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1.
The terrestrial lizard fossil record of Gondwana is very scarce. Few lizards, from mostly fragmentary fossils, have been identified in Madagascar, Tanzania, Morocco, South Africa, India and South America. Among the South American specimens there are basal Squamata forms (Olindalacerta and Tijubina) and a possible iguanid (Pristiguana) form Brazil. In Argentina gondwanian terrestrial lizards are represented by a putative iguanid and a small, poorly preserved dentary. This last specimen, recovered from the Anacleto Formation (Neuquén Group, Río Colorado Subgroup) in vicinity of the locality of Cinco Saltos (Río Negro Province), is here described formally. Comparisons with extant lizards indicate more affinity of the fossil with the Scincomorpha (non Scincophidia) than with any other group of lizards. This finding suggests that lizards were probably better represented and more diverse in the Mesozoic of South America than previously thought, although the fossil record is, at the moment, much poorer than in Laurasia.  相似文献   

2.
Although five genera of procyonids are currently present in South America, only two of the extant genera, Procyon and Nasua are represented in the South American fossil record. A recent discovery of a procyonid lower second molar in Late Pleistocene deposits of Aurora do Tocantins, northern Brazil, offers potential to further our understanding of the stratigraphic and temporal range of South American fossil procyonids. We use geometric morphometric analysis of two-dimensional landmarks and semilandmarks to explore morphological variation in the lower second molars of extant Procyon lotor and Procyon cancrivorus and multivariate methods to support the identification of the Pleistocene specimen as P. cancrivorus. This material represents the second fossil record of P. cancrivorus in South America Procyonids entered South America in two phases: the first comprising by Cyonasua and Chapadmalania during the Late Miocene, and the other recent genera, beginning in the Late Pleistocene. These Late Miocene procyonids were more carnivorous than Late Pleistocene-Recent omnivorous taxa and possible went extinct due to competition with other placental carnivorans that entered South America and diversified during the latest Pliocene-Early Pleistocene.  相似文献   

3.
Anza-Borrego Desert State Park® (ABDSP) has an exceptional proboscidean fossil record. The remains of Gomphotheriidae and Elephantidae span over 10 Ma and encompass over eighty productive sites. The most numerous of the proboscideans are the mammoths. Significant specimens include one of the youngest Gomphotherium and the most complete skeleton of Mammuthus meridionalis in North America.Over 6 km of fossiliferous sediments spans the Miocene–Pliocene and Pliocene–Pleistocene boundaries and provides a continuous record of changing environments. Over 550 taxa of aquatic and terrestrial plants, marine and lacustrine invertebrates, and marine, fresh water and terrestrial vertebrates are represented. Present are both Asian and South American immigrant mammals. M. meridionalis and M. columbi (=M. imperator) regionally co-existed here. A specimen from the Diablo Formation extends the temporal range of Gomphotherium in North America nearly one million years into the middle Blancan.  相似文献   

4.
Catfish bones from Tortonian (Miocene) freshwater beds of central Argentina are here identified as pertaining to a new species of the tropical pimelodid genus Phractocephalus. The new species differs from the other recent and fossil species of the genus in skull, pectoral girdle and spine characters. The material was found in different localities near the city of Paraná, Entre Ríos Province. The bearing horizon is the so-called “Conglomerado osífero” which constitutes the lowermost beds of the fluvial Ituzaingó Formation. The aquatic vertebrate fauna occurring in the bearing bed shows a similar generic composition to several northern South American Miocene units where Phractocephalus remains were found. This report extends the range of Phractocephalus more than 2000 km to the South. The record is in agreement with higher global temperatures and putative ample hydrographic connections of the river basins in the Paraná area with the Amazon basin until at least the early late Miocene.  相似文献   

5.
In 2007, Reguero et al. described a 2.5 Ma erethizontid dentary from the Uquía Formation in Argentina (MACN5376) and referred it to the genus Erethizon, a genus found at present only in North America. They based their generic identification on the presence in the fossil of a markedly inflected angular process. I argue in this commentary that this single trait (the angular process) in a single incomplete fossil is insufficient evidence for the Erethizon attribution, for the following reasons: 1. The trait (the inflected angular process) is variable in modern South and North American porcupines and observer bias and/or allometry may be responsible for observed differences; 2. Among fossil South American porcupines, the inflected angular process is not unique to the Uquían fossil; 3. The fossil possesses other traits (including one trait newly described in this paper) associated with modern South American (Coendou) porcupines and not modern Erethizon; and 4. The traits by which we recognize modern Erethizon, primarily related to the genus’s ability to survive severe winters, are most readily explained by evolution driven by the South American porcupine immigrants’ exposure in North America to that severe weather. The Uquían fossil is not Erethizon and should be attributed for the present to the genus Coendou.  相似文献   

6.
Integration of existing isotopic and geological data allows a reconsideration of the distribution and age of the Río de la Plata Craton within South America. The reinterpretation increases the area of the craton to about 2,400,000 km2 with implications for the tectonic map of South America and for global reconstruction of palaeocontinents. Four areas previously considered as separate cratons (Luís Alves, Curitiba, Tebicuary, and Paranapanema) are interpreted as part of the same Río de la Plata Craton. The craton is organized into six provinces and domains: Buenos Aires–Piedra Alta, Taquarembó, Tebicuary, Luís Alves, Encantadas, and Nico Pérez. The term ‘Transplatense’ is proposed to replace ‘Trans-Amazonian’ for Rhyacian events that occurred within the Río de la Plata Craton. The craton is formed not only by dominant Rhyacian rocks and local Archaean rocks, but also by Statherian and Mesoproterozoic rocks. The domains are all partially to totally covered by Phanerozoic basins (Paraná, Chacoparanense, Claromecó, Salado, Balcarce, and Colorado) which makes their investigation difficult. The Ediacaran–Cambrian collisions of the Brasilian orogen generated tectonic mixtures of orogenic and cratonic zones. This is more evident in the eastern margin of the craton, which behaved as a metacraton.  相似文献   

7.
Between January and September of 1887 Carlos Ameghino carried out his first geologic and paleontological expedition to the Río Santa Cruz, Patagonia. Based on the fossils and geologic information compiled, in 1887 and 1889, Florentino Ameghino named more than 120 new species of extinct mammals and his Formación Santacruceña and Piso Santacruceño (Santacrucian stage). Data published by both brothers state that the specimens were collected in outcrops by the Río Santa Cruz, between 90 and 200 km west of its mouth. However, information in the posthumously published letters and Travel Diary of C. Ameghino allows us to recognize a fourth locality, Río Bote, at about 50 km further southwest. In 1900, 1902, F. Ameghino divided the Piso Santacruceño in a younger étage Santacruzienne and older étage Notohippidéen, restricting the geographical distribution of the latter to Kar Aiken locality, northeast of Lago Argentino. However, 15 of the 54 species that F. Ameghino listed as exclusively Notohippidian stage already had been named on specimens collected South to the Río Santa Cruz in 1887, two year prior to C. Ameghino's first visit to Kar Aiken. Based on historical information and several expeditions to the Río Santa Cruz and its environs, in this contribution we establish the geographical locations of the 1887 localities, formalize their names, evaluate the stratigraphic position of the fossil-bearing levels, and analyze the geographic extension of the Notohippidian, inferring that Río Bote is where C. Ameghino first collected species that came to define the Notohippidian.  相似文献   

8.
A new Ar/Ar date on the Las Flores Tuff (Río Chico Group, Las Flores Fm., central Patagonia, Argentina) yielded an age of 49.512 ± 0.019 Ma. This tuff, which stratigraphically overlies the mammal-bearing deposits that produced the Las Flores fauna, helps constrain the age of the Itaboraian SALMA [South American Land Mammal Age] to which that fauna is referred. The new data also have implications for the age of succeeding mammal biochrons, such as the Riochican and “Sapoan” which are revised to being somewhat younger than previously interpreted. Although closer in age than formerly interpreted, they still are biotically distinct. Concomitant evaluations suggest that the Itaboraian SALMA is perhaps more contemporary with the EECO (Early Eocene Climatic Optimum) than previously considered. The Riochican may be interpreted as post-EECO, with its cooler climate consistent in that regard. A recent reconsideration of the chronology of elements of the Salamanca Formation resulted in the downward revision of the ages of the Peligran SALMA and the Carodnia Zone biochrons. These operations, together with our results, reflect a 9 m.y. gap in the late Paleocene and early Eocene land mammal record in South America.  相似文献   

9.
Knowledge of the main aspects of the Great American Biotic Interchange (GABI) concerning the glyptodontine Glyptodontidae (Xenarthra) is very scarce. A bidirectional dispersal process was recently proposed for this clade, with the presence of the North American genus Glyptotherium Osborn recognized in latest Pleistocene sediments of northern South America (Venezuela and Brazil). However, the earliest stages of this paleobiogeographical process remain poorly understood, mainly because of the limited fossil record on this clade in late Pliocene sediments. The goals of this contribution are: a) to present and describe the first record of a glyptodontine glyptodontid from the late Pliocene of northern South America, tentatively assigned to a new species of Boreostemma Carlini et al. (Boreostemma? sp. nov); and b) to analyze its paleobiogeographical implications with respect to the GABI. This new material was recovered from the San Gregorio Formation (late Pliocene, prior the GABI) in northern Venezuela, where it is represented by several osteoderms of the dorsal carapace. A comparison among the three known late Pliocene glyptodontine glyptodontids of a) southern South America (Paraglyptodon), b) northern South America (Boreostemma), and c) southern North America ("Glyptotherium"), reveals a series of shared characters between (b) and (c), not present in (a). The most important of these shared characters in (b) and (c) are: all the osteoderms present a great development of the central figure, which is always larger than the peripherals; the sulcus that delimits the central and peripheral figures is narrower and shallower; and all the osteoderms present are relatively thin. This evidence suggests that the lineage of Glyptodontinae which participated in the GABI and subsequently diversified in North America originated in northern South America. Moreover, the evident morphological differences between these glyptodontines with respect to the southern South American forms show a significant separation of both lineages since at least latest Miocene-early Pliocene.  相似文献   

10.
Sedimentological, ichnological and paleontological analyses of the Early Miocene uppermost Monte León Formation and the lower part of the Santa Cruz Formation were carried out in Rincón del Buque (RDB), a fossiliferous locality north of Río Coyle in Santa Cruz Province, Patagonia, Argentina. This locality is of special importance because it contains the basal contact between the Monte Léon (MLF) and the Santa Cruz (SCF) formations and because it preserves a rich fossil assemblage of marine invertebrates and marine trace fossils, and terrestrial vertebrates and plants, which has not been extensively studied. A ∼90 m-thick section of the MLF and the SCF that crops out at RDB was selected for this study. Eleven facies associations (FA) are described, which are, from base to top: subtidal–intertidal deposits with Crassotrea orbignyi and bioturbation of the Skolithos-Cruziana ichnofacies (FA1); tidal creek deposits with terrestrial fossil mammals and Ophiomorpha isp. burrows (FA2); tidal flat deposits with Glossifungites ichnofacies (FA3); deposits of tidal channels (FA4) and tidal sand flats (FA5) both with and impoverish Skolithos ichnofacies associated; marsh deposits (FA6); tidal point bar deposits recording a depauperate mixture of both the Skolithos and Cruziana ichnofacies (FA7); fluvial channel deposits (FA8); fluvial point bar deposits (FA9); floodplain deposits (FA10); and pyroclastic and volcaniclastic deposits of the floodplain where terrestrial fossil mammal remains occur (FA11).The transition of the MLF–SCF at RDB reflects a changing depositional environment from the outer part of an estuary (FA1) through the central (FA2–6) to inner part of a tide-dominated estuary (FA7). Finally a fluvial system occurs with single channels of relatively low energy and low sinuosity enclosed by a broad, low-energy floodplain dominated by partially edaphized ash-fall, sheet-flood, and overbank deposits (FA8–11). Pyroclastic and volcaniclastic materials throughout the succession must have been deposited as ash-fall distal facies in a fluvial setting and also were carried by fluvial streams and redeposited in both estuarine and fluvial settings. These materials preserve most of the analyzed terrestrial fossil mammals that characterize the Santacrucian age of the RDB's succession. Episodic sedimentation under volcanic influence, high sedimentation rates and a relatively warm and seasonal climate are inferred for the MLF and SCF section.Lateral continuity of the marker horizons at RDB serve for correlation with other coastal localities such as the lower part of the coastal SCF south of Río Coyle (∼17.6–17.4 Ma) belonging to the Estancia La Costa Member of the SCF.  相似文献   

11.
In this note we report new avian remains from the Late Cretaceous Los Alamitos Formation (Campanian-Maastrichtian) at the Los Alamitos locality, Río Negro Province, Argentina. Isolated remains referable to indeterminate Aves, ?Patagopterygiformes, indeterminate Ornithurae, cf. Hesperornithes and cf. Neornithes are described and discussed. The new genus and species Alamitornis minutus is erected to include a minute-sized and gracile bird, probably related to the non-volant ratite-like bird Patagopteryx. If correctly identified, the record of Hesperornithes may be the first for this group in the Southern Hemisphere. The Los Alamitos paleoavifauna represents one of the most diverse fossil bird assemblage from the Mesozoic of Gondwana known to date.  相似文献   

12.
The freshwater ferns (Salviniales) are well represented in the Maastrichtian deposits of Cerro de los Fragmentos in the headwaters of the Río Chico, Golfo San Jorge Basin. The fossil material of Salviniaceae includes complete megaspore apparatuses with attached microspore massulae and dispersed megaspores, float systems and microspore massulae of Azolla. The new species Azolla colhuehuapensis displays distinctive morphological characters including relative small megaspore apparatuses with easily detached floats and usually attached eglochidiate massulae. The Marsileaceae are represented by the megaspore Molaspora lobata, microspores of Crybelosporites and vegetative remains of Marsileaceaephyllum sp. The assemblage also includes zygospores of the green filamentous algae Zygnemataceae, spores of Marchantiophyta and Monilophyta (Schizaeales), megaspores of Isoetalean affinity and Palm-type angiosperm pollen grains. All these organic microfossils and the associated palynomorphs indicate the presence of a freshwater environment where abundant water ferns were developing and reproducing.  相似文献   

13.
In the 80's, Leonardi treated the presence of a vertebrate ichnological locality from the Barremian Corda Formation, Parnaíba Basin, on the left bank of the Tocantins river, near of the São Domingos town, Itaguatins, State of Tocantins, Brazil. Originally, the record was composed of at least seven in situ trackways, accounting for fifty six tracks. Since 2011, the Hydroelectric Power Plant do Estreito has begun to work, causing the development of a water reservoir 160 km upstream to the ichnological site, causing periodic and highly energetic floods over the footprints-bearing level and altering it. The imprints are poorly to moderate preserved, but it is possible to distinguish the general morphology and the spatial arrangement of the footprints. The specimens are represented by pes imprints, mostly circular to subcircular, with no digital and claw impressions. No distinguishable manus imprints are present. The trackways are relative narrow with respect to the size of the tracks, so they are considered into the Parabrontopodus-like category. The São Domingos tracks have been originally assigned to iguanodontid dinosaurs, and posteriorly related to a sauropodian origin. This idea is herein accepted, particularly to a basal sauropod, basal macronarians, or diplodocoids. Up to date, the tracks from the São Domingos locality are the only vertebrate fossil record from the Corda Formation, meaning an important contribution to the Cretaceous ichnofauna from South America.  相似文献   

14.
Aerial photography and satellite imagery reveal manifold geomorphological evidence of a dynamic evolution of past and present rivers in the Bolivian Amazon. Comparison of oxbow lake and meander scar dimensions along an inactive meander belt of the Río Mamoré (Llanos de Moxos, north-eastern Bolivia) and its modern counterpart suggests significant regional paleohydrological variability. We used these features as geomorphological and sedimentary archives to enhance our understanding of longer-term variations of the fluvial system. Late Pleistocene to Holocene hydrological changes of the Río Mamoré are inferred from: (i) the analysis of satellite imagery, (ii) discharge estimates from meander morphology, (iii) stratigraphic, and (iv) chronological information based on luminescence and radiocarbon dating. The combined data from three oxbows indicate that the now abandoned meander belt – the paleo-Mamoré – continued to be active at least until ∼5 ka, and likely even postdating 3 ka. An up to threefold increase in discharge is estimated for the modern Río Mamoré versus the paleo-Mamoré. The altered runoff regime may have triggered an avulsive shift towards the currently active Río Mamoré. The preceding increase in discharge in turn, was possibly related to a shift in climatic conditions, which changed markedly between the mid- and late Holocene in tropical South America. In addition, it may have been the indirect result of capturing the avulsive Río Grande system to the east of the Río Mamoré. Alternative explanations for the differences in dimensions of the paleo versus the modern Río Mamoré, i.e. contemporaneous activity of both rivers or alteration of site factors such as the channel/floodplain relationship, are considered to be unlikely.  相似文献   

15.
A middle Eocene Crassostrea sp. reef near Río Turbio, southwestern Patagonia (Argentina), represents the earliest record of an oyster reef associated with estuarine facies in the southern hemisphere, and also one of the few known worldwide occurring in Paleogene rocks. The reef grew in an outer estuary environment subject to periodic changes in salinity and may have reached a maturing phase. The Río Turbio reef – by its dimensions, geometry, and substrate lithology– would have been located in a tidal channel convergence area. This reef provides new evidence suggesting that estuaries served as refuges for Crassostrea populations allowing them to disperse into fully marine environments many times throughout the Cenozoic.  相似文献   

16.
Upper Cretaceous deposits in Mongolia, Chinese Inner Mongolia, and, more recently, southern China, have yielded individually rich and taxonomically diverser lizard assemblages. Here we describe the remains of a new terrestrial lizard, Asprosaurus bibongriensis gen. et sp. nov., from the Upper Cretaceous of South Korea. It represents the first record of a Mesozoic lizard from the Korean Peninsula and, although incomplete, is exceptional in its very large size. Characters of the mandible support attribution to crown-group Anguimorpha, with the closest similarities being to monstersaurs, the group represented today by the venomous North American Beaded lizard and Gila monster, genus Heloderma. This group is well-represented in the Upper Cretaceous fossil record in of eastern Asia, and the remains of large monstersaurs have been recovered from several dinosaur egg localities, suggesting dietary preferences similar to those of the living genus. The new Korean lizard, recovered from the Boseong Bibong-ri Dinosaur Egg Site, fits the same pattern.  相似文献   

17.
The Sierra Nevada of southern Spain is a landscape with a rich biological and cultural heritage. The range was extensively glaciated during the late Pleistocene. However, the postglacial paleoecologic history of the highest range in southern Europe is nearly completely unknown. Here we use sediments from a small lake above present treeline – Laguna de Río Seco at 3020 m elevation – in a paleoecological study documenting over 11,500 calendar years of vegetation, fire and climate change, addressing ecological and paleoclimatic issues unique to this area through comparison with regional paleoecological sequences. The early record is dominated by Pinus pollen, with Betula, deciduous Quercus, and grasses, with an understory of shrubs. It is unlikely that pine trees grew around the lake, and fire was relatively unimportant at this site during this period. Aquatic microfossils indicate that the wettest conditions and highest lake levels at Laguna de Río Seco occurred before 7800 cal yr BP. This is in contrast to lower elevation sites, where wettest conditions occurred after ca 7800. Greater differences in early Holocene seasonal insolation may have translated to greater snowpack and subsequently higher lake levels at higher elevations, but not necessarily at lower elevations, where higher evaporation rates prevailed. With declining seasonality after ca 8000 cal yr BP, but continuing summer precipitation, lake levels at the highest elevation site remained high, but lake levels at lower elevation sites increased as evaporation rates declined. Drier conditions commenced regionally after ca 5700 cal yr BP, shown at Laguna de Río Seco by declines in wetland pollen, and increases in high elevation steppe shrubs common today (Juniperus, Artemisia, and others). The disappearance or decline of mesophytes, such as Betula from ca 4000 cal yr BP is part of a regional depletion in Mediterranean Spain and elsewhere in Europe from the mid to late Holocene. On the other hand, Castanea sativa increased in Laguna de Río Seco record after ca 4000 cal yr BP, and especially in post-Roman times, probably due to arboriculture. Though not as important at high than at low elevations, fire occurrence was elevated, particularly after ca 3700 years ago, in response to regional human population expansion. The local and regional impact of humans increased substantially after ca 2700 years ago, with the loss of Pinus forest within the mountain range, increases in evidence of pasturing herbivores around the lake, and Olea cultivation at lower elevations. Though human impact was not as extensive at high elevation as at lower elevation sites in southern Iberia, this record confirms that even remote sites were not free of direct human influence during the Holocene.  相似文献   

18.
《Gondwana Research》2003,6(1):65-77
A sedimentary succession included in the lower section of the Playa Hermosa Formation from the Playa Verde Basin, Uruguay, is described. Two facies associations, one mainly coarse- to medium-grained and other one fine-grained, have been defined (FA I-II). In the first one, breccias, conglomerates, sandstones and minor mudstones were deposited in a subaqueous depositional setting (proximal) suggesting slope instability and resedimentation processes. The second one contains diamictites, rhythmites, sandstones and mudstones and presents abundant evidence of soft-deformation, also interpreted to be deposited in a subaqueous environment (distal). Dropstones, clast layers, diamictites, rhythmites and varve-like deposits are interpreted as ice rafting processes generated during a glacial episode. This glacial-related succession constitutes the first record from the Varanger glaciation at the Río de la Plata Craton of the late Neoproterozoic age and also represents one of the oldest sedimentary records after the collision of the Río de la Plata and Kalahari Cratons. A combined interaction of extensional faulting and glaciation in a tectonically active basin with locally high subsidence rates, resulted in high rates of sedimentation and resedimentation processes. As a whole, the sedimentary succession sets a relevant datum to be used in future paleogeographic reconstructions of the Vendian glacial record in southern South America.  相似文献   

19.
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20.
Pipid frogs are fully aquatic frogs that today inhabit freshwater environments of southern continents on both sides of the Atlantic Ocean, with a fairly good fossil record dating back to the Cretaceous. Here I report on fossils from the Allen Formation (upper Campanian–lower Maastrichtian), Río Negro Province, Argentina, that are ascribed to a new genus and species of pipid. In order to assess the evolutionary relationships of the new taxon, which is represented by sphenethmoids, otic capsules, ilia, humeri, and vertebrae, cladistic analyses of a data matrix of 165 osteological characters scored for 36 taxa were performed. The results are congruent with previous hypotheses of pipoid interrelationships and consistently place the new taxon as part of the lineage today represented by the African xenopodines. Temporal calibration of the phylogenetic tree based on the fossil record imply that the origin and early diversification of crown-group Pipidae might have occurred during the Early Cretaceous, prior to the final breakup of western Gondwana. This study highlights the importance of including fossils, even fragmentary ones, directly in phylogenetic analyses in order to disentangling how, when, and where pipid frogs diversified.  相似文献   

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