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1.
Postcranial material of a crocodylomorph from the Danian of Oulad Abdoun Basin (Morocco) is described. Several characters, in particular the shape of its dorsal osteoderms, allows its attribution to Pholidosauridae. Up till now, the latest known pholidosaurid was Terminonaris, from the Early Turonian of North America, so, the Moroccan pholidosaurid extends the stratigraphic range of the group to more than 20 million years younger, and shows that the pholidosaurids survived the K-Pg crisis. The reevaluation and the phylogenetic analysis of Dakotasuchus kingi, Woodbinesuchus byersmauricei, and Sabinosuchus coahuilensis, previously considered as goniopholidids and dyrosaurid, respectively, revealed rather that they are pholidosaurids. This analysis also suggests that at least two independent pholidosaurid lineages reached the Maastrichtian, among which one crossed the K-Pg boundary. This study proposes the first analysis of tethysuchian diversity from the Late Jurassic to the Early Paleogene. Two diversity peaks are observed during the Oxfordian and Cenomanian, two stages of high paleotemperatures evaluated with the δ18O. The tethysuchian diversity strongly decreases after the Cenomanian, a decline that may be correlated with the “Ocean Anoxic Event” (OAE 2), which caused the strong marine faunal turnover during the Cenomanian-Turonian time interval. The large Turonian-Coniacian oceanic regression could also be a factor in the tethysuchian decline. After that, tethysuchian diversity remains low until the Maastrichtian-lower Paleocene marine tethysuchian dyrosaurid diversification, correlated with the Late Cretaceous-Paleocene transgression. At least one pholidosaurid lineage crossed the K-Pg boundary, confirming the weak impact of this crisis on the crocodylomorphs. Pholidosaurids seem to go extinct at the beginning of the Paleocene, possibly due to the strong radiation of dyrosaurids in marine environments and crocodylians in fresh-water during this time period.  相似文献   

2.
A new teleosaurid from the Lower Cretaceous of Tataouine (Tunisia), Machimosaurus rex sp. nov., definitively falsifies that these crocodylomorphs faced extinction at the end of the Jurassic. Phylogenetic analysis supports its placement closer to M. hugii and M. mosae than M. buffetauti. With the skull length up to 160 cm and an estimated body length of 10 m, M. rex results the largest known thalattosuchian, and the largest known crocodylomorph at its time. This giant thallatosuchian probably was an ambush predator in the lagoonal environments that characterized the Tethyan margin of Africa during the earliest Cretaceous. Whether the Jurassic-Cretaceous mass extinction was real or artefact is debated. The discovery of M. rex supports that the end-Jurassic crisis affected primarily Laurasian biota and its purported magnitude is most likely biased by the incomplete Gondwanan fossil record. The faunal turnovers during the J-K transition are likely interpreted as local extinction events, triggered by regional ecological factors, and survival of widely-distributed and eurytypic forms by means of habitat tracking.  相似文献   

3.
The latest Cretaceous continental vertebrate faunas of the wider Transylvanian area figured prominently in discussions concerning the Cretaceous–Paleogene Boundary (K-Pg) events when they were first described by Nopcsa between 1897 and 1929, because they were assumed to be late Maastrichtian in age. Subsequently their age was reconsidered as early Maastrichtian, and were thus regarded of lesser importance in understanding the K-Pg boundary events in Europe and worldwide. Moreover, Transylvanian continental vertebrate assemblages (the so-called ‘Haţeg Island’ faunas) were often lumped together as a temporally restricted assemblage with a homogenous taxonomic composition. Recent fossil discoveries and more precise dating techniques have considerably expanded knowledge of the Transylvanian vertebrate assemblages, their ages, and their evolution. A synthesis of the available stratigraphic data allows development of the first comprehensive chronostratigraphic framework of the latest Cretaceous Transylvanian vertebrates. According to these new data, expansion of continental habitats and emergence of their vertebrate faunas started locally during the latter part of the late Campanian, and these faunas continued up to the second half of the Maastrichtian. During this time, long-term faunal stasis appears to have characterized the Transylvanian vertebrate assemblages, which is different from the striking turnovers recorded in western Europe during the same time interval. This suggests that there was no single ‘Europe-wide’ pattern of latest Cretaceous continental vertebrate evolution. Together, the available data shows that dinosaurs and other vertebrates were relatively abundant and diverse until at least ca. 1 million years before the K-Pg boundary, and is therefore consistent with the hypothesis of a sudden extinction, although this must be tested with future discoveries and better age constraints and correlations.  相似文献   

4.
The Lechówka section comprises the most complete Cretaceous–Paleogene (K-Pg) boundary succession in Poland and is among 29 sites worldwide with the youngest ammonite record. Here, cephalopods (ammonites and nautilids), organic-walled dinoflagellates (dinocysts) and foraminifera from the uppermost Maastrichtian interval are studied. In terms of ammonite biostratigraphy, the upper Maastrichtian Hoploscaphites constrictus crassus Zone is documented up to a level 120 cm below the K-Pg boundary. There is no direct, ammonite-based evidence of the highest Maastrichtian H. constrictus johnjagti Zone. However, the predominance of the dinocyst marker taxon Palynodinium grallator suggests the presence of the equivalent of the uppermost Maastrichtian Thalassiphora pelagica Subzone, which is correlatable with the H. c. johnjagti ammonite Zone. The planktonic foraminiferal assemblage is coeval with that from the H. c. johnjagti Zone as well. These data indicate that the top of the Maastrichtian at Lechówka is complete within the limits of biostratigraphic resolution, albeit slightly condensed. The dinocyst and foraminiferal assemblages are dominated by taxa that are characteristic of high-energy, marginal marine environments. A reduction in test size among the calcareous epifaunal benthic foraminifera is observed at a level 50 cm below the K-Pg boundary, which is possibly related to environmental stress associated with Deccan volcanism.  相似文献   

5.
An integrated study of palynology, Mössbauer spectroscopy, mineralogy and osmium isotopes has led to the detection of the first K-Pg boundary clay layer in a Southern Hemisphere terrestrial setting. The K-Pg boundary layer was independently identified at centimetre resolution by all the above mentioned methods at the marine K-Pg boundary site of mid-Waipara and the terrestrial site of Compressor Creek (Greymouth coal field), New Zealand. Mössbauer spectroscopy shows an anomaly of Fe-containing particles in both K-Pg boundary sections: jarosite at mid-Waipara and goethite at Compressor Creek. This anomaly coincides with a turnover in vegetation indicated by an interval dominated by fern spores and extinction of key pollen species in both sections. In addition to the terrestrial floristic changes, the mid-Waipara section reveals a turnover in the dinoflagellate assemblages and the appearance of global earliest Danian index species. Geochemical data reveal relatively small iridium enrichments in the boundary layers of 321 pg/g at mid-Waipara and 176 pg/g at Compressor Creek. Unradiogenic 187Os/188Os values of the boundary clay reveal the presence of a significant extraterrestrial component. We interpret the accumulation of Fe nano-phases at the boundary as originating from both the impactor and the crystalline basement target rock. The goethite and jarosite are interpreted as secondary phases formed by weathering and diagenesis. The primary phases were probably controlled by the initial composition of the vapor plume and condensation kinetics rather than condensation thermodynamics. This investigation indicates that identification of Fe in nano-phases by Mössbauer spectroscopy is an accurate and cost-effective method for identifying impact event horizons and it efficiently complements widely used biostratigraphic and geochemical methods.  相似文献   

6.
Ostracode assemblages of the Poty quarry, Pernambuco-Paraíba Basin, northeastern Brazil, record an abrupt faunal change near the Cretaceous-Paleogene (K-Pg) boundary. Paleoecological and stable isotope analyses of ostracode carapaces were used to interpret the paleoenvironmental changes that took place from the Late Maastrichtian to the Early Danian. Ostracode distribution demonstrates substantial changes at and around the boundary with the last occurrences of most species and appearance of new ones. Stable isotope analyses carried out for four ostracode species distributed along the entire succession exhibit oscillations in the isotopic composition before the K-Pg boundary, thus suggesting that environmental changes may have begun slightly prior the boundary event itself. These changes may have triggered the extinctions that culminated at the K-Pg boundary, marked by positive δ18O and negative δ13C excursions. The K-Pg boundary is marked by cooling of bottom seawaters and a decrease in productivity, as well as the beginning of an extensive marine regression.  相似文献   

7.
A review of the tetrapod (amphibian and amniote) record across the Permo-Triassic boundary (PTB) indicates a global evolutionary turnover of tetrapods close to the PTB. There is also a within-Guadalupian tetrapod extinction here called the dinocephalian extinction event, probably of global extent. The dinocephalian extinction event is a late Wordian or early Capitanian extinction based on biostratigraphic data and magnetostratigraphy (the extinction precedes the Illawara reversal), so it is not synchronous with the end-Guadalupian marine extinction. The Russian PTB section documents two tetrapod extinction events, one just before the dinocephalian extinction event and the other at the base of the Lystrosaurus assemblage. However, generic diversity across the latter extinction remains essentially the same despite a total evolutionary turnover of tetrapod genera. The Chinese and South African sections document the stratigraphic overlap of Dicynodon and Lystrosaurus. In the Karoo basin, the lowest occurrence of Lystrosaurus is in a stratigraphic interval of reversed magnetic polarity, which indicates it predates the marine-defined PTB, so, as previously suggested by some workers, the lowest occurrence of Lystrosaurus cannot be used to identify the PTB in nonmarine strata. Correlation of the marine PTB section at Meishan, southern China, to the Karoo basin based primarily on magnetostratigraphy indicates that the main marine extinction preceded the PTB tetrapod extinction event. The ecological severity of the PTB tetrapod extinction event has generally been overstated, and the major change in tetrapod assemblages that took place across the PTB was the prolonged and complex “replacement” of therapsids by archosaurs that began before the end of the Permian and was not complete until well into the Triassic. The tetrapod extinctions are not synchronous with the major marine extinctions at the end of the Guadalupian and just before the end of the Permian, so the idea of catastrophic causes of synchronous PTB extinctions on land and sea should be reconsidered.  相似文献   

8.
The Bálvány North Permian-Triassic boundary sediments were deposited on a carbonate platform in the tropical part of the western Paleo Tethys ocean.The overall elemental geochemistry of the detailed two-metre-thick section across the boundary that we studied shows that the clastic content of the sediments came from dominantly silica-rich continental sources though with some more silica-poor inputs in the uppermost Permian and lowest Triassic limestones as shown by Ni/Al and Nb/Ta ratios.These inputs bracket, but do not coincide with, the main extinctions and associated C, O and S changes.Increased aridity at the Permian-Triassic boundary with increased wind abrasion of suitable Ti-bearing heavy minerals accounts for both the high Ti/Al and Ti/Zr ratios.Various geochemical redox proxies suggest mainly oxic depositional conditions, with episodes of anoxia, but with little systematic variation across the Permian–Triassic extinction boundary.The lack of consistent element geochemical changes across the Permian-Triassic boundary occur not only in adjacent shallower-water marine sections, and in other marine sections along the SW Tethys margin such as the Salt Range sections in Pakistan, but also in deeper shelf and oceanic sections, and in non-marine African and European continental sediments.In the absence of significant changes in physical environments, chemical changes in the atmosphere and oceans,reflected in various isotopic changes, drove the Permian–Triassic extinctions.  相似文献   

9.
白垩纪-古近纪(K-Pg)生物大灭绝事件前后的孢粉植物群的变化,可以详细揭示植被对一场全球环境危机(6550万年前小行星撞击现今的墨西哥)的响应。在K-Pg界线处有一些植物门类灭绝了,因此孢粉地层学是确定非海相序列中白垩纪-古近纪界线的主要工具。南半球包括以下植物区:热带至亚热带棕榈植物大区,高纬度Nothofagidites/Proteacidites大区,以及一个含有混合的植物组分的过渡区域。在棕榈植物大区,一些马斯特里赫特期的关键物种在白垩纪-古近纪界线处灭绝了,包括Aquilapollenites magnus、Buttinia andreevi、Crassitricolporites brasiliensis、Proteaci ditesdehaani和Gabonisporis vigourouxii。在Nothofagidites/Proteacidites大区,Tricolporite slilliei、Triporopollenites sectilis、Quadraplanus brossus、Nothofagidites kaitangata和Grapnelispora evansii等物种最后出现于白垩纪-古近纪界线处。由此可见,我们需要对南半球与中国的K-Pg界线处的孢粉信息进行更详尽的分析,才能更详细地了解不同纬度、不同地点的植被对墨西哥尤卡坦撞击事件的响应。  相似文献   

10.
The Molí del Baró-1 site (north-eastern Iberia) has yielded one of the richest fossil assemblages of the continental upper Maastrichtian from western Europe. It includes abundant plant (charophytes, sporomorphs, angiosperm leaves, seeds and logs), invertebrate (mollusc shells, partial insect exoskeletons and eggs) and vertebrate (teeth, bones, eggshells) fossils. The fossil assemblage is composed of both parautochthonous and allochthonous remains that mostly deposited in a pool formed at the edge of a meandering river. Plant megafossil evidence suggest that at least three angiosperm species (a willow-like dicot, palms and a likely typhacean monocot) formed the plant community growing in this fluvial environment. Tooth remains indicate that crocodylomorphs were diverse (including atoposaurids, ziphosuchians and eusuchians) which together with small theropod dinosaurs likely consumed prey carcasses or scavenged near the depositional setting. Prismatoolithid eggshells also suggest the presence of nearby nesting area of theropods. On the other hand, hadrosauroids apparently dominated phytophagous faunas including lambeosaurines as well as more basal forms. The taxonomic composition of the Molí del Baró-1 site matches the expected for the timespan and environment and represents a relevant locality to know the diversity of the latest Cretaceous terrestrial ecosystems in south western Europe.  相似文献   

11.
The uppermost Cretaceous (upper Campanian–Maastrichtian) marine deposits of the central south Pyrenees host a rich larger benthic foraminiferal fauna and several rudist-rich levels. These marine deposits are directly overlain by the continental facies of the Arén and Tremp Formations, which are famous for their fossil dinosaur remains. Larger benthic foraminiferal distribution documents an important faunal turnover in all the carbonate platform environments within the photic zone, from open marine to littoral areas. Biostratigraphy indicates that this turnover occurred close to the Campanian-Maastrichtian boundary. This is also confirmed by strontium isotope stratigraphy which indicates an earliest Maastrichtian age for the appearance of the larger benthic foraminiferal assemblage constituted by Lepidorbitoides socialis, Clypeorbis mammillata, Wannierina cataluniensis, Orbitoides gruenbachensis, Siderolites aff. calcitrapoides, Fascispira colomi, Omphalocyclus macroporus and Laffiteina mengaudi. In particular, a numerical age of 71 Ma is obtained for the Hippurites radiosus level, just a few meters below the first continental deposits of the Arén sensu stricto Formation. The youngest marine sediments of the central south Pyrenees are early Maastrichtian in age. This is also an important constraint for the age of the end-Cretaceous dinosaur fossil localities of the Tremp basin.  相似文献   

12.
The continental climatic evolution of Anatolia has been reconstructed quantitatively for the last 45 million years using the coexistence approach. Although there were some regional effects, the Anatolian Cenozoic continental climate record correlated with the European climatic condition and the global oxygen isotope record from marine environments. From middle Eocene to late Miocene, continental warming in Anatolia was pronounced for inferred winter temperature and mean annual temperature as in Europe. Generally, the palaeoclimatic property of Anatolia resembles the European climatic changing and marine temperature changing based on the oxygen isotope record; however, climatic values of the terrestrial area in Anatolia are higher from Lutetian to Aquitanian and these values are lower than European values from Aquitanian to Tortonian. Correspondingly, Cenozoic climatic cooling in Anatolia is directly associated with an increase of seasonality, palaeogeographic position and terrestrial condition. Furthermore, mean annual precipitation values of Anatolia remained relatively stable during the Eocene–Oligocene; however, these values indicated changing throughout middle–late Miocene. Moreover, in this study, decline of abundance and variables for the mangrove and back mangrove palaeocommunities during the last 45 million years is recorded because of the decreasing of humidity, temperature and increasing of terrestrial condition.  相似文献   

13.
In the locality of Rambla de los Gavilanes (Betic Chain Murcia, Spain), six new outcrops with dinosaur footprints are reported from an uppermost Cretaceous succession of coastal marine deposits. Four of these sites are found in the upper part of the lower Maastrichtian and the other two in the upper Maastrichtian, these latter located only a few meters below the Cretaceous-Paleocene boundary (K/Pg). Traces appear as impressions or as natural casts, on tabular limestone beds consisting of micritic carbonate facies, these deposited in coastal marine areas that include a variety of environments (coastal lakes, tidal flats, salt marshes, and shallow coastal lakes). Two main types of traces have been recognized, respectively attributed to sauropod footprints and large ornithopod footprints with remarkable pointed hoof marks. The Maastrichtian sites with large ornithopod footprints in Europe are reported from Spain, Poland and Romania. Sauropod (titanosaurian), theropod (Irenesauripus and other undetermined theropods), and ornithopod footprints have been found at sites located up to 20 m below the K/Pg boundary. The Spanish dinosaur ichnites known closest to the K/Pg boundary are in the Pyrenean upper Maastrichtian, and are tridactyl, theropod and ornithopod (Hadrosauropodus and Amblydactylus) footprints. In this work, the assignation of the Rambla de los Gavilanes ornithopod footprints has been made following the recommended synthesis in several recent papers which reduce the large ornithopod ichnogenus to four.  相似文献   

14.
The Paleocene-Early Eocene larger benthic foraminifera (LBF) in the far eastern Neo-Tethyan Ocean of Tibet still remain poorly known. Here, we present a novel, high-resolution larger foraminiferal biozonation from the shallow-water limestones in Tibet, which will improve our current understanding of the larger foraminiferal evolution in the eastern Neo-Tethyan Ocean. Based on one continuous section at Tingri and three separate sections at Gamba, ten Shallow Benthic Zones (SBZ 1–10) have been designated in Tibet by following the principle of Oppel Zone. In contrast to those in Europe, the Paleocene LBFs in Tibet are characterized by high diversification of Lockhartia, Kathina, Daviesina, Miscellanea, Ranikothalia, and Operculina and show progressively increasing diversity of genera and species during SBZ 2–5. Adult dimorphism and large shell size of some LBFs as well as differentiation of the diversity between genera and species initiated as early as SBZ 3. It suggests that the occurrence of the Larger Foraminifera Turnover (LFT) was probably not synchronous in the entire Neo-Tethyan Ocean, because in Europe, the LFT was generally thought to occur at the beginning of SBZ 5. During the Early Eocene, the LBFs in Tibet decreased markedly on the generic level and increased on the species level, and some new genera (Alveolina, Orbitolites, Nummulites, Assilina, Discocyclina) have gained predominance in Tibet. It is nearly identical to the evolution of the LBFs in Europe and indicates a high-degree homogenization of the LBFs in the entire Neo-Tethyan Ocean. Furthermore, the Paleocene-Eocene (P-E) boundary in shallow-water environments has been clearly identified by us, and it is situated in the upper part of SBZ 5 and associated with no evident biotic turnover of shallow benthic foraminiferal communities. The possible diachroneity of the LFT in the Neo-Tethyan Ocean and the evident lagging of the Paleocene-Eocene Thermal Maximum (PETM) behind the LFT imply that the LFT could only be the result of a natural evolutionary process and has no linkage with the PETM. Notably, a transient but distinct larger foraminiferal extinction and origination (LFEO) event has been found in Tibet, which is characterized by a sudden disappearance of all Paleocene lamellar-perforate LBFs, such as Lockhartia, Kathina, Daviesina, Miscellanea, Ranikothalia, and Operculina, and the initial dominance of the Early Eocene porcellaneous-walled Alveolina. The LFEO marks the boundary between SBZ 5 and 6, and might only occur in the low-latitude areas of the Neo-Tethyan Ocean. Surprisingly, the LFEO coincides with the initial recovery of the Carbon Isotope Excursion (CIE), and their synchronicity implies that some possible mechanisms causing the rapid recovery of the CIE probably had also led to the LFEO in the shallow-water environments.  相似文献   

15.
We describe remains of a new crocodyliform found from the Lower Cretaceous (Aptian) Khok Kruat Formation, northeastern Thailand. Remains consist of two caudal ends of mandibles, two rostral symphyseal parts of right rami of mandibles, a dorsal part of postorbital, a cranial end of squamosal and one osteoderm. Phylogenetic analyses supported inclusion of this crocodyliform into the Eusuchia as it shares several morphological characters with other eusuchians, such as a dorsocaudally oriented retroarticular process, smooth lateral surface of the caudoventral region of mandible, and a craniocaudally oriented ridge on the dorsal surface of retroarticular process. The shape of symphyseal region showed this crocodyliform had a longirostrine snout shape, which is uncommon in early eusuchians. Finding of this crocodyliform draws back the oldest record of Asian eusuchians, which was Tadzhikosuchus, approximately 30 million years and it is the only Mesozoic eusuchian found in East and Southeast Asia.  相似文献   

16.
The poor preservation and apparent monospecifity of Permian tetrapod footprints from eolian paleoenvironments have thus far hampered their reliable interpretation. This study clarifies how this is due to distinct and repeated ichnotaphonomic effects on trackway pattern and footprint morphology on originally inclined planes. Once these effects are excluded, the anatomy-consistent ichnotaxobases useful for ichnotaxonomy can be recognized. Several nomina dubia are identified, among these the ichnogenus Chelichnus, here considered a taphotaxon. The eolian ichnoassociations from the Lopingian of Scotland and Germany include six different ichnotaxa: cf. Capitosauroides isp. (?eutheriodont therapsid), Dicynodontipus geinitzi (cynodont therapsid), Dolomitipes isp. (dicynodont therapsid), Pachypes loxodactylus n. comb. (pareiasaurian parareptile), Procolophonichnium isp. (small parareptile) and Rhynchosauroides isp. (non-archosauriform neodiapsid). This is completely different from the interpretations of the last 20 years, which postulated that these paleoenvironments comprised monospecific associations of synapsid tracks. These ichnoassociations are instead moderately diverse, similar to low-latitude marginal marine to floodplain ichnoassociations and belong to the Lopingian Paradoxichnium footprint biochron. The Cornberg Formation of Germany, being constrained between the Illawarra reversal and the mid-Wuchiapingian Kupferschiefer at the Rotliegend/Zechstein transition, constitutes the earliest evidence of Lopingian tetrapod faunas at low paleolatitudes and the first evidence of low-paleolatitude faunal turnover related to the end-Guadalupian mass extinction from both the skeleton and the track record. This suggests a global extension of the dinocephalian extinction event, which occurred at high-mid (South Africa and Russia) and low (Western Europe) paleolatitudes of Pangea about 259–260 Ma and was probably triggered by the eruption of the Emeishan Large Igneous Province of SW China, which considerably changed global environmental conditions in both marine and continental settings.  相似文献   

17.
A review of the tetrapod (amphibian and amniote) record across the Permo-Triassic boundary (PTB) indicates a global evolutionary turnover of tetrapods close to the PTB. There is also a within-Guadalupian tetrapod extinction here called the dinocephalian extinction event, probably of global extent. The dinocephalian extinction event is a late Wordian or early Capitanian extinction based on biostratigraphic data and magnetostratigraphy (the extinction precedes the Illawara reversal), so it is not synchronous with the end-Guadalupian marine extinction. The Russian PTB section documents two tetrapod extinction events, one just before the dinocephalian extinction event and the other at the base of the Lystrosaurus assemblage. However, generic diversity across the latter extinction remains essentially the same despite a total evolutionary turnover of tetrapod genera. The Chinese and South African sections document the stratigraphic overlap of Dicynodon and Lystrosaurus. In the Karoo basin, the lowest occurrence of Lystrosaurus is in a stratigraphic interval of reversed magnetic polarity, which indicates it predates the marine-defined PTB, so, as previously suggested by some workers, the lowest occurrence of Lystrosaurus cannot be used to identify the PTB in nonmarine strata. Correlation of the marine PTB section at Meishan, southern China, to the Karoo basin based primarily on magnetostratigraphy indicates that the main marine extinction preceded the PTB tetrapod extinction event. The ecological severity of the PTB tetrapod extinction event has generally been overstated, and the major change in tetrapod assemblages that took place across the PTB was the prolonged and complex “replacement” of therapsids by archosaurs that began before the end of the Permian and was not complete until well into the Triassic. The tetrapod extinctions are not synchronous with the major marine extinctions at the end of the Guadalupian and just before the end of the Permian, so the idea of catastrophic causes of synchronous PTB extinctions on land and sea should be reconsidered.  相似文献   

18.
The Early Triassic Induan–Olenekian Stage boundary (Dienerian–Smithian sub-stage boundary) has been identified at a depth of 2719.25 m in the petroleum exploration well Senecio-1 located in the northern Perth Basin, Western Australia. Conodont faunas represent three conodont zones in ascending order, the Neospathodus dieneri Zone, the Neospathodus waageni eowaageni Zone and the Neospathodus waageni waageni Zone. The Induan–Olenekian (Dienerian–Smithian) boundary is placed at the base of the Neospathodus waageni eowaageni Zone equivalent to the first appearance of Neospathodus ex. gr. waageni utilised elsewhere and adopted by the IUGS ICS Triassic Subcommission to define the base of the Olenekian. Bulk kerogen δ13C carbon isotopes define a positive peak of c. 4 per mille that essentially coincides with the Induan–Olenekian boundary as seen in proposed Global Stratotype Sections and Points (GSSPs) in South China and Spiti, India demonstrating the global utility of this level for correlation. An anoxic zone is recognised in the lower part of the Senecio-1 core and the upper limit of this zone is dated as late Induan (late Dienerian). Temporal and spatial mapping of marine anoxia and dysoxia globally demonstrates that pulses of dysoxia/anoxia affected shallow-marine zones at different times in different locations. Dysoxia/anoxia in the shallow-marine environment appeared in the latest Permian at the extinction level, later than in the deep-marine environment, and appears to be largely restricted to the Induan (Griesbachian and Dienerian) and early Olenekian (Smithian). Temporally and geographically restricted upwelling of an oxygen minimum zone into the ocean surface layer due to environmental perturbations including extreme global warming, increased terrestrial chemical weathering intensity and continental erosion, sea level rise, and changes in marine nutrient inventories and productivity rates, is interpreted as a likely cause of observed variation in shallow-marine dysoxia/anoxia in the Early Triassic.  相似文献   

19.
We sampled 150 sites in fine-grained Plio-Pleistocene sediments of the Palm Springs and Imperial Formations. Sampling was confined to 3000 m of stratigraphically continuous section containing abundant vertebrate fossil remains of the Vallecito Creek, Arroyo Seco, and Layer Cake local faunas of the Irvingtonian and Blancan Land Mammal Ages. The magnetic stability of these sediments was sufficient to delineate the magnetic stratigraphy, which ranges from below the Cochiti event at the base, to the Matuyama reversed magnetic epoch at the top of the section. Eight faunal events are placed relative to the magnetic polarity sequence. They are cf. Pliohippus extinction, Geomys appearance, cf. Equus appearance, Tremarctos appearance, Hypolagus extinction, cf. Odocoileus appearance, Smilodon and ? Euceratherium appearance. The latter two faunal events characterize Irvingtonian Land Mammal Age. The transition from the Blancan to Irvingtonian Land Mammal Age occurs in the lower Matuyama magnetic epoch close to the Pliocene-Pleistocene boundary. The appearance of European migrants during the lower Matuyama magnetic epoch indicates that the Bering land bridge was exposed for animal migration between Europe and North America at this time.  相似文献   

20.
We propose a population model for Middle Pleistocene Europe that is based on demographic “sources” and “sinks”. The former were a small number of “core” or populations in glacial refugia in southern Europe from which hominins expanded northwards in interstadial and interglacial periods; occupation outside glacial refugia would have been restricted to warm or temperate periods, and populations at the northern limit of the Middle Pleistocene range would have been “sink” populations in that they depended upon recruitment from source populations further south. Southwest Asia would also have been a likely source of immigrant, source populations. We argue as an alternative to an “ebb and flow” model in which groups retreated to refugia when conditions worsened that local extinction outside refugia would have been frequent. In extreme situations, Europe may have been a population “sink” (i.e. unpopulated) that was replenished from source populations in Southwest Asia. We suggest that this pattern of repeated colonisation and extinction may help explain the morphological variability of European Middle Pleistocene hominins, particularly Homo heidelbergensis and its apparent non-lineal evolution towards Homo neanderthalensis.  相似文献   

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