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1.
Costate forms of Lagena (benthonic foraminifer) have been studied in an uppermost Maastrichtian bore-hole core sequence (28 levels) from southern Sweden. Analysis of a large sample shows that consistent assignment to previously described “species” of costate Lagena is not possible. This observation is supported by multivariate analysis of five morphologic variables which indicates that there are no distinct clusters of specimens along principal component axes associated with general size, number of costae, and relationships between roundness of test/length and number of costae. Specimens with a basal ring differ from specimens with an apical spine and those with no basal structure with regard to average number of costae. They also differ from specimens with no basal structure with regard to relationships between roundness of test/length and number of costae. These characters grade into each other thus preventing them from being used as taxonomic criteria.Mean ratios of benthonic foraminifers to total benthonic and planktonic formaminifers determined individually for each level are inversely related to mean relative frequencies of costate Lagena and directly related to mean general sizes of costate Lagena. The benthonic/planktonic ratio has been suggested by other workers to represent a paleobathymetric index across a continental slope and shelf when open marine conditions prevailed. If this were so in the Danish basin at the end of the Cretaceous, costate Lagena would have been relatively less frequent and larger at shallower depths.As a result of the study, the analyzed specimens are considered as representing a single morphospecies of the costate species Lagena sulcata. Between-level variation is regarded as ecophenotypic, and within-level variation as natural biologic variation of no taxonomic significance. A total of about 1000 species of Lagena has been described in the literature which is probably many orders of magnitude too great for a genus with such a simple morphology as Lagena. A revision of the taxonomy of Lagena is, therefore, probably necessary on a wide scale.  相似文献   

2.
The restudy of the Late Cretaceous rudist fauna of the Chiapas Central Depression, considered Campanian-Maastrichtian with no more precisions in literature, reveals that, in fact, three successive rudist assemblages occur: (1) a lower one, early and middle Campanian, in the uppermost part of the Sierra Madre Formation (distinguished as Suchiapa Formation); (2) a middle one, early Maastrichtian, in the upper part of the Ocozocoautla Formation; and (3) an upper one, late Maastrichtian, in several horizons within the Angostura Formation. Published data on planktic and benthic foraminifers and inoceramid bivalves, as well as new findings of ammonites, helped bracketing the age of the three rudist assemblages. Their stratigraphic position fits with a depositional model assuming a basal carbonate platform fragmented into blocks each following a different tecto-sedimentary evolution: (1) a drowned block constituting the Tuxtla Gutiérrez Basin, deepening towards the neighbor raised block and receiving terrigenous material from the emerged Sierra Madre de Chiapas, that, after a long hiatus, was filled up and, subsequently, a marginal shallow carbonate platform, surrounded by a detritic belt and prograding towards the basin, installed on top; (2) a raised block constituting a high where, after a long hiatus, an insular shallow carbonate platform installed.  相似文献   

3.
Five radiolitid rudist species are described from the Turonian sequence of Abu Roash area. They are recognized in three rudist biostromes that occur in two informal members of Abu Roash Formation; the Rudist- and the Actaeonella-bearing limestone–marl members. The three biostromes show autochthonous and parautochthonous fabrics and moderate to high packing potential. The first rudist biostrome at the base of the Rudist-bearing limestone–marl member (Middle Turonian) contains Durania gaensis, Praeradiolites ponsianus and Bournonia fourtaui. The second biostrome in the same member consists of Bournonia roashensis. The third biostrome that recognized in the Actaeonella-bearing limestone–marl member (Late Turonian) consists of Durania arnaudi. Rudist biostromes in the Rudist-bearing limestone–marl member were deposited on subtidal rudist shoals with moderate to high energy versus that of the Actaeonella-bearing limestone–marl member that deposited in low to moderate energy on deeper part of subtidal rudist shoals. The exposed Turonian succession at Abu Roash area could be divided into three depositional sequences bounded by three sequence boundaries (paleosols and angular unconformity).The first rudist biostrome in the Rudist-bearing limestone–marl member represents the lower part of the transgressive systems tract of the first depositional sequence. The deepening upward trend of the transgressive systems tract is due to increase of accommodation space in transgressive context during relative sea-level rise episode. On the other hand, the second rudist biostrome in the Rudist-bearing limestone–marl member and the third rudist biostrome in the Actaeonella-bearing limestone–marl member are in shallowing-upward set sequence forming the highstand systems tract of the first and third depositional sequences. This indicates that, the accommodation space was being filled more rapidly than was being created during the highstand stage.  相似文献   

4.
The M 7.0 Haiti earthquake of 2010 in the Greater Antilles is a reminder that the northeastern Caribbean is at a high risk for seismic and tsunami hazards. The Greater Antilles consist of the Hispaniola microplate to the west and Puerto Rico–Virgin Islands to the east and are situated between two subduction zones with the Puerto Rico Trench to the north and the Muertos Trough to the south. Although there is no active volcanism on Puerto Rico, earthquake depths and previous seismic tomography results imply that the slabs of Caribbean and North American Plates exist at depth. However, how far the east Muertos Trough subduction of the North Caribbean Plate has extended has not been fully addressed. In addition, the Puerto Rico–Virgin Islands are bounded by extensional regimes to both the west (Mona Rift) and east (Anegada Passage). The cause of the extension is still under debate. In this paper, we use new 3D seismic tomography and gravity data to carry out an integrated study of the geometry of the subducting slabs of the North American and North Caribbean Plates in the Puerto Rico–Virgin Islands area. The results indicate that both slabs have an increase of dip westward, which is strongly controlled by the subduction rollback of the North American Plate. These variations affected the tectonic evolution of the Puerto Rico–Virgin Islands. Thus, the results of this research advance our understanding of the kinematic evolution of the Puerto Rico–Virgin Islands and associated natural hazards. Copyright © 2015 John Wiley & Sons, Ltd.  相似文献   

5.
The uppermost Cretaceous (upper Campanian-Maastrichtian) pelagic successions from the Malatya Basin (NW Malatya, eastern Anatolia) were studied by 688 samples, which were collected from five stratigraphic sections in the Hekimhan area. The pelagic deposits conformably overlie rudist bearing shallow-water limestones and are overlain conformably by Maastrichtian dolomites and unconformably by Paleocene-Eocene deposits, respectively.The pelagic successions in the Hekimhan area comprise the Kösehasan Formation at the base and the Zorbehan Formation at the top and reach up to 1100 m in thickness. The Kösehasan Formation rests over the neritic rudist-bearing limestones of the Güzelyurt Formation along a sharp contact and consists mainly of flysch-type sandstone-mudstone alternation with complete and partial Bouma sequences. The carbonate content of abundant planktonic foraminifera and nannoplankton-bearing 980-m-thick succession increases upwards and the formation passes gradually to the clayey limestones and marlstones of the Zorbehan Formation to the top. Occurrences of nannoplankton Lithraphidites quadratus Bramlette and Martini and Micula praemurus (Bukry) in the first beds of the Kösehasan Formation indicate that the age of the Kösehasan Formation and overlying Zorbehan Formation is of late Maasthrichtian. Another late Maastrichtian taxa Cribrosphaerella daniae Perch-Nielsen and Arkhangelskiella maastrichtiana Burnett are observed from the lowermost part of the succession. Maastrichtian planktonic foraminifera such as Contusotruncana walfischensis (Todd) and Globotruncanita pettersi (Gandolfi) were recorded through the successions. Although planktonic foraminifera are diverse and abundant particularly in the Kösehesan Formation, index late Maasthrichtian species were not encountered. Campanian and Santonian-Campanian planktonic foraminifera, e.g. Radotruncana calcarata (Cushman) and Globotruncanita elevata (Brotzen), obtained particularly from the lower part of the succession and calcareous nannofossils such as Broinsonia parca parca Bukry, Reinhardtites anthophorus (Deflanre) and Eiffellithus eximius (Stover) are interpreted as reworked from older strata. Trace fossils are common throughout the succession.Rareness of planktonic foraminifera and nannoplankton in the uppermost part of the succession (Zorbehan Formation) indicates maximum shallowing of the latest Maastrichtian sea in this part of the basin. Rare echinoids, bivalves and ammonites are observed in that part of the sequence.The obtained data indicate that sediment accumulation rate of the pelagic deposits is rather high and about 27.5 cm/ky for this part of the basin. Changes in thickness of the formations along short distances in the five stratigraphic sections analysed in this study should be related to the diachroneity of the depositional and erosional events.  相似文献   

6.
The biostratigraphy and the response of calcareous nannofossils to the End Cretaceous warming are investigated in the lower boundary of Kalat formation through the record of species richness, diversity, distribution patterns, and statistical treatments. The Kalat formation comprised of coarse-grained detritus limestone with subordinate sandstone intercalations. In the studied sections, the number of ten samples were taken and prepared with smear slide. In Dobaradar, section 22 species; in Kalat, section 25 species; and in Chahchaheh, section 32 species have been determined. Based on nannoplanktons and as a result of biostratigraphic studies, the nannofossil standard zones (CC25–CC26) were identified in all of sections. According to these zones in all of sections, the age of the studied thickness is Late Maastrichtian–Late Late Maastrichtian. In these sections, the presence of Micula murus at the end of Neyzar formation and the presence of this species at the lower part of Kalat formation indicate that the investigated boundary is Late Maastrichtian in age. The paleoecological results point to warm climate. The presence of warm water indicators (M. murus and Micula prinsii) and the absence of cool water indicators (Ahmuellerella octoradiata, Kamptnerius magnificus, and Nephrolithus frequens) suggest warm surface water conditions in these areas. In the lower boundary of Kalat formation, base on Lithraphidites spp. and Watznaueria barnesae, lowered fertility condition with low productivity at the end of the Maastrichtian were suggested, and the studied area was deposited in shallow marine environment in relatively low latitude.  相似文献   

7.
A numerical-dynamic, tropical storm surge model, SLOSH (Sea, Land, and Overland Surges from Hurricanes), was originally developed for real-time forecasting of hurricane storm surges on continental shelves, across inland water bodies and along coastlines and for inland routing of water -either from the sea or from inland water bodies. The model is two-dimensional, covering water bodies and inundated terrain. In the present version available at the University of Puerto Rico a curvilinear, polar coordinate grid scheme is used. The grid cells are approximately 3.2 × 3.2 km in size.The model has been used in a revision of all coastal Flood Insurance Rate Maps (FIRM) for Puerto Rico and the U.S. Virgin Islands, and in hurricane evacuation studies. The FIRM's, since they are based on the 100 year stillwater elevation, are also used by the state Planning Board for regulatory purposes. The hurricane evacuation studies are used by emergency planners and personnel to assign shelters, escape routes, and delimit coastal zones that need to be evacuated during a hurricane threat.Recently, the acquisition of data from hurricane Hugo has allowed the first comparison of model results and observations for Puerto Rico and the other islands.  相似文献   

8.
The siderolitids from the uppermost Campanian and Maastrichtian deposits of the Pyrenees have been re-studied. This has revealed a high diversity and rapid replacement of taxa, confirming the group as a good tool for high resolution biostratigraphy. Two genera have been found in the uppermost Campanian–Maastrichtian interval in the Pyrenean deposits: Siderolites Lamarck, and Wannierina Robles-Salcedo. Siderolites, with canaliferous spines or denticulate periphery, is represented by four species replacing each other from the latest Campanian to Maastrichtian: Siderolites praecalcitrapoides (latest Campanian), S. pyrenaicus sp. nov. (early Maastrichtian), Siderolites calcitrapoides (late Maastrichtian) and Siderolites denticulatus (late Maastrichtian). Wannierina is characterised by well-developed keels and ramified marginal canals. Two species of Wannierina have been identified and they succeeded one another from latest Campanian to early Maastrichtian: Wannierina vilavellensis sp. nov. (latest Campanian) and Wannierina cataluniensis (early Maastrichtian). The species of the genus Siderolites inhabited shallow waters of tropical to subtropical platforms with moderate-to-high water-energy conditions and those of the genus Wannierina are typical of deep–water low-energy environments but still in the eutrophic zone.  相似文献   

9.
This study is based on calcerous nannofossil assemblages changes and fluctutions of stable carbon and oxygen isotopes was collected clayey limestones, limestones, and marls in the Maastrichtian to Selandian from Akveren Formation (Western Black Sea). As the relative abundances of species of Micula spp, Watznauera barnesiae, and Arkhangelskiella cymbiformis, which tolerated changes of temperature and nutrition, carbon and oxygen isotopes compositions, and low species richness imply strong diagenesis effect at the Maastrichtian, there is no important diagenesis effect at Paleocene. Just after the Cretaceous/Tertiary (K-Pg) boundary, Thoracosphaera spp. and Braarudosphaera bigelowi were dominant species; Danian is characterized by Thoracosphaera, Ericsonia ovalis, Cruciplacolithus spp., Coccolithus pelagicus, and Ericsonia subpertusa. Generally, the nutrition productivity is good–moderate in Lower Maastrichtian, and decreasing carbon isotope values during the Uppermost Maastrichtian shows the presence of oligotrophic environmental conditions suitable with global nutrition crises before the K-Pg boundary and diagenesis in study area. Throughout the Danian, mesotrophic–oligotrophic environmental conditions dominate; however, the decrease in nutrition before Selandian represents oligotrophic environmental conditions. The increasing nutrition at Selandian is related to the change in the environmental conditions.  相似文献   

10.
A year-long trawl survey of the mangrove-fringed Laguna Joyuda, Puerto Rico yielded 41 species of juvenile and adult fishes. Twelve percent of the species and 55% of the individuals were residents in the lagoon; 56% of the species and 44% of the individuals were cyclic visitors, mostly juveniles of species which spawn offshore. The lagoon yielded fewer species than tropical estuaries in continental regions of the Caribbean and temperate estuaries of North America. However, paralleling other estuarine habitats, Laguna Joyuda supported three primary types of fishes, a resident small flatfish (Achirus lineatus), a complex of transient juveniles (Gerreidae, paralleling the Sciaenidae of higher latitudes), and small planktivores (Anchoa spp.). Seasonal patterns in the abundance of individual species were not strong, but overall abundance showed a wet season maximum, particularly because of recruitment of anchovies and cyclic visitors in April, May, and June. Wet season abundance corresponded with highest water column productivity and detrital input to the benthos. Fishes associated with the water column appeared to show greater variation in abundance than those associated with more stable benthic food webs.  相似文献   

11.
The stratigraphic and palaeoenvironmental implications of a section in the Upper Senonian to Lower Eocene carbonates of the Tripolitza Platform in central Crete are discussed in this paper. The lower part (upper Campanian–lower Maastrichtian) of the succession consists of about 75 m of thickly bedded, light to dark grey limestones, dolomitic limestones and dolomites that were deposited on an inner carbonate platform. It is characterized by stratified bioclastic rudist facies (shelly limestone) associated with foraminifera and especially with species of the Rhapydioninidae family. These are overlain by 35 m of crystalline light grey dolomites that were deposited on a very restricted internal platform, characterized by intertidal-supratidal facies. The overlying 75-m-thick light grey dolomitic limestones and dolomites are characterized by the presence ofRhapydionina liburnicaStache, and in the uppermost part byNeobalkhania bignotiCherchi, Radoicic & Schroeder, dating it as Late Maastrichtian. The facies, cryptalgal laminites with fenestrae of varying dimensions, suggest relatively extensive subaerial exposure. Possible pedogenic textures are common in this sequence and especially in the uppermost part, which coincides with the K/T boundary. The presence ofPseudonummoloculina heimi(Bonet) at two levels in the Maastrichtian succession suggests transportation of Middle-Late Cenomanian sediments from emergent blocks of the platform during this period. A gap is suspected, for regional stratigraphic reasons, between this horizon and the next which containsSpirolinasp. and “Pseudochrysalidina” sp., dating it as Early-Middle Eocene.  相似文献   

12.
Numerical Simulation of the 1918 Puerto Rico Tsunami   总被引：1，自引：0，他引：1  
Mercado  A.  McCann  W. 《Natural Hazards》1998,18(1):57-76
The Caribbean Sea region is well known for its hurricanes, and less known for tsunamis. As part of its responsibilities in hazard assessment and mitigation, the U.S.A. Federal Emergency Management Agency, and the Puerto Rico Civil Defense, funded a pilot study to perform a numerical simulation of the 1918 Puerto Rico tsunami, one of the most deadly in the region. As part of the study a review has been made of the tectonic and tsunamigenic environment around Puerto Rico, the fault parameters for the 1918 event have been estimated, and a numerical simulation has been done using a tsunami propagation and runup model obtained through the Tsunami Inundation Modeling for Exchange (TIME) program. Model results have been compared with the observed runup values all along the west coast of Puerto Rico.  相似文献   

13.
This study of the upper Maastrichtian to Danian sedimentary succession from the northern part of the Romanian Eastern Carpathians (Varniţa section) aims to establish an integrated biostratigraphy based on calcareous nannofossils, organic-walled dinoflagellate cysts (dinocysts) and foraminiferal assemblages, and to reconstruct the depositional environments of the interval. The stratigraphic record across the studied section is incomplete, considering that an approximately 16 m thick strata interval from the top of the Maastrichtian to lowermost Danian cannot be analyzed due to a landslide covering the outcrop. The upper Maastrichtian is marked by a succession of biostratigraphic events, such as the First Appearance Datum (FAD) of the nannoplankton taxon Nephrolithus frequens and FAD of the dinocyst species Deflandrea galeata and Disphaerogena carposphaeropsis, and the Last Appearance Datum (LAD) of Isabelidinium cooksoniae in the lower part of the section. These bioevents are followed by the LAD of the Dinogymnium spp. and Palynodinium grallator dinocyst markers in the top of the Maastrichtian deposits analyzed. In terms of foraminiferal biostratigraphy, the upper Maastrichtian Abathomphalus mayaroensis Zone is documented in the lower part of the studied section. Some bioevents, such as the bloom of the calcareous dinoflagellate genus Thoracosphaera and the FAD of the organic-walled dinocysts Damassadinium californicum, Senoniasphaera inornata, Xenicodinium lubricum and X. reticulatum suggest an early Danian age for the middle part of the section. From the Danian deposits in the Varniţa section, we describe a new organic-walled dinocyst species, Pentadinium darmirae sp. nov., which is until now the only species of the Pentadinium genus discovered in the Paleocene. The occurrence of the global Danian dinocyst marker Senoniasphaera inornata in the top of the section, suggests an age not younger than middle Danian (62.6 Ma) for the analyzed deposits.The palynofacies constituents, as well as the agglutinated foraminiferal morphogroups, used to reconstruct the depositional environments, show that the late Maastrichtian sediments were deposited in an outer shelf to distal (bathyal) environment, followed by a marine transgression during the Danian.  相似文献   

14.
We compared species presence, abundance, and size characteristics of fish in three brackish, coastal marshes at Humacao, Roosevelt Roads, and Boqueron, Puerto Rico, in February and March 1988. The three marsh ecosystems were similar with respect to the presence of large expanses of open water bordered by emergent vegetation, creeks, and mangroves, and all had some recreational use. We sampled fish using gill nets. Tilapia (Oreochromis) mossambica were the most abundant fish, accounting for 55–79% of the samples at all three marshes. Overall, tilapia were both the largest (North Lagoon) and the smallest (Frontera Creek) at Humacao. Tilapia were most common in open lagoons rather than creeks or bays (except for Mandri Creek), and their distribution seemed unrelated to salinity. Tarpon (Megalops atlantica) were more abundant at low salinities, whereas other fish were more abundant at higher salinities.  相似文献   

15.
A detailed biostratigraphic study of four sections (El Kantara, Menaa, Tighanimine and El Gaâga) in the Campanian–Maastrichtian of the Aures Basin in Algeria allows the identification of 109 ostracod species belonging to 50 genera. From a biochronostratigraphic point of view, none of the identified ostracod assemblages are typically restricted to the Campanian or to the Maastrichtian, but rather characterize a range covering the Campanian–Maastrichtian. As regards the paleobiogeography, the presence of species in common between Algeria and other countries or regions enables us to compile four maps showing the distribution of two provinces that developed during the Santonian-Paleocene, i) the South-West and South Tethyan Margins Province, including North Africa, the Middle East and Arabian Peninsula, and ii) the East Atlantic Margin Province, bordering West and Central-West Africa. We can also recognize relationships between the different regions making up these provinces, which remained under more or less arid, subtropical to tropical climates, with ostracods migrating according to the directions of ocean currents during the Late Cretaceous. The dispersal of the ostracods reflects marine communication between these provinces and India, the southern USA and Jamaica, Brazil and Europe.  相似文献   

16.
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17.
The Aqra Formation represents a succession that was deposited over most of Northern Iraq and adjacent regions. In north Iraq, in the core of NW–SE trending Aqra anticline, a 438-m-thick section of the Aqra Formation crops out at Geli Zanta corge. The base of the Aqra Formation is not exposed here. The upper contact is unconformably overlain by Paleocene–Lower Eocene formations (Kolosh and Khurmala formations). A hundred and one samples were collected from the section and used for biostratigraphic and microfacies analysis. According to the occurrence of larger Foraminifera (Orbitoides media and Orbitoides apiculata) and planktonic Foraminifera (Abathomphalus mayaroensis), Late Upper Campanian–Maastrichtian age was determined for Aqra Formation. Fifteen facies were distinguishable throughout the formation, representing tidal flat (supratidal), restricted marine shelf (lagoon) and shelf margin rudist reef, and its related debris. These environments were used to interpret three depositional sequences which correlate with those of Aruma Formation (KSA), Simsima Formation (UAE) in Arabian Plate, and with Iraqi formation sequences. Three maximum flooding surfaces were recognized as MFS 175, MFS 180, and MFS 190.  相似文献   

18.
Island arc volcanism in the Greater Antilles persisted for >70m.y. from Middle Cretaceous to Late Eocene time. During theinitial 50 m.y., lavas in central Puerto Rico shifted from predominantlyisland arc tholeiites (volcanic phase I, Aptian to Early Albian,120–105 Ma), to calc-alkaline basalts (phase II, LateAlbian, 105–97 Ma), and finally to high-K, incompatible-element-enrichedbasalts (phases III and IV, Cenomanian–Maastrichtian,97–70 Ma). Following an island-wide eruptive hiatus, geochemicaltrends were reversed in the Eocene with renewed eruption ofcalc-alkaline basalts (phase V, 60–45 Ma). Progressiveincreases in large-ion lithophile elements (LILE)/light rareearth elements (LREE), LILE/high field strength elements (HFSE),LREE/HFSE, and HFSE/heavy rare earth elements (HREE) characterizethe compositional evolution of the first four volcanic phases.The shift in trace element compositions is mirrored by increasingradiogenic content of the lavas. Pb  相似文献   

19.
The Cape Lamb Member of the Snow Hill Island Formation at The Naze on the northern margin of James Ross Island, east of the Antarctic Peninsula, yielded a theropod dinosaur recovered near the middle of a 90 m thick section that begins at sea level, ends below a basalt sill, and is composed of interbedded green–gray massive and laminated fine-grained sandstones and mudstones. Sixteen palynoassemblages were recovered from this section, which yielded moderately diverse assemblages with a total of 100 relatively well-preserved species. The principal terrestrial groups (32%) are represented by lycophytes (8 species), pteridophytes (15 species), gymnosperms (13 species), angiosperms (21 species) and freshwater chlorococcaleans (3 species). Marine palynomorphs (68%) belong to dinoflagellates (61 species), chlorococcaleans (6 species), and one acritarch. The vertical distribution of selected species allows the distinction of two informal assemblages, the lower Odontochitina porifera assemblage from the base to its disappearance in the lower part of the section, and the remaining section characterized by the Batiacasphaera grandis assemblage. The global stratigraphic ranges of selected palynomorphs suggest an early Maastrichtian age for this section and the entombed dinosaur that is also supported by the presence of the ammonoid Kitchinites darwinii. These assemblages share many species with latest Campanian–early Maastrichtian palynofloras from Vega and Humps Islands, New Zealand, and elsewhere in the Southern Ocean, establishing a good correlation among them. The dominance or frequent presence of dinoflagellates throughout the section supports the general interpretation of a shelf marine depocenter. The consistent presence of terrestrial palynomorphs suggests contributions from littoral/inland environments.  相似文献   

20.
Charophytes bearing small sized fructifications dominated in fluviatile floodplain facies (red beds) from the Maastrichtian of Coll de Nargó and neighbouring basins in the southern Pyrenees (Catalonia, Spain). These charophytes mainly belong to the genus Microchara and are often disregarded in biostratigraphic studies, which focus their attention instead on facies from permanent lakes that are richer in species which usually bear fructifications with a larger size range. However, small sized gyrogonites are also significant for biostratigraphic purposes and even include some of the index species for Maastrichtian biozones in Europe. Indeed, the charophyte assemblages from the Maastrichtian of Coll de Nargó belong to the Microchara punctata biozone, recently calibrated to the middle-upper Maastrichtian.Floodplain ponds from the Maastrichtian red beds of Coll de Nargó (Lower Red Unit) were extremely shallow, received considerable terrigenous influx and were frequently exposed, probably resulting in turbid, warm waters with high light availability. These conditions could explain the almost exclusive occurrence of charophytes with small fructifications in the Lower Red Unit. The available data, mainly based on oospores from extant species, indicate that the small size observed in gyrogonites from temporary ponds may represent an adaptation to environmental stress. Fossil species with small gyrogonites of Microchara cristata, M. punctata and Microchara nana would thus develop massively in stressed shallow ponds on fluvial floodplains. To contrast these hypotheses, we compared our results to those of four well-known case studies with similar sedimentological contexts, ranging from the Lower Cretaceous to the upper Eocene–lower Oligocene. Gyrogonite size patterns were similar in all cases, possibly suggesting that characeans display a long history of adaptation to shallow, temporary and turbid floodplain ponds by means of producing a high number of small gyrogonites, probably representing short life cycles and opportunistic strategies.  相似文献   

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