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Verónica Díez Díaz Géraldine Garcia Fabien Knoll Xabier Pereda Suberbiola Xavier Valentin 《Proceedings of the Geologists' Association. Geologists' Association》2012,123(4):626-637
The Fox-Amphoux Syncline of the Aix Basin (SE France) has yielded a diverse Late Cretaceous vertebrate assemblage, including several taxa of dinosaurs. Here, we report on cranial material of titanosaurian sauropods, which consist of a partial braincase, two fragmentary skull roofs, and nine teeth, from the Métisson locality (Var Department). The braincase differs from those found previously in Europe (i.e., Ampelosaurus atacis, Lirainosaurus astibiae, and an unnamed juvenile skull from Romania) as well as from other titanosaurian braincases in having a groove that extends along the ventral surface of the occipital condyle neck (this feature may be autapomorphic). One of the fragmentary skull roofs may belong to the same taxon, whereas the other suggests the presence of a second titanosaur at Métisson very close to Ampelosaurus. Two dental morphologies are present in the sample; one of them includes teeth of different sizes. We suggest that this could be accounted for by age differences within a single taxon, or be due to different positions in the tooth row. The presence of a new, still unnamed titanosaurian taxon in the Ibero-Armorican Island supports previous works indicating a high titanosaurian diversity during the Campanian-Maastrichtian in southern Europe. 相似文献
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Yabeinosaurus was the first lizard genus described from the Early Cretaceous Jehol Biota of China. The holotype of the type species, Y. tenuis, is an immature skeleton but it has been lost for decades. A second species, Y. youngi, was erected based on another immature skeleton and is distinguished by its longer, more gracile limbs. In 2001, a juvenile skeleton from the Jingangshan locality of Liaoning was designated as the neotype for Y. tenuis. Subsequently, several further specimens from other localities, juvenile and adult, have been attributed to this species. Here we describe a second lizard specimen from the neotype locality. In anterior tooth shape, finger-like mandibular angular process, cranial sculpture, maxilla shape, and short, robust humerus, the new specimen resembles adult and subadult material currently attributed to Y. tenuis, but it differs in having bicuspid posterior teeth and a straight rather than a hooked angular process of the mandible. We propose a new species (Y. bicuspidens sp. nov.) for this specimen which represents the first occurrence of bicuspid teeth in an Early Cretaceous lizard from China. However, the recognition of a second species at the neotype locality raises a taxonomic problem. Due to the immaturity of the designated holotype and neotype of Y. tenuis, they cannot be coded for the jaw and dental characters that distinguish the two robust-limbed species. This renders Yabeinosaurus tenuis a nomen dubium. Here we propose that Yabeinosaurus tenuis should be treated as a historic taxon, permitting retention of the generic name under ICZN rules, and erect a second new species (Y. robustus sp. nov.) for the principal robust-limbed morphotype in the Jehol Biota with monocuspid posterior teeth and a hooked angular process. 相似文献
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The type braincase of Viavenator exxoni (MAU-Pv-LI-530) was recovered complete and isolated from most of the other skull bones. Although the braincase is crossed by numerous fractures, using CT scans allowed the generation of 3D renderings of the endocranial cavity enclosing the brain, cranial nerves, and blood vessels, as well as the labyrinth of the inner ear. Within the abelisaurids, the only taxon with a complete braincase and known endocranial morphology is Majungasaurus crenatissimus, from Madagascar. In turn, in Argentina, partial endocranial morphology is known for another two Cretaceous forms: Abelisaurus comahuensis and Aucasaurus garridoi. Here, we present the most complete reconstruction of the neuroanatomy for a representative of the clade in South America. These findings add knowledge to the field of theropod paleoneuroanatomy in general, and abelisaurid diversity in particular. Comparisons of Viavenator with other abelisaurids indicate greater similarity with Aucasaurus than with Majungasaurus, suggesting that South American forms shared the same neurosensorial capabilities, which include larger flocculus of the cerebellum and larger olfactory ratios than the form from Madagascar. 相似文献
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Pachycephalosaurs, a group of ornithischian dinosaurs with distinctive cranial ornamentation and skull domes, underwent dramatic changes in cranial morphology during ontogeny. This has caused debate about whether some specimens belong to juveniles or adults, which impacts studies of pachycephalosaur phylogeny and evolution. One such debate concerns a small skull roof specimen from the Campanian (Upper Cretaceous) of New Mexico, NMMNH P-33898, which was originally described as an indeterminate juvenile but later regarded as a mature adult and erected as the holotype of a new small-bodied species, Stegoceras novomexicanum. We restudied NMMNH P-33898 using computed tomography scanning, morphometric and phylogenetic analyses, and comparisons to growth series of other pachycephalosaurs (Stegoceras validum, Pachycephalosaurus wyomingensis). We conclude that two purported paratype specimens of Stegoceras novomexicanum cannot be referred to the same taxon as the holotype, that the holotype and controversial paratypes all belong to immature specimens and not aberrant small-bodied adults, but that current evidence cannot clearly determine whether NMMNH P-33898 is a juvenile belonging to its own diagnostic species (S. novomexicanum) or is a juvenile of Stegoceras validum, Sphaerotholus goodwini, or another known taxon. We review the pachycephalosaur record of New Mexico and demonstrate that pachycephalosaurs were important components of dinosaur faunas in the southern part of Western North America during the ∼15 million years before the end-Cretaceous extinction, just as they were in roughly contemporaneous northern localities. 相似文献
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The Early Cretaceous lizard Yabeinosaurus was one of the first taxa described from the now famous Jehol Biota of northeast China. Misinterpreted for more than 60 years and misclassified as a gekkotan based on juvenile specimens, it is now recognised to be a large, well-ossified lizard with an extended period of skeletal maturation. Here we describe two additional complete specimens of Yabeinosaurus that provide new information on skeletal morphology, most notably of the skull, pectoral girdle, and tail. Both specimens also preserve gut contents, showing that large individuals took vertebrate prey, including fish. A more complete understanding of Yabeinosaurus permits a review of the type specimen of Yabeinosaurus youngi. Skull traits used to distinguish Y. youngi from Yabeinosaurus tenuis are invalid but, as noted by Hoffstetter, the two species differ markedly in limb proportions. Attribution of Young’s specimen to Yabeinosaurus is equivocal, but could potentially extend the temporal range of the lineage into the Jurassic. A new phylogenetic analysis based on a morphological data set places Yabeinosaurus on the stem of Scleroglossa, as the sister taxon of the contemporaneous Japanese lizard Sakurasaurus. 相似文献
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A new ctenochasmatoid pterosaur from the Early Cretaceous Yixian Formation of western Liaoning,China
A new ctenochasmatoid pterosaur, Gladocephaloideus jingangshanensis gen. et sp. nov. from the Yixian Formation of western Laioning Province is erected based on a complete skull and partial postcranial skeleton. It is characterized by following features: about 50 total teeth with sharp tips; small nasoantorbital opening, occupying approximately 13% of the skull length; ratio of prenarial length to skull length approximately 0.63. The diagnoses of the Ctenochasmatoidea and Gallodactylidae are amended based on the new taxon. Gladocephaloideus jingangshanensis is the first gallodactylid pterosaur found in Asia. Its discovery not only provides much more osteological information about the Gallodactylidae but also indicates that the ctenochasmatoid pterosaurs were highly diverse in the Early Cretaceous. The filamentous structures preserved near the dorsal and posterior margins of the posterior portion of the skull and around the neck indicate that it had an epidermal covering and may have been a warm-blooded animal. 相似文献
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Metriorhynchidae is a clade of marine-adapted crocodilians known from several Middle Jurassic–Early Cretaceous specimens collected predominantly in South America and Europe, but poorly known in the northern margin of Gondwana. The “Portomaggiore crocodile” is the most complete specimen of an Italian metriorhynchid to date: it consists of a partial skeleton that has been provisionally referred to an unnamed species of Late Jurassic Metriorhynchus or Geosaurus. The specimen is preserved in the reddish, nodular limestone of the Rosso Ammonitico Veronese Formation (Bajocian–Tithonian); new data on microfossil associations constrain the age of the metriorhynchid to the late Bajocian–earliest Bathonian. On the basis of cranial synapomorphies, the “Portomaggiore crocodile” falls as the closest sister-taxon of the Late Jurassic–Early Cretaceous geosaurines, and is referred to Neptunidraco ammoniticus gen. et sp. nov. It is unique among Middle Jurassic metriorhynchids in showing an incipient streamlining of the skull, shared with Late Jurassic and Cretaceous taxa. Since Neptunidraco is the oldest known member of Metriorhynchidae, its phylogenetic position supports the hypothesis that the timing of the initial metriorhynchid and geosaurine diversifications should start in the Bajocian. 相似文献
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Recent work in the Tropic Shale by the Museum of Northern Arizona reveals a high biodiversity for plesiosaurs along the western margin of the Cretaceous Western Interior Seaway during the Cenomanian and the Turonian. This paper describes a new species of polycotylid plesiosaur from the Tropic Shale, Dolichorhynchops tropicensis, which adds to the known biodiversity from this time period. The identification is based on two specimens, a well-preserved, nearly complete skeleton including the skull and an additional specimen with only fragmentary skeletal elements. The material shares several synapomorphic characters with Dolichorhynchops, including the shape of the temporal fenestrae, the shape of the sagittal crest, the trend of the ectopterygoids, the morphology of the teeth, and the number of teeth within the mandibular symphysis. D. tropicensis differs from other species of Dolichorhynchops in a greater size range of the teeth, moderately constricted dorsal vertebrae, angled anterior processes on the coracoids, and the presence of well-defined facets on the propodials. The specimens are of Early Turonian age (based primarily on ammonite biostratigraphy and bentonite marker beds), and their discovery extends the known stratigraphic range for Dolichorhynchops back by approximately seven million years. This suggests that Dolichorhynchops tropicensis and Trinacromerum bentonianum, a closely related polycotylid, co-existed in the Western Interior Seaway. 相似文献
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The systematics of elasmosaurids are not well resolved partially because of the scarcity of well-preserved skull material. Among Weddellian elasmosaurids, one exception to this is the holotype of Tuarangisaurus keyesi from upper Campanian–lower Maastrichtian levels of the Maungataniwha Sandstone Member of the Tahora Formation, Mangahouanga Stream, inland Hawke's Bay, New Zealand. The material preserves an almost complete cranium and mandible. This material is re-described here, based on new observations and digital reconstruction of the internal anatomy. The result adds one new autapomorphy to the diagnosis of the taxon: ectopterygoid with large boss on the ventral surface and a posteriorly directed process. Additionally new features are recorded: presence of stapes; pterygoid overlapping the vomer, medial sulcus on the dorsal surface of the vomer, parasphenoid–basisphenoid complex covering the ventral surface of the body of the basioccipital. The presence of stapes in Tuarangisaurus keyesi is surprising as they were previously considered absent among elasmosaurids. Phylogenetic analysis indicates a clear elasmosaurid affinity, although there is low resolution within the Elasmosauridae. The previously proposed close relationship with the early Aptian Callawayasaurus colombiensis is rejected. 相似文献
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S.P. Bennett P.M. Barrett M.E. Collinson S. Moore-Fay P.G. Davis C.P. Palmer 《Proceedings of the Geologists' Association. Geologists' Association》2012,123(1):146-154
This paper describes a new ichthyosaur specimen from the Stonebarrow Marls Member (Charmouth Mudstone Formation) of Charmouth, Dorset. It is substantially complete and various characters of the skull (snout ratios and basioccipital morphology) and appendicular skeleton (the number of digits on the hind paddle, which can be used to infer the number of digits on the front paddle) indicate that the specimen is referable to Ichthyosaurus communis. This provides the first direct evidence for the presence of I. communis in the Pliensbachian, extending the stratigraphic range of the species. The specimen also preserves a probable gullet content, which demonstrates that fish formed part of the diet of this individual. 相似文献
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根据发现于辽西朝阳九佛堂组一具有部分头骨的不完整骨架,命名一新的帆翼龙--赵氏龙城翼龙(新属新种).头骨和牙齿形态显示龙城翼龙与宽齿帆翼龙的关系要比与其他翼龙的关系密切,龙城翼龙是目前辽西九佛堂组中发现的唯一的帆翼龙类.帆翼龙科原仅一属种,发现于英国早白垩世地层中,辽西早白垩世新帆翼龙的发现,不但扩大了其地理分布,也证明辽西是晚侏罗世至早白垩世翼龙辐射、分异的重要地区.这对研究翼龙的起源和演化有重要意义. 相似文献
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Four new genera and five new species of Archegocimicidae are described from the Lower Cretaceous of England: namely Mortalia martini gen. et sp. nov., Tyrion lannister gen. et sp. nov., T. cersei sp. nov., Stannis baratheon gen et sp. nov., Daenerys khaleesi gen. et sp. nov. A transitional position of the complex between Asian Late Jurassic and Early Cretaceous entomofaunas is indicated by this new material. 相似文献
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新铺龙是产于我国西南地区的海龙目的一个属,最早发现于贵州省关岭晚三叠世卡尼期小凹组地层。虽然新铺龙材料众多,但其未成年个体骨骼学特征及个体发育模式并未被详细地记述。本文报道了一个保存完好的孙氏新铺龙未成年标本,是目前发现最小的完整的孙氏新铺龙个体,标本产自贵州省关岭县新铺乡。依据其体长小于正型标本的一半,头骨长度和躯干比值大,头骨骨缝清晰等特征,判断该标本为孙氏新铺龙的幼年个体。新标本补充了孙氏新铺龙幼年个体的形态学信息,如上颞骨内侧突短于外侧突,翼骨无牙齿发育,神经弓与椎体未融合,尺骨两端微凸,五边形坐骨等。此外,结合已发表的新铺龙标本数据进行统计分析发现,该物种吻部相对头部呈负异速生长,证明了吻部占头骨比例的大小属于新铺龙种内个体发育差异,而并非之前所认为的新铺龙种间差异。 相似文献
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A new species of hybodontid shark is described for the Lower Cretaceous of Brazil. The type specimen is derived from pre-Aptian strata of Malhada Vermelha Formation, Lima Campos Basin (Ceará State, northeastern Brazil), with referred material from the Missão Velha Formation of Araripe Basin, northeastern Brazil. The new taxon differs from other Planohybodus species by the presence of a stronger tooth ornamentation characterized by simple, usually non-branching cristae that nearly reach the apex of the main cusp as well as distinctly divergent lateral cusplets. In addition, certain North-American specimens formerly attributed to Hybodus are identified as Planohybodus. The new species presented herein, in addition to the North-American remains, represents an important contribution to the knowledge of the distribution of Planohybodus, expanding the geographic range of the genus to South and North America in the Late Jurassic and Early Cretaceous. 相似文献
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A Corythosaurus skull (UALVP 13) was collected in 1920 from what is now Dinosaur Provincial Park in southern Alberta, Canada and was designated the holotype of Corythosaurus excavatus, a new hadrosaur species by Gilmore (1923). In 1992, a previously uncovered, weathered, headless skeleton was found. This was collected in 2012 by the University of Alberta as there was potential it could belong to the holotype skull. In addition, an isolated hadrosaur dentary had been found in 1992 close to the articulated postcranial skeleton and may be one of the missing jaws of Corythosaurus excavatus. The hypothesis that it may be the skeleton of the holotype of Corythosaurus excavatus is tested using anatomical information and statistical analyses. Statistical comparisons suggest it is possible that the skull and dentary belong to the same specimen. Furthermore, bivariate plots and percentage prediction errors also indicate that the postcranial material could belong to the UALVP 13 skull. Because many large vertebrate fossil specimens (even types) are not always collected in their entirety, this method may be used as a line of evidence to determine whether independently collected specimens potentially belong to the same individual. The problems described herein highlight the need to collect specimens in their entirety, and for good field documentation, including the spatial and stratigraphic context of all finds. 相似文献
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Jean-Paul Billon-Bruyat Jean-Michel Mazin Joane Pouech 《Swiss Journal of Geoscience》2010,103(2):143-153
The spotty nature of the terrestrial fossil record for the Mesozoic hinders a more complete understanding of dinosaur diversity. For stegosaurs (Ornithischia), the plated dinosaurs, only a few and fragmentary remains are reported from the Early Cretaceous of Europe. A recent revision concluded that only a partial vertebra of the nomen dubium Craterosaurus (?Aptian, England) could be considered as stegosaurian. Here we report on a stegosaur tooth from the Early Cretaceous (Berriasian) Purbeckian deposits of Cherves-de-Cognac (Charente), southwestern France. This tiny tooth was examined in detail using microtomography. Comparisons being limited by the rarity of stegosaur tooth rows material (e.g., from the skull of the holotype of Stegosaurus stenops) and dental material, notably from Europe, we observed new material of cf. Stegosaurus armatus and Hesperosaurus mjosi from the Upper Jurassic Morrison Formation of Wyoming (USA). The tooth shows the most similarities to the Late Jurassic genera Stegosaurus and Hesperosaurus, but differs in having a distinctive downwardly arched (V-shaped) cingulum on the ?lingual face (maxillary tooth hypothesis). It is referred to as Stegosauria indeterminate, a medium-sized quadrupedal herbivore that inhabited an emerged land between the Armorican Massif and the Massif Central. This finding is the first evidence of a stegosaur from the Early Cretaceous of France and a welcome addition to the meagre European record of that time. In addition, it is the second stegosaurian tooth crown reported from Europe. The assemblage of ornithischians of Cherves-de-Cognac shares some similarities with that of the Early Cretaceous (Berriasian) of the Purbeck Limestone Group, southern England. The relative rarity of ornithischian osteological remains in both Purbeckian environments suggests that most of these dinosaurs were mainly inhabitants of inland terrestrial palaeoenvironments. 相似文献
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A New Troodontid Dinosaur from the Lower Cretaceous Yixian Formation of Liaoning Province, China 总被引：1，自引：1，他引：0
SHEN Caizhi Lü Junchang LIU Sizhao Martin KUNDRáT Stephen L. BRUSATTE GAO Hailong 《《地质学报》英文版》2017,91(3):763-780
A new troodontid dinosaur,Daliansaurus liaoningensis gen.et sp.nov.,is erected based on a nearly complete specimen from the Lower Cretaceous Yixian Formation of Beipiao City,Liaoning Province,China.This well preserved skeleton provides important new details of the anatomy for Liaoning troodontids,and gives new insight into their phylogenetic relationships and evolution.Daliansaurus is distinguished from other troodontids by an enlarged ungual on pedal digit IV,which is approximately the same size as the sickle-shaped second ungual,and is differentiated from other Liaoning troodontids by a number of characters of the skull,manus,pelvis,and hindlimb.A phylogenetic analysis recovers Daliansaurus within a subclade of Liaoning troodontids that also includes Sinovenator,Sinusonasus,and Mei.We erect a name for this group—Sinovenatorinae—and argue that it reflects a localized radiation of small-bodied troodontids in the Early Cretaceous of eastern Asia,similar to previously recognized radiations of Liaoning dromaeosaurids and avialans.As more Liaoning theropods are discovered,it is becoming apparent that small,feathered paravians were particularly diverse during the Early Cretaceous,and future work is needed to clarify how this diversity arose,which species coexisted,and how these numerous species partitioned niches. 相似文献