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1.
Cretadilar olei gen. et sp. nov. and Burmopsychops groehni sp. nov. (Neuroptera: Dilaridae) are described from the Late Cretaceous (earliest Cenomanian/late Albian) Burmese amber. Their dilarid affinities are supported by two autapomorphies of the family: the reduction of the ligula and the presence of pectinate antennae in males. They are considered to belong to two new subfamilies, Cretanallachiinae subfam. nov. and Cretadilarinae subfam. nov., respectively. These subfamilies are easily distinguished from other dilarids (Dilarinae, Nallachiinae, and Berothellinae) by possessing siphonate mouthparts; however they strongly differ from each other. In particular, the broad gonocoxites 9 of Cretanallachiinae females are not fused to form a distinct ovipositor, and bear long gonostyli, whereas a distinct saber-like ovipositor (short, lacking gonostyli) is present in Cretadilarinae. It is assumed that Cretanallachiinae primarily fed on pollen and Cretadilarinae on the nectar of flowers.  相似文献   

2.
A new species, Eoptychoptera cantabrica sp. nov. is described from the Albian El Soplao amber-bearing deposits based on a single male with an elongated proboscis, unknown among extant members of Ptychopteridae. It represents the youngest record of the species-diverse genus Eoptychoptera. A pupa of Eoptychoptera sp. with a long respiratory horn, typical of extant ptychopterids, is described from Las Hoyas limestones, where adult fossil ptychopterids have yet to be discovered. A key to species of Eoptychoptera, based on wing venation, and a distribution map of Cretaceous Ptychopteridae are provided. The climatic preferences and mouthparts of Mesozoic and recent Ptychopteridae are discussed.  相似文献   

3.
In the course of evolution, mutualism between pollinators and plants was likely first developed between insects and gymnosperms, since the occurrence of long-proboscid Mecoptera, Neuroptera and Diptera predates the diversification of flowering plants in the Early Cretaceous by at least 60 million years. Here we report one of the most advanced pre-angiosperm pollinator, the Late Jurassic acrocerid fly Archocyrtus kovalevi (Nartshuk, 1996). Re-examination of the holotype specimen has shown that A. kovalevi had an extremely long siphonate proboscis (12 mm) almost twice the length of its body. Relatively, this kind of mouthpart was the longest of all long-proboscid Mesozoic insects hitherto. While long-proboscid species of extant Acroceridae are known as pollinators of flowers with long corolla tubes, we hypothesize that A. kovalevi pollinated bisexual bennettitalean cones such as Williamsoniella karataviensis Tur.-Ket., 1963 from the same deposits.  相似文献   

4.
Oxyporinae is a distinct staphylinid subfamily characterized by a large body, remarkably long and projecting mandibles, and large, crescent-shaped terminal labial palpomeres. Previously, only three compression fossils from the Lower Cretaceous Yixian Formation in China were known for the subfamily from the Mesozoic Era. This study describes a remarkable new species, Oxyporus cretaceous sp. nov., based on a single specimen in Upper Cretaceous Burmese amber. This finding is the earliest record of Oxyporinae preserved as amber inclusion. The well preserved mouthparts suggest that mycophagous feeding habits had already been acquired by the mid-Cretaceous.  相似文献   

5.
A new family of Scarabaeoidea is revealed from the Cretaceous amber of Myanmar (earliest Cenomanian, near 100 Ma), on a series of males and females, most in perfect condition. Passalopalpidae fam. nov., Passalopalpus cheni gen. et sp. nov., are probably extinct since a long time. Observations were made on the whole body (head, thorax, abdomen) and appendages (mouthparts, antennae, legs, wings). Most characters are remarkable and plesiomorphic, compared to Passalidae Leach, as well as to some other Scarabaeoidea. Morphological characteristics demonstrate that the fossil species is typically « passaloid », but that it cannot belongs to Passalidae. Passalopalpidae and Passalidae are clearly distinct families, with strong assumption to be sister groups. The systematic interpretation of the fossil could fill a historic gap: the sister group of Passalidae had been established previously never in a solid way. Passalids are Gondwanian, whereas passalopalpids lived on former eastern Laurasia. Vicariance of both taxa, each on one of the super-continents, seems to support recent hypotheses of phylogeny and paleobiogeography for Passalidae. Other preliminary notes are given on the morphology, biology and phylogenetic relationships of passalopalpids.  相似文献   

6.
A new species of Elcanidae (Orthoptera: Elcanoidea), Panorpidium yixianensis sp. nov., is described based on two new specimens from the Lower Cretaceous Yixian Formation of northeastern China. It differs from other species in forewing characters and spines on the hind tibiae. In addition, a new specimen Burmelcana sp., is described and figured based on an amber inclusion from the mid-Cretaceous Burmese (Myanmar) amber. P. yixianensis sp. nov. represents the first definite record of Elcanidae in the Early Cretaceous Jehol biota, and Panorpidium is the only genus of Ensifera to be found in the Early Cretaceous faunas of England, Russia and China.  相似文献   

7.
A new genus and species of female phantom midge Neuseptychoptera carolinensis gen. et sp. nov. in the fossil subfamily Eoptychopterinae is described and illustrated; this ptychopterid is from Upper Cretaceous (Campanian) amber of North Carolina. This is the first member of Eoptychopterinae from North America and the first found in Campanian age amber. We propose to treat the extinct subfamily Eoptychopterininae Lukashevich, 1995, as a junior synonym of the subfamily Eoptychopterinae Handlirsch, 1906, syn. nov. The elongated mouthparts of the new species have broad labella, and these are functionally interpreted as feeders for honeydew, fermenting plant juices, and nectar rather than for biting.  相似文献   

8.
A new genus and species of aphids, Tanyaulus caudisetula gen. et sp. nov. (Hemiptera: Aphidoidea: Burmitaphidae), is described from mid- Cretaceous Myanmar amber. Autapomorphies for the new genus are stub-like hind wings, 7-segmented antenna, long rostrum equal to body length, 4 veins departing from the main vein, cubital veins distinctly separated, with a very short, curved basal proximal cubitus, one-branched M vein with distal fork arched at the base, distinct claval folds at wing bases and caudal protuberance with extended setae. The new fossil adds to our knowledge of the morphological diversity of Mesozoic Aphidoidea.  相似文献   

9.
Although recent molecular phylogenetic analyses suggest a great antiquity, going into the Jurassic, for the small modern fly family Hilarimorphidae, no fossil was attributed to this group. The first fossil hilarimorphid Cretahilarimorpha lebanensis gen. et sp. nov., is described, based on a specimen from the Lower Cretaceous Lebanese amber. Its external morphology is analysed and compared with that of Hilarimorpha, unique modern hilarimorphid genus. The differences concern the wing venation, antenna and especially mouthparts. Cretahilarimorpha has very elongate mouthparts, adapted for nectar feeding or (less probably) for predation on other insects. Several other Lower Cretaceous lineages have developed similar elongate mouthparts, viz. nemestrinid and xylomyiid flies, and the Mecoptera: Aneuretopsychina, probably adapted to still unknown deep nectar-producing flowers. A checklist of species belonging to the Hilamorphidae is given.  相似文献   

10.
Prosisyrina sphinga sp. nov. (Neuroptera: Sisyridae) is described from Upper Cretaceous (Santonian) Taimyr amber of northern Siberia (Yantardakh locality). The new species is preliminary assigned to this Cretaceous genus, mainly based on character states of the maxillary palpus and the hind wing vein M and crossvein 1r-m, and the presumable absence of tibial false spurs. However, the structure of the costal space and the RP branching in the hind wing indicate that it may belong to a new, closely related genus. The discovery of this second species from the locality might suggest that sisyrids were usual members of the Santonian riparian biocenoses of northern Siberia.  相似文献   

11.
The family Mesopsychidae Tillyard,1917 presently consists of ten described genera from the Early Permian to the Early Cretaceous of Australia,China,Kyrgyzstan,Russia,South Africa,Tajikistan,and Ukraine.Herein,a new genus and a new species of fossil mesopsychid,Epicharmesopsyche pentavenulosa gen.et sp.nov.,is described from the Middle Jurassic Jiulongshan Formation of Inner Mongolia,China,supplementing the family-level diversity.The well-preserved,new material reveals many characters,including antennae,wing venation,shape and genitalia that increase our morphological understanding of the family,although unfortunately the mouthparts and most of the legs are not preserved.Diagnosis of the Mesopsychidae is emended to include a four-or five-branched MP in both the fore-and hind wings.This is the first documentation of a wing coupling structure in the Mesopsychidae,consisting of three to four frenula bristles on the humeral lobe at the base of the costal margin of both hind wings.Asymmetrical shape and size of the left and right wings on a specimen of E.pentavenulosa gen.et sp.nov.,seems to be a common condition for mesopsychid taxa from northeastern China.  相似文献   

12.
The Cretaceous lacewing family Babinskaiidae comprises nine species in seven genera, of which two new genera and one new species are described herein: Parababinskaia elegans gen. et sp. nov. from the late Aptian Crato Formation of Brazil, and Pseudobabinskaia gen. nov. (with the only species Pseudobabinskaia martinsnetoi Lu et al., 2017, comb. nov.) from mid-Cretaceous Burmese amber. The reduction of the veins AA2 and AA3 in the hind wing is an autapomorphy of the family. Babinskaiidae are most closely related to Nymphidae, and have probably evolved as a side branch of these. The superfamily Myrmeleontoidea is divided into two epifamilies: Nymphidoidae whose MP (or M) is long, not fused with CuA, and trichosors are present (Nymphidae, Babinskaiidae), and Myrmeleontoidae whose MP is crossvein-like, then fused with CuA, and trichosors are lost (Araripeneuridae, Ascalaphidae, Myrmeleontidae, Nemopteridae, Palaeoleontidae). The aligned crossveins 1r-m and 1m-cu in the forewing is probably the only venational autapomorphy of the Myrmeleontoidea.  相似文献   

13.
Two new genera and three new species of the Mesozoic family Aeotheogrammatidae are described from the Lower Cretaceous of Yixian Formation (Liaoning Province, China): Cyclicogramma rotundum gen. et sp. nov. and Aetheogramma bistriatum sp. nov. from Huangbanjigou, and Curtogramma ovatum gen. et sp. nov. from Dawangzhangzi. The hind wing of Cyclicogramma rotundum gen. et sp. nov. differs from that of other species by its smaller size, its overall shape, RP bearing only two oblique radial branches, and its configurations of CuP, AA1, and AA2. Aetheogramma bistriatum sp. nov. is closely related to A. speciosum Ren and Engel, 2008, but distinguished from the latter by its distinctive configuration of RP1, and the narrower and longer AA2 space in the hind wing. Curtogramma ovatum gen. et sp. nov. is distinguished from all other species of the family by the forewing with M forking very far from the wing base.  相似文献   

14.
This paper describes a new ichthyosaur specimen from the Stonebarrow Marls Member (Charmouth Mudstone Formation) of Charmouth, Dorset. It is substantially complete and various characters of the skull (snout ratios and basioccipital morphology) and appendicular skeleton (the number of digits on the hind paddle, which can be used to infer the number of digits on the front paddle) indicate that the specimen is referable to Ichthyosaurus communis. This provides the first direct evidence for the presence of I. communis in the Pliensbachian, extending the stratigraphic range of the species. The specimen also preserves a probable gullet content, which demonstrates that fish formed part of the diet of this individual.  相似文献   

15.
The fossil snakefly family Baissopteridae from the Lower Cretaceous (upper Barremian) of the Yixian Formation of Liaoning Province, China is reviewed. Two genera and five species of Baissopteridae are recorded, including a new genus, namely Microbaissoptera gen. nov., and three new species, i.e., Baissoptera bicolor sp. nov., Baissoptera sinica sp. nov., and Microbaissoptera monosticha sp. nov. Moreover, Baissoptera euneura Ren, 1997 is herein treated as a new junior synonymy of Baissoptera grandis Ren in Ren et al., 1995. Baissoptera minima Ponomarenko, 1993 is transferred to the presently described new genus Microbaissoptera gen. nov. based on the small body-size and the presence of a single gradate series of crossveins in radial and medial areas of posterior part of both fore- and hind wings. A key to genera of Baissopteridae is provided.  相似文献   

16.
A strange wingless female parasitic wasp from mid-Cretaceous Burmese amber is described as Aptenoperissus burmanicus sp. et gen. nov. in the new family Aptenoperissidae (Hymenoptera, Ceraphronoidea). Diagnostic characters of the female Aptenoperissus burmanicus sp. et gen. nov. include its wingless, streamlined and heavily sclerotized body lacking any apparent trace of a wasp waist, and geniculate antenna composed of a long, thin, stick-like scape, standard pedicel and 22 uniform flagellomeres. Also the body has 9 externally visible segments with no evidence of segment fusion implying the presence of a completely hidden segment. All tibiae have paired spurs and the hind femora are saltatory and incrassate. The double fore-tibial spur combined with unquestionable diagnostic features of Apocrita (primarily an internalized needle-like thin and acute ovipositor) suggest placement within the superfamily Ceraphronoidea s.str. with the Maimetshidae as a sister group of the crown Ceraphronoidea, composed of the Ceraphronidae, Megaspilidae, Stigmaphronidae, and Radiophronidae. The fossil is hypothesized to live semicryptically on the forest floor or tree trunk and to parasitize immature holometabolous insects. Diagnostic features of a respective male are suggested to test the hypothesized position of the new taxon.  相似文献   

17.
The first definitive fossil of the peculiar monotypic rove beetle subfamily Megalopsidiinae is described as a new species Megalopinus extinctus sp. n. It represents a stem lineage valuable for the study of the early diversification of Staphylinidae, where sister group relationships of Megalopsidiinae currently remain a big controversy. This discovery corroborates the Mesozoic origin of this subfamily implied by its presumably basal phylogenetic position within Staphylinidae and hitherto available fossil record for the family. Well preserved peculiar mouthparts of M. extinctus specialized similarly with recent Megalopinus suggest the same mode of feeding in Megalopsidiinae for nearly a hundred million years.  相似文献   

18.
19.
P. Nel  D. Azar  A. Nel   《Cretaceous Research》2007,28(6):1033-1038
A new family of thrips, Moundthripidae, is described on the basis of a new genus and species, Moundthrips beatificus, from Early Cretaceous Lebanese amber. This taxon has plesiomorphic prognathous mouthparts, a unique type of wing venation and the main apomorphies of the Thysanoptera of the legs and mouthpart structures, suggesting that they were acquired very early during the evolution of this order.  相似文献   

20.
New Mesozoic Mesopsychidae(Mecoptera)from Northeastern China   总被引:2,自引:2,他引:0  
<正>The Mesozoic family Mesopsychidae Tillyard,1917 presently consists of seven described genera and 13 species from the mid-Triassic to the Early Cretaceous of Australia,South Africa and Eurasia.In the present paper one new genus and three new species of fossil mesopsychids are described that add significant distributional and stratigraphic extensions to the family.This finding documents the first formal record of fossil Mesopsychidae in China.Both Lichnomesopsyche gloriae gen.et sp.nov.and L.daohugouensis gen.et sp.nov. were found from the Middle Jurassic Jiulongshan Formation of Inner Mongolia,whereas Vitimopsyche kozlovi sp. nov.,of mid Early Cretaceous age,was collected from the Jehol Biota of northern Hebei Province.This new,well-preserved material from China reveals complete and previously unknown body features,including head, antennae,mouthparts,legs and abdomen.The delicate and long proboscides of these new taxa indicate that they were feeding on externally exposed,nutrient-rich fluids of gymnospermous ovulate fructifications,and incapable of piercing surface epidermis-attributable principally to the absence of stylets.These proboscides originated, perhaps multiply,among basal Mecoptera and are functionally and structurally convergent with equivalent mouthparts borne by fossil and extant Diptera,Lepidoptera,Neuroptera and Coleoptera.  相似文献   

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