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We describe two new species of extinct biting midges in the fossil genus Protoculicoides: Protoculicoides hispanicus Szadziewski and Arillo, sp. nov. and Protoculicodes sanjusti Szadziewski and Arillo, sp. nov., from Albian (101–113 Ma) amber of San Just, Spain. Atriculicoides Remm, 1976 is recognized as a new junior synonym of Protoculicoides Boesel, 1937 (NEW SYNONYM). Devalquia Choufani et al., 2013 is regarded here as a new junior synonym of the extant genus Culicoides Latreille, 1809 (NEW SYNONYM). Metahelea roggeroi Choufani et al., 2013 from Upper Cretaceous French amber, is excluded from the tribe Heteromyiini and placed in the extant genus Stilobezzia Kieffer, 1911, tribe Ceratopogonini: Stilobezzia roggeroi (Choufani et al., 2013), comb. nov. The extinct tribe Atriculicoidini Szadziewski, 1996 is regarded as a new subfamily, Atriculicoidinae, stat. nov., that includes the fossil genus Protoculicoides, with 13 species reported from Albian, Turonian, Cenomanian, Coniacian, Santonian and Campanian ambers (78–113 Ma). Protoculicoides skalskii Szadziewski & Arillo, 1998 from Lower Cretaceous amber of Álava, Spain, Protoculicoides succineus Szadziewski, 1966 from Lower Cretaceous Lebanese amber and Protoculicoides burmiticus Szadziewski & Poinar, 2005 from Upper Cretaceous Burmese amber are transferred to the extinct genus Archiculicoides Szadziewski, 1996, comb. nov. Protoculicoides krzeminskii Choufani et al., 2014 from Lower Cretaceous Lebanese amber is transferred to the extinct genus Archiaustroconops Szadziewski, 1996, comb. nov. The Cretaceous subfamily Atriculicoidinae forms an unresolved trichotomy with the extant subfamilies Forcipomyiinae and Dasyheleinae, both of which date to the Eocene. The fossil record indicates that wings with macrotrichia in biting midges evolved during the mid-Cretaceous greenhouse climate. We suggest that this was an evolutionary adaptation to new atmospheric conditions with higher levels of CO2 in order for the antennal Johnston's organs of males to receive the vibrational sex signals produced by females during flight.  相似文献   

3.
A new genus, Cretasyne gen. nov., in the family Lasiosynidae, with two species C. lata sp. nov. and C. longa sp. nov., is described from the Lower Cretaceous Yixian Formation of Inner Mongolia, China. These fossils are the first record of the family from the Yixian Formation and provide significant information on lasiosynid morphology and diversity. The fossil record of Lasiosynidae is also reviewed.  相似文献   

4.
Two new genera from the Lower Cretaceous Yixian Formation of Beipiao City, Liaoning Province, northeastern China are described and illustrated. Paracretocateres gen. nov. (type species P. bellus, sp. nov.) and Yixianteres gen. nov. (type species Y. beipiaoensis sp. nov.) are assigned to the subfamily Lophocaterinae based on the state of the procoxae which are not projecting, the transverse procoxal cavities, and the exposed protochantins. The new taxa broaden the diversity of this family and provide further morphological characters for phylogenetic studies of Trogossitidae. They also provide evidence that lophocaterines well-diversified prior to the trogossitines and their biologies may have been similar to extant forms.  相似文献   

5.
Seven specimens of fossil scorpionflies (Mecoptera) not assignable to any known family were discovered in the Wealden Supergroup (Lower Cretaceous) of southern England. They were found at Rudgwick Brickworks, West Sussex and Smokejacks Brickworks, Surrey and came from the Upper Weald Clay Formation, dated as Barremian (∼129.4–125 Ma). A new family – Englathaumatidae fam. nov., new genus – Englathauma gen. nov. and two new species E. crabbi sp. nov. and E. mellishae sp. nov. are described. A discussion of systematic position of these new taxa within the order Mecoptera is given. Englathaumatidae fam. nov. has been a nomen nudum since the year 2002, due to the first author's untimely death.  相似文献   

6.
Specimens of the Venicoridae (Heteroptera) occur in the Lower Cretaceous Yixian Formation in northeastern China. We describe a new genus and two new species Halonatusivena shii gen. et sp. nov. and Halonatusivena nervosus sp. nov. from this formation. The veins of the membrane are distinctive; the edges of veins are not simple but resemble an irregular stain, with a reduced vasiform structure previously unknown in the Heteroptera. Remarkably, the antennae of one of the specimens of Halonatusivena shii exhibit morphological abnormality. An identification key for the known species of the Venicoridae is provided.  相似文献   

7.
One new genus with three new species of the family Prionoglarididae, Palaeosiamoglaris leinhardi Azar, Huang et Nel gen. et sp. nov.; P. burmica Azar, Huang et Nel sp. nov. and P. inexpectata Azar, Huang et Nel sp. nov. are characterised, illustrated and described from Cretaceous Burmese amber. Palaeosiamoglaris gen. nov. share several characters with the recent genus Siamoglaris Lienhard, 2004, endemic to the Oriental (Sino-Indian) Region. A new tribe Siamoglaridini Azar, Huang et Nel trib. nov. is established to accommodate the group Siamoglaris + Palaeosiamoglaris; the other Prionoglaridinae being attributed to Prionoglaridini. This discovery implicates a palaeobiogeographic repartition for this particular tribe (Siamoglarini) that is still the same currently. Most diagnostic features show a nearly stasis state, supporting the notion that these characters appeared at least by the early Cenomanian. A checklist of all prionoglaridids is given.  相似文献   

8.
A new genus Barbderma gen. nov. with a new species, Barbderma oblonguata sp. nov., and a new species, Sinoprotodiplatys ellipsoideuata sp. nov., of the family Protodiplatyidae are described from the Lower Cretaceous Yixian Formation at the Huangbanjigou Village, Beipiao City, Liaoning Province, China. Both new species are assigned to Protodiplatyidae mainly based on their diagnostic characters of antenna, pronotum, tegmina, tarsi and the distinct long, slender, multi-segmented cerci. These findings of fossil Protodiplatyidae provide more evidence to confirm the existence of these basal earwigs in the Early Cretaceous. Key diagnostic characters for genera of Protodiplatyidae are compared to highlight the generic variations and similarities. Evolutionary trend of cerci morphology suggests that it is likely that the cerci were evolved from long, slender and segmented to the stout terminal forceps without segmentation.  相似文献   

9.
A new genus and species of Scoliidae (Hymenoptera), Sinoproscolia yangshuwanziensis gen. et sp. nov., is described from the Lower Cretaceous Yixian Formation in Inner Mongolia, China and attributed to the subfamily Proscoliinae. Some characteristics, including complete crossveins 1r-rs and 2r-m in the forewing, and a free section present in almost all longitudinal veins indicate a basal position of the new genus in Proscoliinae. The presence of vein 2A suggests its atavistic origin in Sinoproscolia gen. nov. rather than a true plesiomorphy.  相似文献   

10.
Although the Cretaceous is characterized by a rich fish diversity, Cretaceous continental fishes from Gondwana are poorly known and comparatively scarce. Among these fishes, the family Pleuropholidae is only known by a few species relatively poorly preserved, from the Middle Jurassic to Lower Cretaceous of Europe, Africa, North America, and South America. In this paper, two new species of the pleuropholid new genus Zurupleuropholis are described, Z. quijadensis gen. et sp. nov. and Z. decollavi gen. et sp. nov. The new fishes were recovered in the Lower Cretaceous lacustrine Lagarcito Formation of central-west Argentina. This taxon constitutes a relevant finding considering that the representation of the family Pleuropholidae is rare worldwide. Zurupleuropholis gen. nov. appears to be the youngest known member of Pleuropholidae, and it represents the second record of the family in South America and the first record in the Cretaceous of the continent.  相似文献   

11.
A new genus with a new species, Brachypelecinus euthyntus gen. et sp. nov., and two new species, Abropelecinus tytthus sp. nov. and Zoropelecinus periosus sp. nov., are described and figured from three exquisitely preserved pelecinid wasps in the Upper Cretaceous Myanmar (Burma) amber. These taxa, owing to their well-preserved characters in amber, provide a better understanding of morphological changes and relationships among the constituent groups, while further highlighting the diversity of Pelecinidae in the Mesozoic.  相似文献   

12.
One new genus and two new species of fossil Cerophytidae, Baissophytum convexus gen. et. sp. nov. and Baissophytum amplus sp. nov. are described and illustrated from Zaza Formation (Lower Cretaceous), Eravnensky (Bauntovsky) District, Buriatskay Autonomous Republic (Transbaikalia), Russia.  相似文献   

13.
A new species, Shoushida infera sp. nov. and a new genus with a new species, Stelepelecinus longus gen. et sp. nov., both in Pelecinidae, are described and illustrated. All specimens were collected from the Lower Cretaceous of Yixian Formation, Jehol Biota at Huangbanjigou, Beipiao City, western Liaoning Province, China. The forewing of Shoushida infera sp. nov. has a rudimentary “X” pattern, formed by veins of Rs, Rs1, Rs2 and 2r-rs, which is similar to but slightly different from that of Shoushida regilla Liu, Shih et Ren, 2009. Consistent with the phylogeny of Pelecinidae reported by Shih et al., 2010, this new species and S. regilla represent a transition between basal and crown pelecinids and the rudimentary “X” pattern evolved later to a more developed and robust “X” pattern in more derived pelecinids. In addition, the long petiole of the basalmost segment of metasoma, present in the male Stelepelecinus longus gen. et sp. nov., is documented for the first time, suggesting a likely Early Cretaceous origination of metasomal evolution leading to the thin and long petiole structure in the extant male Pelecinus thoracicus. We proposed two probable pathways of the transformation of the pelecinid male metasoma from the most plesiomorphic state to the more apomorphic states.  相似文献   

14.
New fossil genus with a new species, Stellularis longirostris Chen, Yao and Ren, gen. et sp. nov., from the family Procercopidae, are described. New specimens were collected from the Lower Cretaceous non-marine sedimentary strata from the Yixian Formations of northeast China. The exceptionally well-preserved materials reveal fine details of the body structure and morphology including rostrum, legs, abdomen, anteclypeus and ovipositor. The genera Luanpingia Hong, 1983 and Cathaycixius Ren, Lu, and Guo, 1995 are discussed.  相似文献   

15.
A new subfamily of Ichneumonidae, Novichneumoninae subfam. nov., is established based on two new genera with two new species: Novichneumon longus gen. et sp. nov. and Caloichneumon perrarus gen. et sp. nov. These two new species are the first ichneumonids described from the Upper Cretaceous Myanmar (Burmese) amber. A list of all described Mesozoic ichneumonid fossil species with their respective localities and ages is summarized. The distribution of Ichneumonoidea during the Cretaceous indicates that Cretaceous ichneumonids were documented from localities at high latitudes while braconids were distributed worldwide, a pattern consistent with the distribution of extant ichneumonids and braconids.  相似文献   

16.
A new fossil soldier beetle Myamalycocerus vitalii gen. et sp. nov., is described and illustrated from an inclusion in Upper Cretaceous Burmese amber (Myanmar). It differs from all extant genera that have elytra adorned with small striae by possessing raised punctation, very probably an ancient character as it is absent among current species. It differs from the fossil Ornatomalthinus Poinar et Fanti, by the long elytra and relief points which are less raised and more numerous. This new genus appears vaguely related (not necessarily phylogenetically) to the current genus Lycocerus Gorham.  相似文献   

17.
Two new genera and two new species of fossil Throscidae: Potergosoma gratiosa gen. et sp. nov. and Rhomboaspis laticollis gen. et sp. nov. are described from the Lower Cretaceous Lebanese amber and are compared with extant and extinct genera. The described amber inclusions are the oldest known representatives of the family Throscidae. Some hypotheses on the phylogeny of the family Throscidae and the position of it in the superfamily Elateroidea are discussed.  相似文献   

18.
Four new genera and five new species of Archegocimicidae are described from the Lower Cretaceous of England: namely Mortalia martini gen. et sp. nov., Tyrion lannister gen. et sp. nov., T. cersei sp. nov., Stannis baratheon gen et sp. nov., Daenerys khaleesi gen. et sp. nov. A transitional position of the complex between Asian Late Jurassic and Early Cretaceous entomofaunas is indicated by this new material.  相似文献   

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The Glandulariini (=Cyrtoscydmini) includes about 60 genera comprising over 70% of the extant species of Scydmaeninae. This successful group is today very common and abundant in forests of all continents. However, while extinct representatives of much smaller extant tribes, as Clidicini, Eutheiini and Cephenniini, were described from Cretaceous deposits, named genera and species of Glandulariini were so far known only from much younger Baltic, Sicilian and Dominican ambers. Here we report the first genus and species of definite Glandulariini from the Upper Cretaceous Burmese amber, Scydmobisetia vetutissima Jałoszyński and Yamamoto, gen. and sp. nov. Scydmobisetia is the first known glandulariine beetle with a pair of very long vertexal sensilla chaetica, previously not found in this tribe, but common among unrelated Staphylinidae. The new genus shows similarities to the extant genera Anthicimimus, Sciacharis, Spinosciacharis and Horaeomorphus, but S. vetutissima has protibial modifications typical of males of some extant species of Euconnus. We discuss the apparent disproportion in the oldest fossil record of various tribes of scydmaenines as a result of identification problems, and not the lack of known fossils.  相似文献   

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