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1.
Nineteenth-century references to clavate borings in woody substrates in the Lower Greensand of the Isle of Wight used a variety of names, but Teredo (a wood-boring bivalve, not a boring), Teredolithes (a junior synonym of Teredolites) and Gastrochaena (a bivalve borer of rock and shelly substrates, not a boring in wood) are all nomenclatorially incorrect. Borings in a beach clast derived from the Lower Greensand Group and recently collected from Sandown Bay, Isle of Wight, are referred to Teredolites isp. cf. T. longissimus Kelly and Bromley. This specimen confirms the presence of Teredolites in the Lower Greensand Group and demonstrates a common ichnological problem of beach clasts; borings, either fossil or modern, are incompletely preserved, making confident classification below the level of ichnogenus problematic.  相似文献   

2.
Tylosaurus nepaeolicus (Cope, 1874), from the lower Smokey Hill Chalk upper Coniacian of Kansas is reassessed and compared to T. kansasensis Everhart, 2005, its sympatric contemporary from the same formation; both are compared to a later species from the upper Smoky Hill Chalk, T. proriger (Cope, 1869). Tylosaurus nepaeolicus (Cope, 1874) is virtually indistinguishable from T. kansasensis Everhart, 2005, and both show important similarities with T. proriger, particularly the smaller individuals of T. kansasensis. Many of the anatomical features of T. kansasensis are indicative of a juvenile stage based on comparisons to T. proriger. In addition to the aforementioned spatial and temporal sympatry between T. nepaeolicus and T. kansasensis, it is anatomically difficult to distinguish the two species from each other, with the few notable differences being ontogenetically variable, and possibly indicating allometric changes during post-embryonic development; in addition, T. nepaeolicus is known from fragmentary remains of very large individuals, while T. kansasensis is known from a small number of complete and recently collected skulls, though of much smaller size than the type materials of T. nepaeolicus. We suggest that T. kansasensis is a juvenile of Tylosaurus nepaeolicus, and is thus the junior synonym of the latter. Furthermore, we find ontogenetic evidence to suggest that T. proriger is likely a paedomorph of T. nepaeolicus, albeit, a gigantic one.  相似文献   

3.
The origin of the genus Bos is a debated issue. From ∼ 0.5 Ma until historic times, the genus is well known in the Eurasian large mammal assemblages, where it is represented by Bos primigenius. This species has a highly derived cranial anatomy that shows important morphological differences from other Plio-Pleistocene Eurasian genera of the tribe Bovini such as Leptobos, Bison, Proamphibos-Hemibos, and Bubalus. The oldest clear evidence of Bos is the skull fragment ASB-198-1 from the middle Pleistocene (∼ 0.6-0.8 Ma) site of Asbole (Lower Awash Valley, Ethiopia). The first appearance of Bos in Europe is at the site of Venosa-Notarchirico, Italy (∼ 0.5-0.6 Ma). Although the origin of Bos has traditionally been connected with Leptobos and Bison, after a detailed anatomical and morphometric study we propose here a different origin, connecting the middle Pleistocene Eurasian forms of B. primigenius with the African Late Pliocene and early Pleistocene large size member of the tribe Bovini Pelorovis sensu stricto. The dispersal of the Bos lineage in Western Europe during middle Pleistocene times seems to coincide with the arrival of the Acheulean tool technology in this continent.  相似文献   

4.
The species of the genus Apocricetus are considered to form the phyletic lineage A. aff. plinii (MN11)–A. plinii–A. alberti–A. barrierei–A. angustidens (MN16). Along this lineage, gradual morphological and biometrical changes occur, but not all the species are represented by rich populations. The assemblage of Apocricetus alberti from Venta del Moro is by far the most abundant collection of this species. This population shows a great morphological variability in some characters like the morphology of the anteroconid and the anterolophulids in m1 and the shape of the anterolophule in M1, with morphotypes that resemble both older and younger populations of Apocricetus. Along the phyletic lineage of Apocricetus, a change in the shape of m3 occurs, from predominantly subtriangular forms in the oldest populations to predominantly subrectangular in the youngest. To quantify this feature we use the posterior width of the molar and the anterior width/posterior width ratio, proposing five morphological categories, from extremely triangular to subrectangular.  相似文献   

5.
The taxonomic status of specimens referred to Tarchia kielanae, T. gigantea, and Minotaurasaurus ramachandrani has been contested. The holotype of T. kielanae, Zaklad Paleobiologii (Institute of Paleobiology)-Polish Academy of Sciences (ZPAL) MgD I/111, is redescribed. It has features common to both Palaeontological Institute-Russian Academy of Sciences (PIN) 3142/250, long considered Tarchia, and to M. ramachandrani but which are lacking in Saichania chulsanensis. The specimen PIN 3142/250 is not referable to Saichania but instead represents a new species, Tarchia teresae sp. nov. Similarly, the holotype skull of M. ramachandrani is not referable to T. kielanae, so Minotaurasaurus is a valid taxon. Tarchia is more derived than either Saichania or Minotaurasaurus while sharing features of both.  相似文献   

6.
Vegetation dynamics during the earliest part of the Holocene (11,250-10,250 cal yr BP) have been reconstructed from a lacustrine sequence on Sandoy, the Faroe Islands, using detailed plant macrofossil and pollen evidence. The plant macrofossils suggest the initial vegetation was sparse herb and shrub tundra, with Salix herbacea and open-ground species, followed by the development of a denser and more species-rich arctic heathland after 11,150 cal yr BP. Despite high pollen values for Betula nana, macrofossils are rare. The bulk of the macrofossils recorded are S. herbacea and Empetrum leaves with numerous herb taxa and an abundance of Racomitrium moss. Conditions start to change around 10,800 cal yr BP, with increased catchment erosion and sediment delivery to the lake from ca. 10,600 cal yr BP, and a transition to alternating Cyperaceae and Poaceae communities between ca. 10,450 and 10,250 cal yr BP. This vegetation change, which has been recorded throughout the Faroes, has previously been interpreted as a retrogressive shift from woody shrubs to a herbaceous community. The detailed plant macrofossil data show the shift is the replacement of an Empetrum arctic heathland by grassland and moist sedge communities. These taxa dominate the modern landscape.  相似文献   

7.
Indigofera melanadenia and Tephrosia longipes plant species, collected from Cu–Ni mining area, were evaluated for accumulation of Cu and Ni. The total and bioavailable concentrations of Cu and Ni in the host soils were also determined. Flame Atomic Absorption Spectrometry was used for all metal determinations. The total and bioavailable concentrations of Cu in the soils were in the range 900–9000 μg/g and 200–2000 μg/g respectively. For Ni, the total and bioavailable concentrations were in the range 900–2000 μg/g and ∼ 40–100 μg/g respectively. The concentrations of Cu and Ni in the leaves of I. melanadenia were higher than in the roots with a range 80–130 μg/g in the leaves and 20–80 μg/g in the roots for Cu and a range of 150–200 μg/g in the leaves and 20–60 μg/g in the roots for Ni. Concentration of Cu in T. longipes was in the range of 37–240 μg/g and 150–200 μg/g in the leaves and roots respectively while the concentration of Ni was 80–140 μg/g in the leaves and 25–100 μg/g in the roots. Results indicate that both species have a potential for accumulating Cu and Ni. Translocation factor, a ratio of shoots to roots metal concentration, was used to evaluate the translocation properties of the plants from roots to shoots. Translocation factors of the plants were ≥ 1 suggesting efficient translocation of metals from roots to shoots.  相似文献   

8.
A new partial skeleton of the armoured ornithischian dinosaur Polacanthus found in the Wadhurst Clay Formation (Valanginian stage) of Bexhill, Sussex is the oldest recorded occurrence of this taxon. Previous discoveries suggested that at least two armoured ornithischians occur in the Wealden succession: Polacanthus, which was mostly restricted to the Barremian, and Hylaeosaurus, which was recorded as present only in the Valanginian. The new discovery extends the stratigraphic range of Polacanthus into the Valanginian. Although these two taxa appear to be closely similar anatomically, their osteology now suggests they are not synonymous. The new specimen includes the first known jugal as well as a comparatively rare polacanthid plate/spine (splate) which probably comes from the shoulder (pectoral) area of these animals.  相似文献   

9.
The skeleton of a young prime adult cave bear, Ursus spelaeus, was found in Chiostraccio Cave (Siena, Tuscany, central Italy), only slightly buried under rock falls. The specimen was dated yielding a conventional age of 24,030 ± 100 14C yr BP (29,200–28,550 cal yr BP), which makes it the latest known representative of the species in Italy. The skeleton was accompanied by the remains of wolf (Canis lupus), wild boar (Sus scrofa), aurochs (Bos primigenius), red deer (Cervus elaphus), roe deer (Capreolus capreolus), bat (Vespertinus murinus), and crow (Corvus monedula). The site seems confirming that the latest Italian U. spelaeus populations shared the risk of intrusion. The association of the cave bear with other animals suggests that the assemblage is an attritional palimpsest of remains of different species not originally associated in life. Cave bears were probably more vegetarian than brown bears and possibly became extinct when plant productivity dropped at the onset of MIS 2. Central and southern Italy may have offered isolated and sheltered refugia for cave bears.  相似文献   

10.
Understanding late Holocene extinctions on islands requires accurate chronologies for all relevant events, including multiple colonisations by humans and the introduction of alien species. The most widely held hypothesis on the causes of Holocene island vertebrate extinctions incorporates human impacts, although climatic-related hypotheses cannot be excluded. Both hypotheses have been suggested to account for the extinction of the endemic Lava Mouse, Malpaisomys insularis from the Canary Islands. Here we present the first accelerator mass spectrometer (AMS) 14C ages from collagen of M. insularis bones from ancient owl pellets collected at Fuerteventura (Canary Islands, eastern Atlantic Ocean). These new dates contribute to an understanding of the extinction of this species. We are able to exclude climatic causes, predation by invasive species, and competition with the house mouse, Mus musculus. The arrival of Europeans in the Canary Islands correlates with the extinction of Malpaisomys. The introduction of rats, Rattus spp., together with their parasites and diseases, emerges as the most reasonable hypothesis explaining the extinction of M. insularis.  相似文献   

11.
The extent of racemization of aspartic acid (Asp) has been used to estimate the ages of 9 shells of the epifaunal calcitic brachiopod Bouchardia rosea and 9 shells of the infaunal aragonitic bivalve Semele casali. Both taxa were collected concurrently from the same sites at depths of 10 m and 30 m off the coast of Brazil. Asp D/L values show an excellent correlation with radiocarbon age at both sites and for both taxa (r2Site 9 B. rosea = 0.97, r2Site 1 B. rosea = 0.997, r2Site 9 S. casali = 0.9998, r2Site 1 S. casali = 0.93). The Asp ratios plotted against reservoir-corrected AMS radiocarbon ages over the time span of multiple millennia can thus be used to develop reliable and precise geochronologies not only for aragonitic mollusks (widely used for dating previously), but also for calcitic brachiopods. At each collection site, Bouchardia specimens display consistently higher D/L values than specimens of Semele. Thermal differences between sites are also notable and in agreement with theoretical expectations, as extents of racemization for both taxa are greater at the warmer, shallower site than at the cooler, deeper one. In late Holocene marine settings, concurrent time series of aragonitic and calcitic shells can be assembled using Asp racemization dating, and parallel multi-centennial to multi-millennial records can be developed simultaneously for multiple biomineral systems.  相似文献   

12.
Baeomorphinae Yoshimoto, 1975, based on Baeomorpha Brues, 1937, is transferred from Tetracampidae Förster, 1856 and recognized as a junior synonym of Rotoitidae Bouček and Noyes, 1987 (Hymenoptera: Chalcidoidea) syn. nov. following ICZN (1999) article 35.5. Inclusions in Taimyr amber (84–100 Ma) assigned to Baeomorpha are reviewed and the following eleven new species, all of Gumovsky, are described: B. avamica sp. nov., B. baikurenis sp. nov., B. bianellus sp. nov., B. caeleps sp. nov., B. gracilis sp. nov., B. ingens sp. nov., B. quattorduo sp. nov., B. quattoruno sp. nov., B. popovi sp. nov., B. yantardakh sp. nov., and B. zherikhini sp. nov. The recognized species are differentiated in separate keys to females and males and illustrated through microphotography. Two of four previously described Baeomorpha species from Campanian Canadian amber are synonymized: B. distincta Yoshimoto and B. elongata Yoshimoto under B. ovatata Yoshimoto (syn. nov.). One enigmatic rotoitid inclusion, which differs from Baeomorpha species in the possession of very short stigmal vein, is described as Taimyromorpha pusilla Gumovsky gen. et sp. nov. Inclusions containing specimens identified as Baeomorpha and Taimyromorpha are found in amber from Taimyr and Canada that originated from Laurasia, not Gondwana. Two Realms are newly proposed to recognize different Cretaceous faunal elements, a more northern Baeomorpha Realm that is characterized by a temperate or warm temperate climate and very abundant aphid fossils, and the Isoptera Realm, an opposing southward territory with a warmer climate and common termite but rare aphid fossils. The newly described fossils indicate the southern hemisphere distribution of extant Rotoitidae is relictual with the pattern observed being formed at least in part by extinction events, though distributions of the only two extant rotoitid genera, Rotoita Bouček and Noyes, 1987 (New Zealand) and Chiloe Gibson and Huber, 2000 (small area in the southern Chile) may have been more extensive in the past. Both of known regions of extant Rotoitidae have highly suppressed ant faunas, which may suggest that their survival there depended on low biocenotic pressure by ants, perhaps as low as is hypothesized for the Late Cretaceous. The Canadian amber genera Distylopus Yoshimoto, 1975 (Distylopinae) and Bouceklytus Yoshimoto, 1975 (Bouceklytinae) are excluded from Tetracampidae and regarded as Chalcidoidea incertae sedis.  相似文献   

13.
The palaeontological content (charophytes and vertebrates) of organic-rich layers from the Middle Jurassic (Bathonian) of the Grands Causses at Mostuéjouls (Aveyron, France) has been examined. It is rich in gyrogonites of Porochara douzensis. The vertebrates of the lower layer include a single hybodontiform tooth, “semionotiform” teeth and scales, and one pycnodontiform tooth whereas the upper layer has yielded one hybodontiform tooth, a variety of actinopterygian remains (mostly Caturus sp. and indeterminate pycnodontiforms) and a few possible reptile remains. The variation in fauna between the two layers is ascribed to a different degree of marine influence.  相似文献   

14.
Four juvenile specimens referable to Pinacosaurus grangeri (Ankylosauria: Dinosauria) are described from the Campanian (Upper Cretaceous) locality Bayan Mandahu in northern Inner Mongolia Autonomous Region (People’s Republic of China). All the specimens preserve the skulls as well as, in some cases, mandibles, postcrania, and osteoderm. They are not taphonomically deformed by expanding matrix distortion, unlike many Gobi specimens, including the holotype of P. grangeri. Bayan Mandahu is also the type locality for Pinacosaurus mephistocephalus. The proximity in space and time of these two closely related species warrants a generic and specific revision for Pinacosaurus. The distinction of the two species is based on characters of the squamosal dermal elaborations, cranial roof posterior to the orbits, premaxillary notch, and distal margin of the ilium. Although a relatively well-represented ankylosaur taxon, the phylogenetic position of Pinacosaurus has not been unequivocally resolved. A new analysis recovers Pinacosaurus as the most basal member of the Ankylosaurinae.  相似文献   

15.
The Corumbá Group, cropping out in the southern Paraguay Belt in Brazil, is one of the most complete Ediacaran sedimentary archives of palaeogeographic, climatic, biogeochemical and biotic evolution in southwestern Gondwana. The unit hosts a rich fossil record, including acritarchs, vendotaenids (Vendotaenia, Eoholynia), soft-bodied metazoans (Corumbella) and skeletal fossils (Cloudina, Titanotheca). The Tamengo Formation, made up mainly of limestones and marls, provides a rich bio- and chemostratigraphic record. Several outcrops, formerly assigned to the Cuiabá Group, are here included in the Tamengo Formation on the basis of lithological and chemostratigraphical criteria. High-resolution carbon isotopic analyses are reported for the Tamengo Formation, showing (from base to top): (1) a positive δ13C excursion to +4‰ PDB above post-glacial negative values, (2) a negative excursion to −3.5‰ associated with a marked regression and subsequent transgression, (3) a positive excursion to +5.5‰, and (4) a plateau characterized by δ13C around +3‰. A U-Pb SHRIMP zircon age of an ash bed interbedded in the upper part of the δ13C positive plateau yielded 543 ± 3 Ma, which is considered as the depositional age ( Babinski et al., 2008a). The positive plateau in the upper Tamengo Formation and the preceding positive excursion are ubiquitous features in several successions worldwide, including the Nama Group (Namibia), the Dengying Formation (South China) and the Nafun and Ara groups (Oman). This plateau is constrained between 542 and 551 Ma, thus consistent with the age of the upper Tamengo Formation. The negative excursion of the lower Tamengo Formation may be correlated to the Shuram–Wonoka negative anomaly, although δ13C values do not fall beyond −3.5‰ in the Brazilian sections. Sedimentary breccias occur just beneath this negative excursion in the lower Tamengo Formation. One possible interpretation of the origin of these breccias is a glacioeustatic sea-level fall, but a tectonic interpretation cannot be completely ruled out.  相似文献   

16.
Reworked fossils may be exotic, but more typically are locally derived. Echinoderms have only rarely been identified from beach clasts; most commonly, these are Upper Cretaceous echinoids from the Chalk. A pebble collected from a beach in Estonia has yielded a moderately well preserved specimen of the crinoid stem, Baltocrinus (col.) antiquus (Eichwald). This species is typical of the Baltic region, only being known from the Middle Ordovician (Darriwilian) of Estonia, but is exotic because the local outcrop is Upper Ordovician to Silurian.  相似文献   

17.
Charophytes from uppermost Campanian to Paleocene deposits in the Pingyi Basin (Shandong Province, Eastern China) are studied from the perspectives of taxonomy, paleoecology, biogeography, and biostratigraphy. The taxonomy of charophytes used by previous authors is revisited based on a study of intraspecific gyrogonite polymorphism, facilitating comparisons between China and Europe. A number of synonymies are proposed. Gobichara deserta is confirmed as a younger synonym of Microchara cristata. The genus Euaclistochara Z. Wang is shown to be a younger synonym of Lamprothamnium J. Groves. Charophyte assemblages from the Pingyi Basin were generally species-poor but showed a high degree of variation depending on the paleoenvironment. During the latest Campanian–Maastrichtian, brackish water assemblages were monospecific, formed by Feistiella anluensis. Permanent lakes were dominated by Microchara cristata and Peckichara praecursoria. Lamprothamnium ellipticum and Mesochara voluta inhabited the overbank ponds near braided rivers. The Paleocene was much more homogeneous and was dominated by Peckichara varians in permanent lakes. A new biozonation is proposed which encompasses two biozones based on species with broad paleoecological requirements and a Eurasiatic distribution. These are the Microchara cristata biozone starting in the latest Campanian and lasting at least until the earliest Danian and the Peckichara varians biozone encompassing the late Danian–earliest Eocene. These biozones allow direct correlation between Chinese and European basins.  相似文献   

18.
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19.
New records of Jefferson's ground sloth (Megalonyx jeffersonii) and elk-moose (Cervalces scotti) from Lang Farm provide the first precise temporal correlation of these taxa with the specific environments inhabited by them near the time of their extinction. Six AMS 14C measurements establish an age of 11,405 ± 50 14C yr B.P. for Lang Farm Cervalces and an age of 11,430 ± 60 or 11,485 ± 40 14C yr B.P. for the Megalonyx. These measurements represent the youngest 14C dates for these two genera based on direct dating. Comparison of the dates with pollen data from northern Illinois indicates that these species inhabited a nonanalog environment that was transitional from mid-latitude tundra to mixed conifer and deciduous woodland. Although spruce (Picea sp.) was dominant, it was less abundant than prior to 12,500 14C yr B.P. The presence of black ash (Fraxinus nigra) and fir (Abies sp.) indicates a wet climate and heavy winter precipitation. This may have been the preferred habitat for Cervalces because of its narrow geographic range. However, this habitat type was only one of many occupied by Megalonyx as indicated by its broad geographic distribution.  相似文献   

20.
A new chelid species (Prochelidella cerrobarcinae nov. sp.) are described from the Aptian-Albian? Puesto La Paloma Member, Cerro Barcino Formation, northern of Chubut Province, Argentina. The basal section of this member, which bears the turtle remains, is composed of tuffaceous mudstones with plane parallel lamination, asymmetrical ripples and a chert intercalation suggesting sub-aqueous deposition in a relatively shallow lacustrine environment. Pr. cerrobarcinae nov. sp. is represented by post-cranial remains of several specimens that not only represents the oldest pleurodiran chelid record in the world but, together with the chelid remains of Albian Lightning Ridge, New South Wales, Australia, indicates that chelid diversification began well before the final fragmentation of southern Gondwana.  相似文献   

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