshell membrane in several specimens from Coll de Nargó contributes to understanding the taphonomic process of megaloolithid eggs in this area. 相似文献
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Upper Maastrichtian deposits formed in a nearshore subtidal environment within the Valdenoceda Formation (Castilian Ramp, North Iberian margin) are described together with two recently found selachian assemblages. Rare earth element concentrations (REE) have been used to assess the degree of taphonomic mixing and reworking, observing that it is minor or non-existent, and differences in degree of preservation and ecologic mixing can be explained by biostratinomic processes. The patterns of REE also helped to obtain a better understanding of the depositional environment, including the diagenetic history from burial to final degree of bone preservation.The fossil assemblages here described are close to that of the late Maastrichtian of Albaina (in the enclave of Condado de Treviño, Burgos), both in the Basque-Cantabrian Region, but their age may be slightly older (early late Maastrichtian). In total, the new assemblages consist of 17 taxa, assigned to 11 genera of shallow-water dwellers combined with individuals from the outer shelf. They represent cosmopolitan taxa (Squalicorax pristodontus, Serratolamna serrata and Rhombodus binkhorsti) together with local species (Rhinobatos echavei, Rhinobatos ibericus). Although there are not significant differences between Albaina and Quintanilla la Ojada faunas, the new assemblages add interesting taphonomic and geochemical information to the few existing uppermost Cretaceous deposits with fossil sharks in southwestern Europe. 相似文献
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Nineteen benthonic and planktonic foraminiferal zones and their subzones have been recognized in the Tethyan cretaceous successions along the four sections analyzed in the northwestern Zagros fold–thrust belt within the preforeland–foreland basin. A detailed micropaleontological investigation revealed eight benthonic zones from the Qamchuqa Formation (Barremian to Lower Early Cenomanian) including: the Choffatella decipiens interval zone, C. decipiens/Palorbitolina lenticularis total range zone, C. decipiens/Salpingoporella dinarica interval zone, Mesorbitolina texana total range zone, Mesorbitolina subconcava total range zone, Orbitolina qatarica total range zone, Orbitolina sefini total range zone, and the Orbitolina concava partial range zone. The Rotalipora cushmani total range zone was recorded in the Dokan Formation that overlies the Qamchuqa Formation of the Late Cenomanian age. The Gulneri Formation is represented only by the Whitnella archaeocretacea partial range zone/Heterohelix moremani total range subzone and indicates the Late Cenomanian/Early Turonian age. Six planktonic foraminiferal zones were recorded from the Kometan Formation, indicating the Late Cenomanian to Early Campanian age, and are represented by the R. cushmani/H. moremani subzone, Helvetotruncana helvetica total range zone, Marginotruncana sigali partial range zone, Dicarinella primitiva interval range zone, Dicarinella concavata interval zone, Dicarinella assymetrica total range zone, and Globotruncanita elevata partial range zone. Two planktonic foraminferal zones were recorded also and these are related to the Globotruncana (fornicata, stuartiformis, elevata, and ventricosa) assemblage zone, Globotruncana calcarata total range subzone, from the Shiranish Formation, Lower Late Campanian, while the second zone is nominated as the Globotruncana (arca, tricarinata, esnehensis, and bahijae) assemblage zone, Globotruncana gansseri interval subzone, and Globotruncana contusa total range zone of the Late Campanian to basal middle Maastrichtian age. The last zone is related to the Abathomphalus mayaroensis partial range zone (of Late Maastrichtian age) and occasionally intercalated with the Orbitoides–Loftusia benthic zones. An important hiatus, between the Qamchuqa and Kometan formations was proved and manifests Pre-Aruma unconformity, and is occasionally associated with the global Cenomanian–Turonian Oceanic Anoxic Euxinic Event, while the Maastrichtian red bed of the Shiranish Formations mostly points to Tethyan upper Cretaceous Oceanic Red Bed. 相似文献
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The subject of the study was the taxonomy of the ammonite fauna from the Upper Barremian marls and marly limestones of the Donji Milanovac Formation outcropped at the Boljetin Hill (Danubicum Unit). These sediments yielded a rich ammonite fauna which included also representatives of two superfamilies, Desmoceratoidea and Silesitoidea. The Desmoceratoidea include the family Barremitidae to which belong Plesiospitidiscus boljetinensis n. sp., Barremites balkanicus, Montanesiceras breskovskii n. sp., Barremitites strettostoma strettostoma and B. panae, Torcapella serbiensis n. sp., Pseudohaploceras tachthaliae, P. portaeferreae, Melchiorites haugi and Patruliusiceras cf. crenelatum. The Silesitoidea are represented by the family Silesitidae with Silesites trajani and S. seranonis. With the exception of the new, possibly endemic species, Plesiospitidiscus boljetinensis n. sp., Montanesiceras breskovskii n. sp., and Torcapella serbiensis n. sp., these taxa are common in the Tethyan regions. The studied deposits with ammonites belong to the lower part of the Late Barremian, in particular to the upper part of the Toxancyloceras vandenheckii ammonite Zone up to the lower part of the Imerites giraudi Zone. 相似文献
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In this paper we describe previously unpublished trionychid turtle material, consisting of numerous shell fragments, from the Late Cretaceous (late Turonian) Bissekty Formation of the Dzharakuduk locality in Uzbekistan. This material is assigned to two shell-based taxa: Aspideretoides cf. riabinini and “Trionyx” cf. kansaiensis. The material which cannot be confidently attributed to these two taxa is identified as Trionychidae indet. In addition to these shell-based trionychid taxa, the Dzharakuduk turtle assemblage includes two skull-based taxa of trionychids (Khunnuchelys kizylkumensis and Trionychini indet.). The trionychids from the Bissekty Formation are most similar to trionychids from the younger (Santonian – early Campanian) Bostobe Formation of Kazakhstan, represented by three shell-based taxa (Aspideretoides riabinini, Paleotrionyx riabinini and “Trionyx” kansaiensis), and one skull-based taxon (Khunnuchelys sp.). We provide an improved understanding of the subtle similarities and differences between four closely related Cretaceous turtle assemblages of Middle Asia and Kazakhstan. 相似文献
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A small brachiopod fauna is described from the upper part of the Yenduyet Formation near Son La, northwest Vietnam. The fauna includes the following species: Peltichia kwangtungensis (Zhan), Acosarina minuta (Abich), Rhipidomella hessensis King, Schuchertella cf. cooperi Grant, Derbyia sp., Waagenites soochowensis (Chao), Spinomarginifera chenyaoyanensis Huang, and a Marginiferinae gen. and sp. indet. The fauna is most closely correlated with brachiopods from the lower Changhsingian Stage of South China and the lyttoniid fauna from the Huai Tak Formation of northwest Thailand. In view of these correlations, an early Changhsingian (Late Permian) age is proposed for the Son La brachiopod assemblage. 相似文献
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The recently described clade Allodaposuchidae includes European eusuchian crocodyliforms restricted to the Late Cretaceous (Campanian and Maastrichtian). A new allodaposuchid crocodyliform is here described based on two specimens from the upper Campanian–lower Maastrichtian fossil site of Lo Hueco (Cuenca, Spain). This new taxon, Agaresuchus fontisensis gen. et sp. nov., is described by two complete skulls and a lower jaw associated with one of them. This new species can be distinguished unambiguously from Lohuecosuchus megadontos, the other allodaposuchid known from the same fossil site. The presence of two allodaposuchid crocodyliforms in Lo Hueco allows the recognition of the synchronic and sympatric existence of two representatives of this clade for the first time. The new genus Agaresuchus, comprises a previously described Iberian allodaposuchid species, “Allodaposuchus” subjuniperus, as Agaresuchus subjuniperus, new combination. 相似文献
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Martin A. Becker John A. Chamberlain Jr. Albert J. Robb Dennis O. Terry Jr. Matthew P. Garb 《Cretaceous Research》2009,30(4):1031-1040
The Fairpoint Member of the Fox Hills Formation (upper Maastrichtian) in Meade County, South Dakota, USA, contains an osteichthyan assemblage indicative of transitional to marine shoreface deposits. The fauna consists of: Lepisosteus sp., Paralbula casei, Cylindracanthus cf. C. ornatus, Enchodus gladiolus, Hadrodus sp., and indeterminate osteichthyans with probable affinities to the Siluriformes and Beryciformes. The Fairpoint fauna is of limited species diversity and in this character mirrors many other Upper Cretaceous North American osteichthyan assemblages. Comparison to Upper Cretaceous chondrichthyan diversity and consideration of the structure of Cretaceous marine food webs suggest that osteichthyans are strongly under-represented in the Upper Cretaceous of North America. The small size and poor preservation potential of many Upper Cretaceous North American osteichthyans probably account for much of this observed paucity. Fairpoint osteichthyans are members of families that survive the Cretaceous–Paleocene boundary extinction event. Some of these genera and families are still extant and occur in a wide array of modern fresh, brackish, and shallow marine environments. 相似文献
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The Cape Lamb Member of the Snow Hill Island Formation at The Naze on the northern margin of James Ross Island, east of the Antarctic Peninsula, yielded a theropod dinosaur recovered near the middle of a 90 m thick section that begins at sea level, ends below a basalt sill, and is composed of interbedded green–gray massive and laminated fine-grained sandstones and mudstones. Sixteen palynoassemblages were recovered from this section, which yielded moderately diverse assemblages with a total of 100 relatively well-preserved species. The principal terrestrial groups (32%) are represented by lycophytes (8 species), pteridophytes (15 species), gymnosperms (13 species), angiosperms (21 species) and freshwater chlorococcaleans (3 species). Marine palynomorphs (68%) belong to dinoflagellates (61 species), chlorococcaleans (6 species), and one acritarch. The vertical distribution of selected species allows the distinction of two informal assemblages, the lower Odontochitina porifera assemblage from the base to its disappearance in the lower part of the section, and the remaining section characterized by the Batiacasphaera grandis assemblage. The global stratigraphic ranges of selected palynomorphs suggest an early Maastrichtian age for this section and the entombed dinosaur that is also supported by the presence of the ammonoid Kitchinites darwinii. These assemblages share many species with latest Campanian–early Maastrichtian palynofloras from Vega and Humps Islands, New Zealand, and elsewhere in the Southern Ocean, establishing a good correlation among them. The dominance or frequent presence of dinoflagellates throughout the section supports the general interpretation of a shelf marine depocenter. The consistent presence of terrestrial palynomorphs suggests contributions from littoral/inland environments. 相似文献
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保山西邑铅锌矿区香山组一段灰岩牙形石丰富,包括5属7种1个相似种及2个未定种,分别为:Ancyrodella curvata,A.nodosa,A.spp.,Ancyrognathus triangularis,Icriodus alternatus alternatus,I.sp.,Palmatolepis hassi,Pa.simpla,Polygnathus webbi,Po.cf.tenellus和Po.sp.。该牙形石组合面貌可与上泥盆统Late Palmatolepis rhenana带对比,表明研究区内香山组一段的时代为晚泥盆世弗拉斯期晚期,并非早石炭世。 相似文献
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Mahboobeh FEREYDOONPOUR Hossein VAZIRI-MOGHADDAM and Azizollah TAHERI 《《地质学报》英文版》2014,88(6):1681-1695
Abstract: A rich assemblage of planktonic foraminifera has been studied from an outcrop of the Gurpi Formation, the hydrocarbon source rock in the southwest Iran, Deh Dasht area (Kuh-e Siah anticline). Based on the distribution of the planktonic foraminifera, eight biozones have been recognized that included: Dicarinella concavata Interval Zone (Earliest Santonian), Dicarinella asymetrica Total Range Zone (Santonian to Earliest Campanian), Globotruncanita elevata Partial Range Zone (Early Campanian), Globotruncana ventricosa Interval Zone (Middle to Late Campanian), Radotruncana calcarata Total Range Zone (Late Campanian), Globotruncanella havanensis Partial Range Zone (Late Campanian), Globotruncana aegyptiaca Interval Zone (Late to latest Campanian), Gansserina gansseri Interval Zone (Latest Campanian to Early Maastrichtian). These biozones indicates that the Gurpi Formation deposited during the Early Santonian- Early Maastrichtian. These biozones are compared to the most standard biozones defined in Tethysian domain. Based on distribution of morphotype groups of planktonic foraminifera, planktonic to benthic ratio (P/B) and content of carbonate, nine third-order sequences are recognized. 相似文献
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The uppermost Cretaceous (upper Campanian–Maastrichtian) marine deposits of the central south Pyrenees host a rich larger benthic foraminiferal fauna and several rudist-rich levels. These marine deposits are directly overlain by the continental facies of the Arén and Tremp Formations, which are famous for their fossil dinosaur remains. Larger benthic foraminiferal distribution documents an important faunal turnover in all the carbonate platform environments within the photic zone, from open marine to littoral areas. Biostratigraphy indicates that this turnover occurred close to the Campanian-Maastrichtian boundary. This is also confirmed by strontium isotope stratigraphy which indicates an earliest Maastrichtian age for the appearance of the larger benthic foraminiferal assemblage constituted by Lepidorbitoides socialis, Clypeorbis mammillata, Wannierina cataluniensis, Orbitoides gruenbachensis, Siderolites aff. calcitrapoides, Fascispira colomi, Omphalocyclus macroporus and Laffiteina mengaudi. In particular, a numerical age of 71 Ma is obtained for the Hippurites radiosus level, just a few meters below the first continental deposits of the Arén sensu stricto Formation. The youngest marine sediments of the central south Pyrenees are early Maastrichtian in age. This is also an important constraint for the age of the end-Cretaceous dinosaur fossil localities of the Tremp basin. 相似文献
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A symphyseal region of the fused dentaries of a caenagnathid theropod is described from the Upper Cretaceous Nemegt Formation at the Bugin Tsav locality in the Mongolian Gobi Desert. In contrast to the high diversity of Caenagnathidae in the upper Campanian to Maastrichtian in North America, only specimens of a single caenagnathid, Elmisaurus raurus, have been reported in the coeval strata in Asia. Although dentaries are commonly-found bones in the fossil record of Caenagnathidae, the present specimen is the first discovery of caenagnathid dentaries from the upper Campanian to Maastrichtian in Asia. The Nemegt Formation is unique for its diverse oviraptorosaurian fauna that includes both Caenagnathidae and Oviraptoridae as well as the non-caenagnathoid Avimimus portentosus. Hypothesized coexistence of eolian and fluvial environments in the Gobi Basin during the deposition of the Nemegt Formation might explain such co-occurrence of Caenagnathidae and Oviraptoridae. 相似文献
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An Upper Maastrichtian horizon rich in Loftusia is for the first time described in situ in Greece in the province of Boeotia. It is found in a continuous undisturbed carbonate sequence of the eastern Greece platform (Subpelagonian zone) of Maastrichtian–Paleocene age followed by flysch sedimentation. Loftusia is found in a facies reflecting an outer shelf environment, associated with debris of rudists, Orbitoides spp., Siderolites calcitrapoides, Omphalocyclus macroporus, Hellenocyclina beotica, Sulcoperculina sp., and echinoderms. Similar fauna, but without Loftusia, is found in the surrounding levels of Late Maastrichtian age as well. This recovery of the genus in the western part of its distribution area, where it is not as abundant as in the eastern part, is considered significant for the palaeobiogeography of the genus in Tethys Ocean during Late Cretaceous. To cite this article: A. Zambetakis-Lekkas, A. Kemeridou, C. R. Geoscience 338 (2006). 相似文献