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1.
Yabeinosaurus was the first lizard genus described from the Early Cretaceous Jehol Biota of China. The holotype of the type species, Y. tenuis, is an immature skeleton but it has been lost for decades. A second species, Y. youngi, was erected based on another immature skeleton and is distinguished by its longer, more gracile limbs. In 2001, a juvenile skeleton from the Jingangshan locality of Liaoning was designated as the neotype for Y. tenuis. Subsequently, several further specimens from other localities, juvenile and adult, have been attributed to this species. Here we describe a second lizard specimen from the neotype locality. In anterior tooth shape, finger-like mandibular angular process, cranial sculpture, maxilla shape, and short, robust humerus, the new specimen resembles adult and subadult material currently attributed to Y. tenuis, but it differs in having bicuspid posterior teeth and a straight rather than a hooked angular process of the mandible. We propose a new species (Y. bicuspidens sp. nov.) for this specimen which represents the first occurrence of bicuspid teeth in an Early Cretaceous lizard from China. However, the recognition of a second species at the neotype locality raises a taxonomic problem. Due to the immaturity of the designated holotype and neotype of Y. tenuis, they cannot be coded for the jaw and dental characters that distinguish the two robust-limbed species. This renders Yabeinosaurus tenuis a nomen dubium. Here we propose that Yabeinosaurus tenuis should be treated as a historic taxon, permitting retention of the generic name under ICZN rules, and erect a second new species (Y. robustus sp. nov.) for the principal robust-limbed morphotype in the Jehol Biota with monocuspid posterior teeth and a hooked angular process.  相似文献   

2.
Cretaceous dinosaur tracks from Hunan Province are historically significant as the basis for three ichnotaxa: Xiangxipus chenxiensis, Xiangxipus youngi, and Hunanpus jiuquwanensis all representing theropodan tracks, described from a single site in 1982. Although the type locality has since been destroyed, the type specimen and replicas remained available for restudy in 2006, when paratype Hunanpus tracks and sauropod footprints were described from a second, nearby locality. Material from both localities is here re-described in detail. It is proposed that while Xiangxipus chenxiensis can be regarded as a distinct ichnospecies, probably representing an ornithomimid trackmaker, Xiangxipus youngi cannot be accommodated in the same ichnogenus. We consider it similar to the ichnogenus Wupus from the Lower Cretaceous of Sichuan Province, and therefore of possible avian affinity. We also find no justification for regarding Hunanpus jiuquwanensis as generically distinct from the widespread ichnogenus Grallator, and therefore rename it Grallator jiuquwanensis comb. nov. The Hunan track record is generally consistent with the sparse record of theropod skeletal remains in the province, but adds evidence of sauropods that was previously lacking.  相似文献   

3.
Cervical vertebrae of elasmosaurids (Sauropterygia, Plesiosauroidea) have proven to be informative even at family level. The present paper uses a previously developed methodology based on bivariate graphic analysis that evaluates the proportion indices of cervical vertebrae in elasmosaurids, which is applied to a well-preserved postcranial specimen of this group recovered from upper Maastrichtian beds of central Chile. The plotting of such indices groups the Chilean material with other specimens from the Late Cretaceous of Argentina and Antarctica, the latter referred to the genus Aristonectes. This result allows us to identify the first relatively complete individual of the genus based on non-cranial characters. The identification of the Chilean specimen as Aristonectes sp. provides important new data on the postcranial skeleton of the genus, that has remained poorly known until now.  相似文献   

4.
The fossil skeletal record of birds from the Cenozoic of Switzerland is rather poor, despite the fact that avian tracks have been described from twenty tracksites. We review the Swiss fossil skeletal avifauna with emphasis put on new material discovered in the collection of the Natural History Museum Basel. This material includes two new owls (Strigiformes), one from a Late Eocene fissure filling from the Gösgen canal, and another from the Late Oligocene of Mümliswil. The Eocene owl specimen consists of a partial, distal tarsometatarsus, and is therefore too incompletely preserved to allow for reliable hypotheses concerning its taxonomic affinities. It does, however, display features resembling members of the extinct family Palaeoglaucidae. The Oligocene specimen is tentatively attributed to the genus Oligostrix, and it is the youngest representative of the extinct family Protostrigidae. Based on the first complete tarsometatarsus for this family, we erect a new species, ?Oligostrix bergeri. These two specimens represent the first record of fossil owls from Switzerland.  相似文献   

5.
A new species of Macrocnemus is described on the basis of two incomplete specimens from the Lower Ladinian Prosanto Formation of southeastern Switzerland. The new form can be distinguished by its gracile limb elements and having a noticeably longer tibia than either Macrocnemus bassanii or Macrocnemus fuyuanensis. One of the new specimens exhibits soft part preservation in the region of the pelvic girdle. The hind part of this individual is preserved fully articulated and allows a complete count of the caudal vertebrae to be made for the first time with 52 caudals. It is postulated that this specimen was predated upon and that the anterior part of the animal was consumed by a large predator such as a nothosaurid reptile or the actinopterygian fish Saurichthys.  相似文献   

6.
We re-define the Cretaceous bony fish genus Rhinconichthys by re-describing the type species, R. taylori, and defining two new species; R. purgatorensis sp. nov. from the lowermost Carlile Shale (middle Turonian), southeastern Colorado, United States, and R. uyenoi sp. nov. from the Mikasa Formation (Cenomanian), Middle Yezo Group, Hokkaido, Japan. Rhinconichthys purgatoirensis sp. nov. is designated on a newly discovered specimen consisting of a nearly complete skull with pectoral elements. Only known previously by two Cenomanian age specimens from England and Japan, the North American specimen significantly extends the geographic and stratigraphic range of Rhinconichthys. The skull of Rhinconichthys is elongate, including an expansive gill basket, and estimated maximum body length ranges between 2.0 and 2.7 m. Rhinconichthys was likely an obligate suspension-feeder due to its derived cranial morphology, characterized by a remarkably large and elongate hyomandibula. The hyomandibula mechanically acts as a lever to thrust the jaw articulation and hyoid arch both ventrally and anterolaterally during protraction, thus creating a massive buccal space to maximize filtering of planktonic prey items. Cladistic analysis supports a monophyly of suspension-feeding pachycormids including Rhinconichthys, but further resolution within this clade will require more information through additional fossil specimens.  相似文献   

7.
Recent work in the Tropic Shale by the Museum of Northern Arizona reveals a high biodiversity for plesiosaurs along the western margin of the Cretaceous Western Interior Seaway during the Cenomanian and the Turonian. This paper describes a new species of polycotylid plesiosaur from the Tropic Shale, Dolichorhynchops tropicensis, which adds to the known biodiversity from this time period. The identification is based on two specimens, a well-preserved, nearly complete skeleton including the skull and an additional specimen with only fragmentary skeletal elements. The material shares several synapomorphic characters with Dolichorhynchops, including the shape of the temporal fenestrae, the shape of the sagittal crest, the trend of the ectopterygoids, the morphology of the teeth, and the number of teeth within the mandibular symphysis. D. tropicensis differs from other species of Dolichorhynchops in a greater size range of the teeth, moderately constricted dorsal vertebrae, angled anterior processes on the coracoids, and the presence of well-defined facets on the propodials. The specimens are of Early Turonian age (based primarily on ammonite biostratigraphy and bentonite marker beds), and their discovery extends the known stratigraphic range for Dolichorhynchops back by approximately seven million years. This suggests that Dolichorhynchops tropicensis and Trinacromerum bentonianum, a closely related polycotylid, co-existed in the Western Interior Seaway.  相似文献   

8.
Sclerorhynchiform sawfishes are a diverse and extinct clade of elasmobranchs that is restricted to the Cretaceous. Most taxa are known only by isolated rostral spines, whereas skeletal remains are rare and have been reported from a small number of Upper Cretaceous localities. Here, we describe skeletal remains of the giant sclerorhynchiform Onchosaurus pharao for the first time, which provides new morphological information. The single specimen comes from middle-basal upper Turonian strata of the Lessini Mountains in northeastern Italy and represents the first record of this genus from Italy. The specimen consists of unidentifiable cranial remains, several diagnostic rostral spines, the rostrum with fragments of tessellated calcified cartilage, and 87 disarticulated vertebrae. The rostrum preserves the characteristic sensory system of sclerorhynchiforms. It is devoid of any lateral sockets indicating that rostral spines were attached laterally to its surface. This pattern is identical to most sclerorhynchiforms and extant pristiophoriformes implying also similar replacement patterns as in most other sclerorhynchiforms with the exception for Schizorhiza. Additionally, the bases of two longitudinally arranged rows of ventral rostral spines are identifiable concurring with patterns seen in Sclerorhynchus. The axial skeleton is partly preserved. Re-arranging the disarticulated vertebrae according to their life position in combination with measures of the size and thickness of preserved vertebral centra, and the ratio rostrum length/body size depending on the number of vertebral centra indicate that the specimen was ca. 450 cm long. Growth rings in the vertebral centra show that the specimen was about four years old and thus probably not yet fully sexual mature when it died. This age assumption corresponds well with the calculated size when compared with complete skeletons of extinct sclerorhynchiforms and extant pristiforms. The size of the specimen and its occurrence in hemipelagic rocks corroborates previous assumptions that this sclerorhynchiform was a large and pelagic sawfish.  相似文献   

9.
Upper Cretaceous deposits in Mongolia, Chinese Inner Mongolia, and, more recently, southern China, have yielded individually rich and taxonomically diverser lizard assemblages. Here we describe the remains of a new terrestrial lizard, Asprosaurus bibongriensis gen. et sp. nov., from the Upper Cretaceous of South Korea. It represents the first record of a Mesozoic lizard from the Korean Peninsula and, although incomplete, is exceptional in its very large size. Characters of the mandible support attribution to crown-group Anguimorpha, with the closest similarities being to monstersaurs, the group represented today by the venomous North American Beaded lizard and Gila monster, genus Heloderma. This group is well-represented in the Upper Cretaceous fossil record in of eastern Asia, and the remains of large monstersaurs have been recovered from several dinosaur egg localities, suggesting dietary preferences similar to those of the living genus. The new Korean lizard, recovered from the Boseong Bibong-ri Dinosaur Egg Site, fits the same pattern.  相似文献   

10.
A new species of Elcanidae (Orthoptera: Elcanoidea), Panorpidium yixianensis sp. nov., is described based on two new specimens from the Lower Cretaceous Yixian Formation of northeastern China. It differs from other species in forewing characters and spines on the hind tibiae. In addition, a new specimen Burmelcana sp., is described and figured based on an amber inclusion from the mid-Cretaceous Burmese (Myanmar) amber. P. yixianensis sp. nov. represents the first definite record of Elcanidae in the Early Cretaceous Jehol biota, and Panorpidium is the only genus of Ensifera to be found in the Early Cretaceous faunas of England, Russia and China.  相似文献   

11.
Fossil anguine lizard specimens from several Turkish localities are described in this paper. The material comes from ten different localities, spanning a large geographic area consisting of both parts of the European Turkey and Anatolia, and ranging in age from the Oligocene to the Late Miocene. In certain cases, the generic determination was possible and, accordingly, members of Ophisaurus and Anguis were identified and described in detail. The specimens of Anguis, found in different, Middle and Late Miocene localities from Anatolia, represent two of only a few fossil occurrences of this taxon. Moreover, the material reported herein represents the oldest occurrences of anguine lizards, not only from Turkey, but from southeastern Europe and the Eastern Mediterranean basin as a whole. These rare records provide important information about the dispersal routes of anguines from Europe to Asia and significantly enhance our understanding of their biogeography.  相似文献   

12.
In recent years the Late Cretaceous (Santonian) terrestrial vertebrate locality at Iharkút (western Hungary) has yielded well-preserved remains of lizard taxa besides the remains of fishes, amphibians, turtles, crocodiles, pterosaurs and dinosaurs. Previously the polyglyphanodontine lizard Bicuspidon aff. hatzegiensis has been reported from Iharkút. However, recent excavations at this site produced more lacertilian remains including new polyglyphanodontine material, namely a maxilla and two dentaries which suggest the presence of a new genus in the Iharkút fauna. This previously unknown lizard (described here as Distortodon rhomboideus n. g. n. sp.) is distinct from other polyglyphanodontines such as Bicuspidon, Paraglyphanodon, Polyglyphanodon, Dicothodon and Peneteius. It differs from these genera mainly in having the lingual cusp situated more distally compared to the labial one on its bicuspid teeth located in the distal part of the tooth row, thus the crowns having a unique rhomboidal shape in occlusal view. Distortodon rhomboideus further strengthens the dominance of borioteiioid lizards in the Iharkút fauna. The growing presence of borioteiioids in European localities supports previous theories which suggest some paleobiogeographic connections between the western Tethyan archipelago and North America in the Late Cretaceous.  相似文献   

13.
Herreraichthys coahuilaensis gen. and sp. nov. is described based on a single specimen collected in the Santonian marls strata of the “Los Temporales” quarry, Coahuila State, northern Mexico. This new species shows the diagnostic characters of the Family Lepisosteidae and tribe Lepisosteini, together with Lepisosteus and Atractosteus. This new fish shows two rows of teeth on dentary and lacrimomaxillae bones, including a lingual row of sharp and small regular size teeth, as well as a medial row of longer and fang-like teeth; this also has a lacrimomaxillary series as the main bite element of the upper jaw and the plicidentine structure on teeth. This Mexican fish differs from the other lepisoestinis in two characters; its lacrimomaxillary series is composed of 32 bones that constitute the largest series as far known among lepisosteiformes, and its premaxilla is comparatively wider and shorter. The occurrence of this specimen into an open marine deposit with no freshwater elements suggests that Herreraichthys was a marine inhabitant; however, there is the possibility that this species was able to temporarily survive in the sea, as Atractosteus spatula does now.  相似文献   

14.
Choristoderes are a group of extinct diapsid reptiles that once occupied the freshwater systems in the Northern Hemisphere from the Middle Jurassic through the Miocene. The Early Cretaceous monjurosuchid Philydrosaurus from western Liaoning, China, represents a transitional morphotype between a broad-snouted (crocodile-like) and a narrow-snouted (gavial-like) skull during the evolution of Choristodera. New specimens of the taxon from the Early Cretaceous Jiufotang Formation include an incomplete braincase and a nearly complete juvenile skeleton; the study of these specimens provides new information on the braincase and ontogenetic change of the skeleton of this transitional monjurosuchid. In the braincase floor, the foramen internus canalis caroticus externus opens in a groove lateral to the constricted lateral aspect of the parasphenoid, and ventral exposure of this foramen differs from Champsosaurus, in which parasphenoid/pterygoid fusion completely encloses the canal. On the occiput, the vagus nerve foramen penetrates the basioccipital, while the two foramina for the hypoglossal nerve open at the exoccipital/basioccipital suture. Comparative study of the new juvenile specimen with adult Philydrosaurus reveals previously unknown developmental changes of the cranial and postcranial skeleton of this monjurosuchid. Ontogenetically, Philydrosaurus underwent a dramatic change of skull proportions, including elongation of the antorbital and postorbital regions, and elongation of the jaws, with a great increase of the number of marginal teeth. The lower temporal fenestra is entirely closed in early ontogeny, as in large, fully-grown adults.  相似文献   

15.
The postcranial skeleton of a new specimen of the long-tailed lizard Dalinghosaurus longidigitus was excavated from the Yixian Formation in Lingyuan, western Liaoning. The new specimen provides more anatomical information about this species, especially about the anterior dorsal vertebrae, shoulder girdle and forelimbs. This lizard can be included within the clade Scleroglossa by its 27 or more presacrals, moderately long pubis, and gently notched distal end of tibia. But the detailed systematic position for this taxon remains undetermined. The features of the much longer hind limbs and pes compared with forelimbs and manus, metatarsal Ⅳ longer than Ⅲ, pedal phalanges robust, and penultimate phalanx not longer than other phalanges etc. suggest that this lizard was a running and ground swelling animal.  相似文献   

16.
17.
《Cretaceous Research》2012,33(6):781-785
The terrestrial lizard fossil record of Gondwana is very scarce. Few lizards, from mostly fragmentary fossils, have been identified in Madagascar, Tanzania, Morocco, South Africa, India and South America. Among the South American specimens there are basal Squamata forms (Olindalacerta and Tijubina) and a possible iguanid (Pristiguana) form Brazil. In Argentina gondwanian terrestrial lizards are represented by a putative iguanid and a small, poorly preserved dentary. This last specimen, recovered from the Anacleto Formation (Neuquén Group, Río Colorado Subgroup) in vicinity of the locality of Cinco Saltos (Río Negro Province), is here described formally. Comparisons with extant lizards indicate more affinity of the fossil with the Scincomorpha (non Scincophidia) than with any other group of lizards. This finding suggests that lizards were probably better represented and more diverse in the Mesozoic of South America than previously thought, although the fossil record is, at the moment, much poorer than in Laurasia.  相似文献   

18.
19.
Here we describe the well-preserved skull of a juvenile specimen of Sapeornis, STM 16-18. Only the tail and hindlimb plumage of this specimen have been previously described. It preserves what we consider to be the complete dentition of Sapeornis with four premaxillary teeth, three maxillary teeth and two tiny dentary teeth on each side, the latter feature being previously unknown for this taxon. Based on a study of 71 specimens of Sapeornis preserving dentition, we infer the absence of dentary teeth in previously described specimens of Sapeornis is taphonomic, and consider the true dental formula of Sapeornis to be 4-3-2 (premaxillary teeth number – maxillary teeth number – dentary teeth number). Rostral to the dentary teeth, STM 16-18 preserves three shallow, empty aveoli. We hypothesize that this morphology captures an early stage in dental reduction and indicates that in at least some avian lineages, the loss of teeth was gradual.  相似文献   

20.
The sea nettle,Chrysaora quinquecirrha, can capture ctenophore prey on its exumbrella, tentacles, and oral arms. Once attached to the medusa, the prey is transferred to the oral arms, where it becomes enveloped by the lip-like folds of the oral arms. The oral arms move the prey as far proximally as possible by ciliary creeping and muscular contractions. If the prey is too large to enter the stomach, it is digested within the oral arms by extracellular, proteolytic enzymes that probably originate from the gastrodermis of the oral arms and/or from the gastric cirri. Several large ctenophores can be digested simultaneously, owing to the extensive surface areas of the oral arms. Of the threeC. quinquecirrha medusae maintained in the laboratory for up to 60 d, the smaller two specimens (25–150 g wet weight) showed a biomass doubling in ~14 d; the largest specimen (300 g), a sexually mature female, showed little growth but did release >100 ova 24 h?1. Growth efficiencies were estimated to be ~2% for the largest specimen (not including biomass of gametes) and ~10% for the smaller two specimens. These growth efficiencies are comparable to those of other gelatinous predators.  相似文献   

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