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1.
Leymeriellid ammonite faunas are described from northeast of Esfahan and the Khur area (Central Iran). The faunas comprise Leymeriella (L.) tardefurcata, L. (L.) germanica and L. (L.) acuticostata? as well as L. (Neoleymeriella) regularis, L. (N.) diabola and L. (Neoleymeriella) pseudoregularis?. The leymeriellids of Iran are closely related to faunas from northwest Europe, the Vocontian Basin and Transcaspia (Mangyschlak) and thus allow for biostratigraphic correlations, indicating the earliest Albian Leymeriella tardefurcata Zone with its superimposed L. acuticostata and L. regularis subzones. We suggest that the sudden appearance of common representatives of the genus Leymeriella in the northern and Central Iranian basins (Leymeriella acme) can be used as a useful proxy marker to define the base of the Albian Stage in Iran.  相似文献   

2.
We re-define the Cretaceous bony fish genus Rhinconichthys by re-describing the type species, R. taylori, and defining two new species; R. purgatorensis sp. nov. from the lowermost Carlile Shale (middle Turonian), southeastern Colorado, United States, and R. uyenoi sp. nov. from the Mikasa Formation (Cenomanian), Middle Yezo Group, Hokkaido, Japan. Rhinconichthys purgatoirensis sp. nov. is designated on a newly discovered specimen consisting of a nearly complete skull with pectoral elements. Only known previously by two Cenomanian age specimens from England and Japan, the North American specimen significantly extends the geographic and stratigraphic range of Rhinconichthys. The skull of Rhinconichthys is elongate, including an expansive gill basket, and estimated maximum body length ranges between 2.0 and 2.7 m. Rhinconichthys was likely an obligate suspension-feeder due to its derived cranial morphology, characterized by a remarkably large and elongate hyomandibula. The hyomandibula mechanically acts as a lever to thrust the jaw articulation and hyoid arch both ventrally and anterolaterally during protraction, thus creating a massive buccal space to maximize filtering of planktonic prey items. Cladistic analysis supports a monophyly of suspension-feeding pachycormids including Rhinconichthys, but further resolution within this clade will require more information through additional fossil specimens.  相似文献   

3.
4.
Early and early middle Cenomanian turrilitid ammonoids from the upper part of the Albian–Cenomanian Aitamir Formation of the Koppeh Dagh (northeast Iran) are described, illustrated and placed in an integrated stratigraphic context. The Aitamir Formation represents a graded siliciclastic shelf system and the turrilitid faunas comprise eight different species. Representatives of Mariella and Hypoturrilites have been recorded from a lower Cenomanian horizon in the Ghorghoreh section, corresponding to the Mantelliceras mantelli ammonite biozone. Mariella bicarinata (Kner, 1852) and H. wiedmanni Collignon, 1964 are recorded for the first time from Iran and the larger palaeobiogeographical area, respectively. The horizons with Turrilites costatus and T. scheuchzerianus at Taherabad can be dated as early middle Cenomanian Acanthoceras rhotomagense Zone, T. costatus Subzone. Concentrations of ammonoids commonly occur above regressive sandstone units, at the transition into overlying deeper marine shale units and in their lower parts, respectively. Such intervals represent the early transgressive systems tract of depositional sequences and may thus be regarded as early transgressive, slightly condensed shell accumulations. The uppermost lower to lower middle Cenomanian succession at Taherabad is of striking bio-, event- and sequence stratigraphic similarity to contemporaneous sections in northwest Europe. It reflects deposition during falling and low sea-level associated with the latest early Cenomanian unconformity SB Ce 3 as well as the pulsating transgressive development during the early middle Cenomanian, including levels correlative to the arlesiensis, primus and Mid-Cenomanian events in northwest Europe.  相似文献   

5.
Garnet-bearing micaschists and paragneisses of the Yaounde Group in the Pan-African Central African Orogenic Belt in Cameroon underwent a polyphase structural evolution with the deformation stages D1–D2, D3 and D4. The garnet-bearing assemblages crystallized in course of the deformation stage D1–D2 which led to the formation of the regional main foliation S2. In XCaXMg coordinates one can distinguish several zonation trends in the garnet porphyroblasts. Zonation trends with increasing XMg and variably decreasing XCa signalize a garnet growth during prograde metamorphism. Intermineral microstructures provided criteria for local equilibria and a structurally controlled application of geothermobarometers based on cation exchange and net transfer reactions. The syndeformational PT path sections calculated from cores and rims of garnets in individual samples partly overlap and align along clockwise PT trends. The PT evolution started at ~450 °C/7 kbar, passed high-pressure conditions at 11–12 kbar at variable temperatures (600–700 °C) and involved a marked decompression toward 6–7 kbar at high temperatures (700–750 °C). Th–U–Pb dating of metamorphic monazite by electron microprobe (EMP-CHIME method) in eight samples revealed a single period of crystallization between 613 ± 33 Ma and 586 ± 15 Ma. The EMP-monazite age populations between 613 ± 33 Ma enclosed in garnet and 605 ± 12 Ma in the matrix apparently bracket the high temperature–intermediate pressure stage at the end of the prograde PT path. The younger monazites crystallized still at amphibolite-facies conditions during subsequent retrogression. The Pan-African overall clockwise PT evolution in the Yaounde Group with its syndeformational high pressure stages and marked pressure variations is typical of the parts of orogens which underwent contractional crustal thickening by stacking of nappe units during continental collision and/or during subduction-related accretionary processes.  相似文献   

6.
Late Albian ammonite faunas from the Aitamir Formation of the Koppeh Dagh Basin in northeast Iran are described and illustrated. These comprise 14 taxa, several of which are recorded from Iran for the first time, namely Anahoplites planus (formerly recorded from central Iran in open nomenclature only), Semenoviceras solidus, Epihoplites (Metaclavites) iphitus, Hysteroceras orbignyi and Pseudhelicoceras robertianum. New records of Placenticeras grossouvrei extend the stratigraphic range of this species downwards into the Late Albian; previously it was known from the Early and Middle Cenomanian only. The record of the rare E. (M.) iphitus fills a palaeobiogeographic gap between Crimea and Tajikistan, and the holotype of Spath is re-illustrated here. Additionally, Epihoplites trapezoidalis, from the Late Albian of Tajikistan, is relegated into the synonymy of Spath's species. A large number of taxa typical of the Late Albian (upper part of the Gault Clay Formation) of northwest Europe indicate close palaeobiogeographic affinities with the Koppeh Dagh Basin and faunal exchange across the Russian Platform and Transcaspia. The stratigraphic succession of the ammonite faunas is used for a biostratigraphic subdivision of the upper Aitamir Formation.  相似文献   

7.
8.
The Dalichai Formation with an age of Late Bajocian-Late Callovian was sampled in Central Alborz Mountains of northern Iran and studied for palynological, palaeobiogeographical and palynocorrelation purposes. Palynological studies revealed diverse and well-preserved dinoflagellate cyst assemblages and lead to identification of three zones i.e., Cribroperidiniumcrispum (Late Bajocian), Dichadogonyaulaxsellwoodii (Bathonian to Early Callovian) and Ctenidodiniumcontinuum (Early to Middle Callovian) Zones. Subzone a of the D. sellwoodii Zone (Early to Middle Bathonian) was also differentiated. This biozonation corresponds to those recognised in Northwest Europe. Furthermore, the ammonoid families recorded including Phylloceratidae, Oppeliidae, Reineckeiidae, Perisphinctidae, Haploceratidae, Parkinsoniidae and Sphaeroceratidae, which confirm the Late Bajocian to Late Callovian age, are quite similar to those of Northwest Europe and the northwestern Tethys. The close similarities of the dinoflagellate cyst assemblages and ammonite fauna of northern Iran with those of Northwest Europe and the northwestern Tethys during the Middle Jurassic indicate direct but episodic marine connection and faunal exchange between the two areas.  相似文献   

9.
Univariate and multivariate statistics were applied to analyse the morphometrical variability of 4920 upper cheek teeth (P4, M1 and M2) of cave bears from 123 geographical sites (180 samples) of different Pliocene – Pleistocene ages. The analysed specimens included those belonging to the big cave bears Ursus kudarensis, Udeningeri, Uspelaeus (three subspecies) and Ukanivetz (including Uingressus), as well as the small cave bear Urossicus. The information‐theoretical parameters (Shannon entropy and orderliness (Von Foerster, 1960: On self‐organizing systems and their environments. In Self‐Organizing Systems, 31–50. Pergamon Press, London) were used to estimate tooth diversity in different teeth, different taxa and in selected local chrono‐populations. Multivariate allometry coefficients (Klingenberg, 1996: Multivariate allometry. In Advances in Morphometrics, 23‐49. Plenum Press, New York) were used to describe the relationships of different ‘parts’ of a tooth and to compare allometric patterns amongst species or selected local samples. A multivariate analysis showed a significant overlap of the size/shape parameter ranges in deningeroid and spelaeoid bears within morphological spaces. Within the cave bear lineage, the Deninger's bear has the greatest morphological diversity index (entropy) of all the teeth overall, and the lowest diversity is observed in the final taxon of this lineage – Ukanivetz (=ingressus). The P4 and M2 diversity showed multidirectional correlations with elevation above sea level amongst several ‘local’ populations of Late Pleistocene cave bears. The morphological disparities between the studied taxa are in close agreement with the distances in the available schemes of genetic differentiation based on ancient mitochondrial DNA. The split of Ukudarensis and Udeningeri has a good bootstrap support, which corresponds to the hypothesis about their parallel evolution. The small cave bear Urossicus is placed between Uarctos and Udeningeri. The phylogenetic signal is more pronounced in the variability of teeth in comparison with other skeletal remains of cave bears (cranium, mandible, or metapodial bones).  相似文献   

10.
The footwall of the Main Central Thrust (MCT) Zone along the Bhagirathi valley comprises a wide zone of mylonitic quartzite and deep-level tectonites. The systematic variation of finite strain parameters (Es, k and v) in the mylonites indicates heterogeneous deformation, which is determined to vary between, simple shear and non-coaxial flattening type. In such a strain regime the outer boundary of the quartz clasts are no longer preserved thus leading to an error in finite strain measurement.In order to supplement the finite strain studies, Anisotropy of Magnetic Susceptibility (AMS) analyses were carried out on the mylonitic quartzites. A systematic variation in degree of anisotropy (P′) with distance from the MCT is documented and is interpreted to be tectonic in origin. Based on these results it is concluded that P′ can be used as a strain-intensity gauge at least on an outcrop scale, where a systematic variation in P′ values from one part of the outcrop to the other can be established. However, the quantitative relation between principal axes of finite strain ellipsoid and AMS axes, magnitude of principal susceptibility difference (ΔK1 and ΔK3) and finite strain magnitude (ε1=ln 1 + e1 and ε3=ln 1 + e3) were related by a logarithmic relationship with a correlation coefficient of 0.844.  相似文献   

11.
A new teleosaurid from the Lower Cretaceous of Tataouine (Tunisia), Machimosaurus rex sp. nov., definitively falsifies that these crocodylomorphs faced extinction at the end of the Jurassic. Phylogenetic analysis supports its placement closer to M. hugii and M. mosae than M. buffetauti. With the skull length up to 160 cm and an estimated body length of 10 m, M. rex results the largest known thalattosuchian, and the largest known crocodylomorph at its time. This giant thallatosuchian probably was an ambush predator in the lagoonal environments that characterized the Tethyan margin of Africa during the earliest Cretaceous. Whether the Jurassic-Cretaceous mass extinction was real or artefact is debated. The discovery of M. rex supports that the end-Jurassic crisis affected primarily Laurasian biota and its purported magnitude is most likely biased by the incomplete Gondwanan fossil record. The faunal turnovers during the J-K transition are likely interpreted as local extinction events, triggered by regional ecological factors, and survival of widely-distributed and eurytypic forms by means of habitat tracking.  相似文献   

12.
Three species of a canaliculated rudist Caprinula d'Orbigny, 1847, C. sharpei (Choffat, 1885), C. cedrorum (Blanckenhorn, 1890) and C. cf. boissyi d'Orbigny, 1840 and a radiolitid Sauvagesia sharpei (Bayle, 1857) are described from the Hummar Formation (upper Cenomanian) in NW Jordan, in the vicinity of Ajlun. Caprinula sharpei, C. cedrorum and S. sharpei are described for the first time from Jordan. Many specimens of S. sharpei are characterized by the presence of cavities flanking the lamellar myophores in the left valve and the apparence of the dorsal cavity and teeth/socket system moulds in the inner part of the outer shell layer of the right valve. A hiatus (or erosional unconformity) between Hummar Formation and upper Turonian Wadi As Sir Limestone Formation is suggested by the presence of karstic structures, reworked limestone clasts, and rudist fragments and a sharp boundary. Early diagenetic processes such as dissolution and silicification present in the loose rudist material is described.  相似文献   

13.
Dollodon bampingi was recently named based upon a specimen from the Bernissart Quarry that had previously been referred to Mantellisaurus atherfieldensis. The initial diagnosis of Dollodon did not adequately distinguish it from Mantellisaurus or from other basal iguanodonts, necessitating a reassessment of the material. Firsthand examination of the holotypes of the two taxa and numerous other basal iguanodont specimens, as well as a principal components analysis of basal iguanodont dentaries, did not find any morphological features to justify the distinction of Dollodon from Mantellisaurus. D. bampingi is thus best considered a junior synonym of M. atherfieldensis. Furthermore, the recent referral of the species Iguanodon seelyi to the genus Dollodon is not supported; I. seelyi is indistinguishable from Iguanodon bernissartensis, and is considered a junior synonym of that species. Finally, the recently named taxon Proplanicoxa galtoni, also based upon a specimen formerly attributed to M. atherfieldensis, is considered a nomen dubium and probable junior synonym of M. atherfieldensis. Thus, only two species of large-bodied basal iguanodont should be recognized from the Barremian-Aptian of England and Belgium: M. atherfieldensis and I. bernissartensis.  相似文献   

14.
This research is based on the application of stream sediments to mineral exploration. Identifying the geochemical anomalies from background is a fundamental task in exploration geochemistry. This paper applied the element concentration–area (CA) model, to separating the geochemical anomalies from background based on a fractal approach and for the compilation of geochemical mapping from stream sediment samples (n = 620) of the Ahar region (Iran), where some Cu mineralization occurs. Comparisons of the known copper occurrences against the anomalous area created using thresholds from CA method illustrate these hits. All of known Cu mineralizations and moreover defines two extra Cu anomaly districts. Additional sampling (n = 186) around new Cu anomaly confirms this anomaly within the district.  相似文献   

15.
Seven specimens of fossil scorpionflies (Mecoptera) not assignable to any known family were discovered in the Wealden Supergroup (Lower Cretaceous) of southern England. They were found at Rudgwick Brickworks, West Sussex and Smokejacks Brickworks, Surrey and came from the Upper Weald Clay Formation, dated as Barremian (∼129.4–125 Ma). A new family – Englathaumatidae fam. nov., new genus – Englathauma gen. nov. and two new species E. crabbi sp. nov. and E. mellishae sp. nov. are described. A discussion of systematic position of these new taxa within the order Mecoptera is given. Englathaumatidae fam. nov. has been a nomen nudum since the year 2002, due to the first author's untimely death.  相似文献   

16.
In this paper we describe previously unpublished trionychid turtle material, consisting of skull fragments, from the Late Cretaceous (late Turonian) Bissekty Formation of the Dzharakuduk locality in Uzbekistan. This material is assigned to two taxa: the skull-based Khunnuchelys kizylkumensis Brinkman et al. (1993, Can. J. Earth Sci. 30, 2214-2223) and Trionychini indet. Two specimens which cannot be confidently attributed to these two taxa are considered Trionychidae indet. In addition to these trionychid taxa known from skulls, the Dzharakuduk turtle assemblage includes at least two shell-based taxa, Aspideretoides cf. A. riabinini and “Trionyx” cf. “T.” kansaiensis. For this and other Late Cretaceous localities of Middle Asia and Kazakhstan, we suggest the probable skull-shell associations of Khunnuchelys spp. with “Trionyxkansaiensis-like forms and Trionychini indet. with Aspideretoides-like forms.  相似文献   

17.
Pachycephalosaurs, a group of ornithischian dinosaurs with distinctive cranial ornamentation and skull domes, underwent dramatic changes in cranial morphology during ontogeny. This has caused debate about whether some specimens belong to juveniles or adults, which impacts studies of pachycephalosaur phylogeny and evolution. One such debate concerns a small skull roof specimen from the Campanian (Upper Cretaceous) of New Mexico, NMMNH P-33898, which was originally described as an indeterminate juvenile but later regarded as a mature adult and erected as the holotype of a new small-bodied species, Stegoceras novomexicanum. We restudied NMMNH P-33898 using computed tomography scanning, morphometric and phylogenetic analyses, and comparisons to growth series of other pachycephalosaurs (Stegoceras validum, Pachycephalosaurus wyomingensis). We conclude that two purported paratype specimens of Stegoceras novomexicanum cannot be referred to the same taxon as the holotype, that the holotype and controversial paratypes all belong to immature specimens and not aberrant small-bodied adults, but that current evidence cannot clearly determine whether NMMNH P-33898 is a juvenile belonging to its own diagnostic species (S. novomexicanum) or is a juvenile of Stegoceras validum, Sphaerotholus goodwini, or another known taxon. We review the pachycephalosaur record of New Mexico and demonstrate that pachycephalosaurs were important components of dinosaur faunas in the southern part of Western North America during the ∼15 million years before the end-Cretaceous extinction, just as they were in roughly contemporaneous northern localities.  相似文献   

18.
In Mexico, just 54% of the reported Pleistocene Bison material has been identified to species. Current paleontological research in northwestern Oaxaca, southern Mexico, has allowed collection of several specimens of Bison antiquus that are part of the Viko Vijin Local Fauna. B. antiquus had a very wide geographic distribution, from lowlands to mountainous landscapes of North and Central America. The B. antiquus record from southern Mexico links their former records from central Mexico and middle Central America and confirms this wide geographic distribution. The univariate mesowear score of the B. antiquus specimens from Oaxaca is in the lower extreme of grazers and the upper end of mixed-feeders, suggesting that they had a less abrasive diet than the modern plains Bison, as has been observed in other samples of this species from diverse parts of North America. The presence of B. antiquus in the Viko Vijin L. F. constrains the age of this fossil assemblage within a range from 60 Ka to 11.7 Ka.  相似文献   

19.
This study focuses on the morphometry and taxonomy of the Late Cretaceous coccolith genus Arkhangelskiella. Sixty samples from the Campanian–Maastrichtian interval of DSDP Hole 390A (Blake Nose) were investigated for their contents of Arkhangelskiella spp. In each sample one hundred specimens of Arkhangelskiella spp. were examined by measuring the coccolith length and width, as well as the length and width of the central area. In the samples investigated the Arkhangelskiella group exhibits a large size variation, specimens length varies from 4.95 μm to 14.52 μm. Former taxonomic concepts, based on morphometry, subdivided the Arkhangelskiella group into three species: Arkhangelskiella maastrichtiana, Arkhangelskiella confusa and Arkhangelskiella cymbiformis. Our data show a large variability of the morphometric data (coccolith length, width of the outer rim). There is no indication for three independant species; two of the quoted taxa (1. thick outer rim = Arkhangelskiella maastrichtiana; 2. very thin outer rim = Arkkhangelskiella cymbiformis) seem to be extreme forms of a continuous morphometric lineage. The lower part of the investigated succession (139.92–133.42 mbsf) is dominated by small specimens with an average length of 6.8 μm whereas the upper part (132.86–126.15 mbsf) is characterized by larger specimens (mean 8.7 μm). In DSDP Hole 390A the size increase appears to be very abrupt, within two samples (samples 133.42 mbsf, 132.86 mbsf) the mean size increases by 1.51 μm. Previous morphometric studies of Arkhangelskiella indicate a more continuous size increase throughout the late Campanian–Maastrichtian. The abrupt size increase observed here hints toward a minor hiatus in DSDP Hole 390A separating upper Campanian from lower Maastrichtian sediments. It seems likely that the size increase of Arkhangelskiella reflects changes of various environmental factors like nutrient supply and sea water chemistry (Mg/Ca ratio; Ca concentration). A comparison of morphometric results with previous palaeoecological studies documents a nutrient control for the growth of Arkhangelskiella. Small specimens can be related to more mesotrophic conditions whereas large specimens are linked to oligotrophic surface waters.  相似文献   

20.
The toxodont megaherbivores Toxodon and Mixotoxodon were endemic to South and Central America during the late Quaternary. Isotopic signatures of 47 toxodont teeth were analyzed to reconstruct diet and ancient habitat. Tooth enamel carbon isotope data from six regions of South and Central America indicate significant differences in toxodont diet and local vegetation during the late Quaternary. Toxodonts ranged ecologically from C3 forest browsers in the Amazon (mean δ13C = −13.4‰), to mixed C3 grazers and/or browsers living either in C3 grasslands, or mixed C3 forested and grassland habitats in Honduras (mean δ13C = −9.3‰), Buenos Aires province, Argentina (δ13C = −8.7‰), and Bahia, Brazil (mean δ13C = −8.6‰), to predominantly C4 grazers in northern Argentina (δ13C = −4.4‰), to specialized C4 grazers in the Chaco of Bolivia (δ13C = −0.1‰). Although these toxodonts had very high-crowned teeth classically interpreted for grazing, the isotopic data indicate that these megaherbivores had the evolutionary capacity to feed on a variety of dominant local vegetation. In the ancient Amazon region, carbon isotope data for the toxodonts indicate a C3-based tropical rainforest habitat with no evidence for grasslands as would be predicted from the Neotropical forest refugia hypothesis.  相似文献   

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