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1.
Sediments from the Upper Cretaceous (Cenomanian-Turonian) Tropic Shale were deposited along the western margin of the Western Interior Seaway, in present-day southern Utah. Marine vertebrates from this formation include plesiosaurs, mosasaurs, bony fish, sharks, and turtles. They are concentrated in the lower portion of the Tropic Shale, mostly between Bentonites B and D. Study of the taphonomic condition of these vertebrates has contributed to an understanding of how they were preserved as well as a detailed paleoenvironment for the Tropic Shale. Physical factors played the dominant role in their preservation, with robust and durable skeletal elements, such as teeth and vertebrae, being most common within the shale. Isolated bones and teeth are also relatively common within the formation, while complete and nearly complete skeletons are more rare. Biological factors played a less dominant role, with no evidence of epifaunal or infaunal activity preserved with any of the skeletal remains. In addition, scavenging marks (both bite and gnaw marks) are relatively uncommon, typically only being found on more complete specimens. A signature of post-burial alteration can be recognized as low levels of abrasion, weathering, and compression and high levels of fracturing of vertebrate skeletal material. Slightly higher levels of abrasion and weathering occur to the west, closer to the ancient shoreline, suggesting some pre-burial alteration. The preservation of marine vertebrates in the Tropic Shale suggests a low energy marine environment with some weak bottom currents and low levels of benthic oxygen. The substrate ranged from soft and soupy to firm, with moderate sedimentation rates resulting in relatively rapid burial. 相似文献
2.
The phylostratigraphy, taphonomy and palaeoecology of the Late Cretaceous neoselachian Ptychodus of northern Germany appears to be facies related. Ptychodus is not present in lower Cenomanian shark-tooth-rich rocks. First P. oweni records seem to relate to middle Cenomanian strata. P. decurrens appears in the middle to upper Cenomanian mainly in non-coastal environments of the shallow marine carbonate ramp and swell facies which isolated teeth were found partly in giant ammonite scour troughs on the Northwestphalian-Lippe High submarine swell in the southern Pre-North Sea Basin. They are recorded rare in deeper basin black shales facies (upwelling influenced, OAE Event II). P. polygyrus seems to be restricted to upwelling influenced basin and deeper ramp facies mainly of the uppermost Cenomanian and basal lower Turonian (OAE II Event). P. mammillaris is mostly represented during the lower to middle Turonian in the inoceramid-rich ramp and the near shore greensand facies along the Münsterland Cretaceous Basin coast north of the Rhenish Massif mainland. Finally, P. latissimus is recorded by two new tooth sets and appears in the upper Turonian basin swell facies and the coastal greensands. Autochthonous post-Turonian Ptychodus remains are unrecorded in the Santonian–Campanian of Germany yet. Reworked material from Cenomanian/Turonian strata was found in early Santonian and middle Eocene shark-tooth-rich condensation beds. With the regression starting in the Coniacian, Ptychodus disappeared in at least the Münster Cretaceous Basin (NW-Germany), but remained present at least in North America in the Western Interior Seaway. The Cenomanian/Turonian Ptychodus species indicate a rapid neoselachian evolution within the marine transgression and global high stand. A correlation between inoceramid shell sizes, thicknesses and their increasing size during the Cenomanian and Turonian might explain the more robust and coarser ridged enamel surfaces in Ptychodus teeth, if Ptychodus is believed to have preyed on epifaunistic inoceramid bivalves. 相似文献
3.
A thin phosphate-granule conglomerate within the Upper Cretaceous (middle Campanian) Rattlesnake Mountain sandstone member of the Aguja Formation preserves a diverse chondrichthyan and osteichthyan fauna. This highly fossiliferous deposit (the ‘Ten Bits Microsite’) yielded about 5000 teeth, spines, and denticles in a small amount of matrix (c. 150 kg). About 30 identifiable species of sharks, rays, and bony fishes are recognized. Two of the three most abundant chondrichthyan species at Ten Bits (Scapanorhynchus texanus and Ischyrhiza mira) are also the most common species in other middle to late Campanian marine vertebrate faunas along the Gulf and Atlantic Coastal Plain. The myliobatiform rays Brachyrhizodus and Rhombodus that occur at Ten Bits also appear to be characteristic of the Gulf and Atlantic Coast, as are lamniform sharks such as Cretalamna and Serratolamna. These taxa are entirely absent or extremely rare in Western Interior Campanian faunas. In contrast, some small orectolobiform sharks (Cantioscyllium, Chiloscyllium, Columbusia) and small rays (Protoplatyrhina) found at Ten Bits are common in shallow water faunas of the Western Interior and Texas Coastal Plain, but rarely reported from the eastern Gulf or Atlantic Coast. The common Western Interior ray Myledaphus bipartitus does not occur at Ten Bits or in any Gulf or Atlantic Coast fauna. Ptychotrygon agujaensis is abundantly represented in the Ten Bits fauna, but unknown in correlative marine faunas. Although Ptychotrygon occurs in all Western Interior, Gulf and Atlantic Coastal Plain faunas, it is represented elsewhere by apparently endemic species at each collection site. The differences between Western Interior, Gulf, and Atlantic Coastal Plain faunas probably reflect latitudinal variation in water temperature or salinity, or different oceanic water circulation patterns between the Western Interior Seaway and the Gulf or Atlantic Coast that restricted the distributions of some marine fish species. The Ten Bits fauna shares typical species with both Western Interior and Gulf and Atlantic Coast faunas, reflecting its position at the border between these provinces; however, the dominant taxa found at Ten Bits are the same as those found on the Gulf and Atlantic Coast, and indicate that western Texas was more closely allied biogeographically with that province than with the Western Interior of North America. One species tentatively identified in the Ten Bits fauna on the basis of a single tooth, Igdabatis cf. I. indicus, is otherwise known only from southern Europe and Asia, although a similar large myliobatid ray also occurs rarely in Texas Coastal Plain faunas. These occurrences indicate that western Texas may have been near the northern limit of the range for some tropical Tethyan marine vertebrate species. 相似文献
4.
A historical collection of hesperornithiform fossils from the Gammon Ferruginous, Pembina, and Millwood members of the Pierre Shale (Campanian, Upper Cretaceous) in southern Manitoba, Canada, was examined to revise their taxonomy. Only two species of Hesperornis have been recognized in previous studies on the Pierre Shale in Manitoba, but our study recognizes six species of two genera, including H. lumgairi sp. nov. H. regalis is the most common species but absent in the uppermost unit within the studied sequence. The result of this study supports the paleobiogeographic subdivision of the Campanian vertebrate fauna within the Western Interior Seaway, but not the faunal boundary that distinguishes the avian fauna of Manitoba from that of South Dakota and Kansas. 相似文献
5.
A well-preserved right humerus collected from the upper Austin Group at Ejido Piedritas in Coahuila, Mexico is referable to the carinate bird Ichthyornis. This occurrence extends the known distribution of Ichthyornis to the southwest, and is among a younger (Coniacian-Campanian) group of specimens, recovered generally to the south of older (Cenomanian-Turonian) Ichthyornis specimens. The southward shift in occurrence of Ichthyornis may accord with the withdrawal of subtropical marine biota from the Western Interior Seaway of North America during the Late Cretaceous. 相似文献
6.
D.E. Hattin 《Cretaceous Research》1983,4(3):259-263
The Early Turonian ammonite, Vascoceras (Greenhornoceras) birchbyi Cobban & Scott, has been recorded at nine additional localities in Kansas, Colorado and New Mexico. Most significant is discovery of a specimen in Russell County, Kansas, that doubles the known longitudinal range of the species in the U.S. Western Interior. The species occurs in a single, thin, time-parallel limestone bed that apparently was deposited during a peak pulse of the Cenomanian-Turonian transgression. 相似文献
7.
ELANA L. LEITHOLD 《Sedimentology》1994,41(3):521-542
The Tropic Shale and correlative Tununk Shale Member of the Mancos Shale accumulated during Cenomanian-Turonian time, within prodeltaic environments near the western margin of the Western Interior Seaway of North America. Stratigraphical and sedimentological analysis has revealed a detailed history of relative sea level change in the thick, fine grained succession. The Tropic and Tununk shales were deposited during the Greenhorn second-order sea level cycle, over a time span of about 2–5 million years. In southern Utah, six depositional sequences are superposed upon the record of this long term sea level change. The sequences developed during third-order relative sea level cycles of hundreds of thousands of years duration and are composed of at least 37 parasequences, arranged in retrogradational, aggradational and progradational parasequence sets. The Tropic Shale and Tununk Shale Member accumulated just basinward of the axis of maximum subsidence of a foreland basin. Stratal geometries and facies distribution patterns in the succession indicate that in southern Utah the Greenhorn cycle was tectonically controlled. During the Greenhorn transgression and highstand, rapid rates of tectonic subsidence trapped terrigenous sediment to the west of the study area, in the more proximal foreland. At this time, hemipelagic facies accumulated at relatively slow rates in southern Utah and type 2 sequences developed during third-order sea level cycles. In contrast, during the Greenhorn regression rates of thrust-induced subsidence in the proximal foreland basin evidently slowed, and deltaic clinoforms prograded across the study area. At least one forced regression occurred in southern Utah at this time, and type 1 sequences developed. The formation of type 1 sequence boundaries in the upper part of the Tropic Shale and Tununk Shale Member points to episodes of base level fall and indicates that the six third-order sea level cycles recorded in the succession were not the result of changes in sediment supply alone. The third-order cycles may have been a consequence of episodic tectonism. The timing of these cycles, however, suggests that development of sequences and parasequences in the Tropic Shale and Tununk Shale Member may have been related to orbital forcing in the Milankovitch band. Glacioeustasy or climatically related fluctuations in the amount of groundwater stored on continents may explain these high frequency sea level changes. 相似文献
8.
《Proceedings of the Geologists' Association. Geologists' Association》2022,133(3):212-217
The cirripede Calantica? saskatchewanensis Russell, 1967, from the Maastrichtian Bearpaw Formation of Saskatchewan (Canada), is redescribed on the basis of new material and transferred to the genus Pachyscalpellum Buckeridge, 1991. This considerably extends the known range of the genus from the eastern Tethys (Australasia, Tunisia) to the northern part of the Western Interior Seaway of North America. Species of the genus achieved an exceptionally large size for stalked barnacles, with a projected possible height (capitulum and peduncle) of 150–200 mm. 相似文献
9.
Inoceramid bivalves of the upper Albian and lower Cenomanian of the United States Western Interior are revised, Eleven species-level taxa and three genera are described. Two new species, Gnesioceramus mowriensis, characterizing the Mowry Shale of the early, but not the earliest, Cenomanian, and Posidonioceramus merewetheri, of the lower Cenomanian, and on new genus, Posidonioceramus, are recognised. The Western Interior inoceramid species from this interval are strongly endemic and are not good tools for long-distance correlations, although they are very effective in regional dating.In terms of the inoceramid biostratigraphy, middle and upper parts of the upper Albian can be referred to the Gnesioceramus Biozone, represented by G. comancheanus (Cragin) and G. bellvuensis (Reeside). These taxa are endemic to the Western Interior and some adjacent areas (Gulf Coast; Greenland?), but are closely allied to the cosmopolitan species, Gnesioceramus anglicus (Woods). At approximately the Albian-Cenomanian boundary, the endemic clade of ‘Inoceramus’ nahwisi appears, now referred to the newly erected Posidonioceramus, resulting in a distinct P. nahwisi biozone. This zone corresponds to the lower part of the ammonite Neogastroplites’ stratigraphic range. Gnesioceramids re-appear in the early Cenomanian. Close to base of the Cenomanian, for the first in the Western Interior, the genus Inoceramus, represented by Inoceramus irenensis Warren and Stelck, 1958, apparently immigrated into the Western Interior Basin.The Western Interior inoceramids do not allow for direct correlation to chronostratigraphic standard subdivision. The Albian-Cenomanian boundary, as earlier recognized on geochronologic correlations and confirmed, to some extent, based on ammonites, may approximately be located close to the appearance level of the genus Posidonioceramus. 相似文献
10.
11.
Terry et al. (2001) proposed that the Fox Hills Formation in the area of Badlands National Park, southwestern South Dakota, USA, contains the Cretaceous/Paleogene (K/Pg) boundary, marked by a thick layer of contorted bedding called the Disturbed or Disrupted Zone (DZ). Examination of the ammonites from just below this layer yields Hoploscaphites nicolletii (Morton, 1842), H. spedeni (Landman and Waage, 1993), Discoscaphites gulosus (Morton, 1834), D. conradi (Morton, 1834), and Sphenodiscus lobatus (Tuomey, 1856). The abundance of Discoscaphites and the presence of a coarsely ornamented specimen of H. spedeni suggest that this assemblage corresponds to the upper part of the H. nicolletii Zone in the type area of the Fox Hills Formation. No ammonites are present above the DZ, but previous analyses of the dinoflagellates from just below, within, and above the DZ by Palamarczuk et al. (2004) are consistent with the ammonite results. Together, these fossils indicate that the interval just below and above the DZ represents the upper part of the lower upper Maastrichtian (≈ middle upper Maastichtian). Belemnites are present in the strata just below the DZ and occur as guards either isolated in the matrix or associated with fragmentary ammonites in concretions composed of soft sandy marl. The belemnites are assigned to Belemnitella bulbosa Meek and Hayden, 1857a, and B. badlandsensis n. sp., which is characterized by an unusually large fissure angle. The distribution of ammonites and lithofacies at this time reveals that the western shoreline of the Western Interior Seaway trended northeast–southwest across western South Dakota and adjacent parts of North Dakota. Based on an examination of the oxygen isotopes of the belemnites and scaphites in this area, the seawater temperature was nearly constant along the coast, approximately 17–20 °C. A comparison of the contact between the Pierre Shale and Fox Hills Formation at various localities in the Western Interior Basin indicates that this contact rises in the section toward the east, reflecting the final retreat of the Seaway during the late Maastrichtian. 相似文献
12.
We re-define the Cretaceous bony fish genus Rhinconichthys by re-describing the type species, R. taylori, and defining two new species; R. purgatorensis sp. nov. from the lowermost Carlile Shale (middle Turonian), southeastern Colorado, United States, and R. uyenoi sp. nov. from the Mikasa Formation (Cenomanian), Middle Yezo Group, Hokkaido, Japan. Rhinconichthys purgatoirensis sp. nov. is designated on a newly discovered specimen consisting of a nearly complete skull with pectoral elements. Only known previously by two Cenomanian age specimens from England and Japan, the North American specimen significantly extends the geographic and stratigraphic range of Rhinconichthys. The skull of Rhinconichthys is elongate, including an expansive gill basket, and estimated maximum body length ranges between 2.0 and 2.7 m. Rhinconichthys was likely an obligate suspension-feeder due to its derived cranial morphology, characterized by a remarkably large and elongate hyomandibula. The hyomandibula mechanically acts as a lever to thrust the jaw articulation and hyoid arch both ventrally and anterolaterally during protraction, thus creating a massive buccal space to maximize filtering of planktonic prey items. Cladistic analysis supports a monophyly of suspension-feeding pachycormids including Rhinconichthys, but further resolution within this clade will require more information through additional fossil specimens. 相似文献
13.
Thin-bedded and millimetrically laminated platy marly limestone quarried near Vallecillo, north-eastern Mexico, contains abundant excellently preserved marine fossils. Planktic foraminifers, inoceramids, and ammonites occur throughout the 7.7-m-thick section of this plattenkalk and provide a precise and detailed biostratigraphic zonation from the uppermost Cenomanian to early Turonian, with a mixed assemblage of Tethyan and Western Interior Seaway faunal elements. Five species of inoceramids are present and described herein: Inoceramus pictus pictus, Mytiloides hattini, M. puebloensis, M. goppelnensis, and M. kossmati. The faunal characteristics of the Vallecillo fossil assemblage combined with the monotonous lithology are favourable attributes for correlation with the GSSP at Pueblo, Colorado, and the Eastbourne section in southern England. The first appearances (FAs) of Watinoceras and Mammites nodosoides are considered approximately isochronous and thus suited for long-distance correlation. In contrast, the FAs of Pseudaspidoceras flexuosum, Fagesia catinus, and Helvetoglobotruncana helvetica are clearly diachronous. The range of M. kossmati needs further evaluation. 相似文献
14.
The Cenomanian–Turonian ammonite biostratigraphical framework for the southern Tethys margin (North Africa, Middle East and the Arabian Peninsula) is becoming better understood. A first attempt at a synthetic range chart is presented, with 85 taxa and precise correlations for ammonites along a west–east transect from Morocco to Oman, inclusive of the Trans-Saharan Seaway as far south as northern Nigeria. On the basis of a critical review of ammonite taxonomy, 13 bioevents can be identified in the interval from the Late Cenomanian to the Early Turonian (c. 3.5 myr) with each bioevent corresponding to a time interval of approximately 270,000 years, on average. They are consistent throughout several regions along the southern Tethys margin, though some gaps remain, at least at the stage boundary. These bioevents are correlated with the zonation defined for the stratotype (GSSP) of the base of the Turonian in the Western Interior (USA). The paleobiogeographic distribution of ammonites reveals some endemism but the predominant picture is that of a homogeneous fauna throughout the area, even though distinct Boreal and Western Tethys (Atlantic domain) marine influences are evident. An interpretation of the evolution of conch morphology and ornamentation through the zones of the Late Cenomanian–Early Turonian is proposed. 相似文献
15.
Sampling of Cenomanian fossil-rich horizons within the La Luna Formation of two localities in the Zulia and Trujillo states (northern Venezuela) yielded numerous shark teeth belonging to various species within the order Lamniformes (Mackerel sharks). Twelve lamniform species were identified including three new species (Squalicorax lalunaensis sp. nov., Squalicorax moodyi sp. nov., Acutalamna karsteni gen. et sp. nov.) and the genus Microcarcharias gen. nov. is proposed to accommodate with the peculiar morphology of the small-sized odontaspidid M. saskatchewanensis. Other taxa reported here include Cretoxyrhina mantelli, Cretolamna sp., cf. Nanocorax sp. and five Squalicorax species left in open nomenclature. This is the first report of chondrichthyans from the mid-Cretaceous of Venezuela and one of the few records of this group from the Cenomanian of South America. The composition of these assemblages suggests some degree of endemism in the La Luna Sea but also possible connexions with the Western Interior Seaway. One of the most striking features of these assemblages is the high anacoracid diversity (eight species) despite the corresponding outer shelf/upper slope palaeoenvironments of the La Luna Formation. The high diversity of these opportunistic predators is probably related to the high diversity of medium to large marine vertebrates that provided food resources. 相似文献
16.
An expedition to Melville Island in Nunavut, Canada, recovered the fragmentary fossils of several plesiosaurs from non-marine deposits of the Hauterivian–Aptian Isachsen Formation. These plesiosaur fossils are some of the oldest Early Cretaceous records of the group in North America, and they likely predate the formation of a continuous Western Interior Seaway. The plesiosaurs from Melville Island appear to be primarily juveniles, and would have been living in a region that experienced at least seasonally cool temperatures. The presence of these fossils in a fluvial deposit support previous suggestions that juvenile plesiosaurs may have preferentially inhabited shallower waters rather than open marine environments. These fossils also show that polycotylid plesiosaurs were able to rapidly disperse and colonize high latitude coastal regions, as their occurrence in Arctic Canada only slightly postdates the first confirmed appearance of the group in Australia. 相似文献
17.
The biochronology of Cenomanian-early Turonian ammonite faunas from three key stratotype areas (north-west Europe, central Tunisia and the Western Interior of North America) has been analysed and revised by utilizing the unitary association method. This review is prompted by the huge amount of biostratigraphic data published during recent decades and by a taxonomic homogenisation of the ammonite faunas from these key areas. The Cenomanian and lower Turonian of Tunisia comprise twenty-four Unitary Association zones and the middle Cenomanian-lower Turonian of the Western Interior Basin twenty-three such zones. The unitary association method means a two-fold increase in resolution of these ammonite zonations compared to the standard, empirical schemes. Central Tunisia and the Western Interior are correlated with north-west Europe by constructing a zonation including all taxa common to these areas. These correlations highlight the variable completeness and resolution of the faunal record through space and time, and reveal a significant number of diachronous taxa between the three areas. These correlations enable the designation of a new global marker for the middle/upper Cenomanian boundary, which is characterised by the disappearance of the genera Turrilites, Acanthoceras and Cunningtoniceras and by the appearance of Eucalycoceras, Pseudocalycoceras and Euomphaloceras. The only synchronous datum known is the last occurrence of Turrilites acutus, which may thus be used as a marker for the middle/upper Cenomanian boundary, provided that it does not turn out to be diachronous in the light of any new data. 相似文献
18.
Evidence from the Pierre Shale (Late Cretaceous) of South Dakota is presented for an attack on a juvenile Hesperornis by a polycotylid plesiosaur. The wound healed and the Hesperornis grew to maturity. Evidence of survival provides our best information about predator prey interactions in the fossil record but are rare for birds where survival is an unlikely outcome. 相似文献
19.
The Cenomanian–Turonian boundary was characterized by distinctive positive carbon isotope excursions that were related to the formation of widespread oceanic anoxia. High-resolution geochemical proxies (TOC, CaCO3, δ13Corg, and δ13Ccarb) obtained from bulk rock, planktic foraminifers, and inoceramids from four marine marlstone-dominated stratigraphic sections in the Western Canada Sedimentary Basin (WCSB) were used to establish a regional carbon isotope stratigraphic framework and to investigate paleoenvironmental variability in four different depositional settings. Compared to background δ13Corg, (<−27‰) and δ13Ccarb (<2‰) values which were correlative to stable isotope excursions during Oceanic Anoxic Event (OAE) II worldwide, the δ13Corg (>24‰), and δ13Ccarb (>4‰) derived from inoceramid prisms in the studied sections within WCSB, were elevated during the Late Cenomanian–Early Turonian. During this interval, TOC and CaCO3 values which increased sporadically to >40% and 7%, respectively, were not consistent enough to be used for stratigraphic correlations. Based on the δ13Corg excursions, two bentonite beds were regionally correlated across this portion of the Western Interior Seaway (WIS). The eruption associated with the “Red” bentonite occurred approximately coeval with the maximum δ13Corg-excursion during OAE II in the Neocardioceras juddii Zone, whereas the “Blue” bentonite coincides with the termination of OAE II in the latest Watinoceras devonense zone. During the Late Cenomanian–Early Turonian in the WCSB, benthic foraminifers were sparse or totally absent, indicating the existence of fully anoxic bottom-water conditions. Planktic foraminifera were common in the well-oxygenated surface waters. A benthic oxic zone characterized by several agglutinated species occurs in the eastern part of the WSCB at the beginning of OAE II in the Sciponoceras gracile zone. The termination of the OAE II in the WCSB coincides with the first occurrence of small ammonites (Subprionocyclus sp.) in the western part of the basin. 相似文献
20.
The new caprinoidean rudist bivalve Cobbanicaprina bighornensis gen. nov., sp. nov. is described from the upper middle Cenomanian of Big Horn County, Montana, USA. Cobbanicaprina gen. nov. is closely related to Mexicaprina and differs from that form in the absence of an external ligamental groove. The presence of Cobbanicaprina gen. nov. in the middle Cenomanian indicates that the Caprinuloideidae did not become extinct at the top of the Albian and persisted into the Cenomanian. The presence of this specimen so far north in the Western Interior Seaway is attributed to a dried-out individual specimen having been transported post-mortem by currents. 相似文献