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1.
The Kholokhovchan Flora comes from tuffaceous – terrigenous deposits of the Vetvinskaya Member (Chalbugchan Group) in the Penzhina and Oklan rivers interfluve, Northeastern Russia. The depositional environment of the plant-bearing deposits is interpreted to have been a freshwater lake. The Kholokhovchan Flora hosts 42 fossil plant species belonging to Marchantiopsida, Polypodiopsida, Ginkgoales, Leptostrobales, Bennettitales, Pinales and Magnoliopsida. It is characterised by diverse angiosperms, less diverse conifers and ferns, by the presence of relatively ancient Sphenobaiera, Phoenicopsis and Pterophyllum together with advanced Late Cretaceous Taxodium, Glyptostrobus and angiosperms, among which platanoids are quite diverse. The Kholokhovchan Flora is most similar to Penzhina and Kaivayam floras of the Anadyr-Koryak Subregion and Arman Flora of the Okhotsk-Chukotka volcanogenic belt (Northeastern Russia) and should be dated as Turonian–Coniacian. The Kholokhovchan Flora, that populated volcanic plateaus and intermontane valleys, are characterised by a mixture of ancient “Mesophytic” plants with typical Late Cretaceous “Cenophytic” taxa. This peculiar composition probably reflects a gradual penetration of new angiosperm-dominated plant assemblages into older floras: during the Late Cretaceous, “Cenophytic” assemblages migrated along river valleys and other disturbed habitats into the interior of Asia, eventually occupying volcanogenic uplands, and in places replacing the “Mesophytic” fern-gymnospermous communities that existed there. Two new angiosperm species, as well as four the most characteristic conifers of the Kholokhovchan Flora, are described: Cupressaceae gen. et sp. indet. cf. Widdringtonites sp., Taxodium cf. olrikii, Taxodium sp., Glyptostrobus sp., Ettingshausenia vetviensis sp. nov. and Parvileguminophyllum penzhinense sp. nov.  相似文献   

2.
Metriorhynchidae is a clade of marine-adapted crocodilians known from several Middle Jurassic–Early Cretaceous specimens collected predominantly in South America and Europe, but poorly known in the northern margin of Gondwana. The “Portomaggiore crocodile” is the most complete specimen of an Italian metriorhynchid to date: it consists of a partial skeleton that has been provisionally referred to an unnamed species of Late Jurassic Metriorhynchus or Geosaurus. The specimen is preserved in the reddish, nodular limestone of the Rosso Ammonitico Veronese Formation (Bajocian–Tithonian); new data on microfossil associations constrain the age of the metriorhynchid to the late Bajocian–earliest Bathonian. On the basis of cranial synapomorphies, the “Portomaggiore crocodile” falls as the closest sister-taxon of the Late Jurassic–Early Cretaceous geosaurines, and is referred to Neptunidraco ammoniticus gen. et sp. nov. It is unique among Middle Jurassic metriorhynchids in showing an incipient streamlining of the skull, shared with Late Jurassic and Cretaceous taxa. Since Neptunidraco is the oldest known member of Metriorhynchidae, its phylogenetic position supports the hypothesis that the timing of the initial metriorhynchid and geosaurine diversifications should start in the Bajocian.  相似文献   

3.
The new non-marine bivalve species Nippononaia (Martinsonella) tamurai sp. nov. is described from the Upper Formation of the Mifune Group in Kumamoto Prefecture, Southwest Japan. The specimens originally were identified as Plicatounio (Plicatounio) B sp. by Tamura (1990). The subgenus Nippononaia (Martinsonella) previously was only reported from China, and this is the first record from Japan. The Upper Formation of the Mifune Group is of Late Cenomanian to Early Turonian age, as indicated by ammonites. “Nippononaia” (?) obsoleta Hase, 1960 from the Shiohama Formation of Yamaguchi, Japan, and Plicatounio (s.l.) A sp. of Tamura (1990) from strata northeast of Geoncheonri, South Korea, are re-assigned to Nippononaia (Martinsonella). These occurrences may be of significance for the inter-regional correlation of non-marine Cretaceous strata.  相似文献   

4.
There is little fossil evidence for cave arthropods. Small size, disfunctional wings, extremely long antenna, dense minute setation, palidity and reduced eyes are typical adaptations to life in caves shown by the cockroach Mulleriblattina bowangi gen. et sp. n., found together with epigeic Crenocticola svadba sp. n., both from Myanmar amber and belonging to the cavernicolous cockroach family Nocticolidae. These lineages of earliest still living cavernicoles suggest large, numerous caverns, lava tubes or caves within the source area. They provide the first unequivocal evidence for the Mesozoic origin of any living troglomorphic organism, and explain the “long branches” in DNA analyses. Phylogenetic trees show little hierarchical structure and place Latindiinae and Myrmecoblatta within the explosively radiating Nocticolidae. Biogeography indicates a common cosmopolitan Early Cretaceous ancestor except for 8 (of 49) species of true Nocticolidae, which diverged during the Late Cretaceous breakup of Gondwana. A review of all troglofauna documents no other unequivocal pre-Cenozoic cave biotas (including vertebrates). Stable environments and small populations result in a short-time (˂3 Ma) origination of bizarre forms and long term extinctions (˃30 Ma).  相似文献   

5.
In this paper we describe previously unpublished trionychid turtle material, consisting of numerous shell fragments, from the Late Cretaceous (late Turonian) Bissekty Formation of the Dzharakuduk locality in Uzbekistan. This material is assigned to two shell-based taxa: Aspideretoides cf. riabinini and “Trionyx” cf. kansaiensis. The material which cannot be confidently attributed to these two taxa is identified as Trionychidae indet. In addition to these shell-based trionychid taxa, the Dzharakuduk turtle assemblage includes two skull-based taxa of trionychids (Khunnuchelys kizylkumensis and Trionychini indet.). The trionychids from the Bissekty Formation are most similar to trionychids from the younger (Santonian – early Campanian) Bostobe Formation of Kazakhstan, represented by three shell-based taxa (Aspideretoides riabinini, Paleotrionyx riabinini and “Trionyxkansaiensis), and one skull-based taxon (Khunnuchelys sp.). We provide an improved understanding of the subtle similarities and differences between four closely related Cretaceous turtle assemblages of Middle Asia and Kazakhstan.  相似文献   

6.
The behavior of the leatherjack, Oligoplites saurus, was studied in the field. Four behavioral patterns were identified: (1) cleaning behavior on the redfin needlefish, Strongylura notata, (2) intraspecific schooling, (3) interspecific schooling with the rough and tidewater silversides, Membras martinica and Menidia peninsulae, pinfish, Lagodon rhomboides, and menhaden, Brevoortia sp., (4) swimming along in a “leafmimic” position or next to floating algae or debris. Fishes exhibiting any of these behaviors fed primarily on plankton. Schooling leatherjackets also fed on scales of Strongylurus notata, Brevoortia sp., Menidia peninsulae and Lagodon rhomboides. These observations are supported by stomach content analyses. It is suggested that scale-feeding may be a preadaptation for cleaning behavior among leatherjackets.  相似文献   

7.
The Upper Cretaceous shallow-water carbonates of the Pyrenean Basin (NE Spain) host rich and diverse larger foraminiferal associations which witness the recovery of this group of protozoans after the dramatic extinction of the Cenomanian–Turonian boundary interval. In this paper a new, large discoidal porcelaneous foraminifer, Broeckina gassoensis sp. nov., is described from the middle Coniacian shallow-water deposits of the Collada Gassó Formation, in the Bóixols Thrust Sheet. This is the first complex porcelaneous larger foraminifer of the Late Cretaceous global community maturation cycle recorded in the Pyrenean bioprovince. It differs from the late Santonian–early Campanian B. dufrenoyi for its smaller size in A and B generations and the less developed endoskeleton, which shows short septula. Broeckina gassoensis sp. nov. has been widely employed as a stratigraphic marker in the regional geological literature, under the name of “Broeckina”, but its age was so far controversial. Its middle Coniacian age (lowermost part of the Peroniceras tridorsatum ammonite zone), established in this paper by strontium isotope stratigraphy, indicates that it took about 5 My after the Cenomanian–Turonian boundary crisis to re-evolve the complex test architecture of larger foraminifera, which is functional to their relation with photosymbiotic algae and K-strategy.  相似文献   

8.
Late Barremian ammonite fauna from the epipelagic marlstone and marly limestone interbeds of Boljetin Hill (Boljetinsko Brdo) of Danubic Unit (eastern Serbia) is described. The ammonite fauna includes representatives of three suborders (Phylloceratina, Lytoceratina and Ancyloceratina), specifically Hypophylloceras danubiense n. sp., Lepeniceras lepense Rabrenović, Holcophylloceras avrami n. sp., Phyllopachyceras baborense (Coquand), Phyllopachyceras petkovici n. sp., Phyllopachyceras eichwaldi eichwaldi (Karakash), Phyllopachyceras ectocostatum Drushchits, Protetragonites crebrisulcatus (Uhlig), Macroscaphites perforatus Avram, Acantholytoceras cf. subcirculare (Avram), Dissimilites cf. trinodosus (d'Orbigny) and Argvethites? sp. The taxonomic composition and percent abundance of the identified ammonites indicate that their taxa are predominantly confined to the Tethyan realm. Ammonites with smooth and slightly sculptured shells predominate among the studied fauna. The ammonite-bearing succession from Boljetin represents the lower part of the Upper Barremian, ranging in ammonite zonation from the Toxancyloceras vandenheckei Zone to the lower part of the Imerites giraudi Zone. The associated organic-walled dinoflagellate cysts confirm the Late Barremian age of the ammonite-bearing levels.  相似文献   

9.
A new Late Jurassic assemblage of “conchostracans”, ostracods, bivalves and caddisfly cases from the locality “Estancia La Sin Rumbo”, Chubut Province (Patagonia, Argentina) is recorded. The fossils occur in the upper part of an outcropping 45 m thick volcaniclastic lacustrine sequence of yellowish tuffs and tuffites of the Puesto Almada Member, which is the upper member of the Cañadón Asfalto Formation with U/Pb age of 161 ± 3 Ma. The sequence represents one sedimentary cycle composed of a (lower) hemicycle of expansion and a (higher) hemicycle of contraction of the water body. The invertebrates lived in small freshwater bodies during the periods of expansion of the lake. The occurrence of a great number of small spinicaudatans, associated with mud-cracks, is evidence of dry climatic conditions and suggests several local mortality events. The spinicaudatan record of the fushunograptid–orthestheriid (component of the Eosestheriopsis dianzhongensis fauna) and the presence of Congestheriella rauhuti Gallego and Shen, suggest a Late Jurassic (Oxfordian to Tithonian) age. Caddisfly cases are recorded for the first time in the Cañadón Asfalto Basin.  相似文献   

10.
A new middle Miocene vertebrate fauna from Peruvian Amazonia is described. It yields the marsupials Sipalocyon sp. (Hathliacynidae) and Marmosa (Micoureus) cf. laventica (Didelphidae), as well as an unidentified glyptodontine xenarthran and the rodents Guiomys sp. (Caviidae), “Scleromys” sp., cf. quadrangulatus-schurmanni-colombianus (Dinomyidae), an unidentified acaremyid, and cf. Microsteiromys sp. (Erethizontidae). Apatite Fission Track provides a detrital age (17.1 ± 2.4 Ma) for the locality, slightly older than its inferred biochronological age (Colloncuran-early Laventan South American Land Mammal Ages: ∼15.6–13.0 Ma). Put together, both the mammalian assemblage and lithology of the fossil-bearing level point to a mixture of tropical rainforest environment and more open habitats under a monsoonal-like tropical climate. The fully fluvial origin of the concerned sedimentary sequence suggests that the Amazonian Madre de Dios Subandean Zone was not part of the Pebas mega-wetland System by middle Miocene times. This new assemblage seems to reveal a previously undocumented “spatiotemporal transition” between the late early Miocene assemblages from high latitudes (Patagonia and Southern Chile) and the late middle Miocene faunas of low latitudes (Colombia, Perú, Venezuela, and ?Brazil).  相似文献   

11.
In this paper we describe previously unpublished trionychid turtle material, consisting of skull fragments, from the Late Cretaceous (late Turonian) Bissekty Formation of the Dzharakuduk locality in Uzbekistan. This material is assigned to two taxa: the skull-based Khunnuchelys kizylkumensis Brinkman et al. (1993, Can. J. Earth Sci. 30, 2214-2223) and Trionychini indet. Two specimens which cannot be confidently attributed to these two taxa are considered Trionychidae indet. In addition to these trionychid taxa known from skulls, the Dzharakuduk turtle assemblage includes at least two shell-based taxa, Aspideretoides cf. A. riabinini and “Trionyx” cf. “T.” kansaiensis. For this and other Late Cretaceous localities of Middle Asia and Kazakhstan, we suggest the probable skull-shell associations of Khunnuchelys spp. with “Trionyxkansaiensis-like forms and Trionychini indet. with Aspideretoides-like forms.  相似文献   

12.
13.
The well known Santonian conglomerates from the area north of the Harz Mountains contain holes due to leached dolomitic pebbles of Kimmeridgian age, in which well preserved casts of “worm”-burrows have been observed. In smaller pebbles - caused by the lack of space — they were burrowed in narrow windings resembling the Serpulid genusGlomerula. However, their mode of winding differs from that of these Serpulids: The convex arches made by the boring organism must be regarded as a behavioural response when approaching the surface of the pebbles. In the loops of the burrows often a “Spreite” is found, but it is also observed in the convex windings and even parallel to the straight cylindrical burrows. That is in contradiction to the conventional definition of the “Spreite”. The burrows which may belong to the Polychaetes are described asLapispecus cuniculus n. g. n. sp. Other rare burrows have been identified asDodecaceria (?) sp.  相似文献   

14.
The recently described clade Allodaposuchidae includes European eusuchian crocodyliforms restricted to the Late Cretaceous (Campanian and Maastrichtian). A new allodaposuchid crocodyliform is here described based on two specimens from the upper Campanian–lower Maastrichtian fossil site of Lo Hueco (Cuenca, Spain). This new taxon, Agaresuchus fontisensis gen. et sp. nov., is described by two complete skulls and a lower jaw associated with one of them. This new species can be distinguished unambiguously from Lohuecosuchus megadontos, the other allodaposuchid known from the same fossil site. The presence of two allodaposuchid crocodyliforms in Lo Hueco allows the recognition of the synchronic and sympatric existence of two representatives of this clade for the first time. The new genus Agaresuchus, comprises a previously described Iberian allodaposuchid species, “Allodaposuchussubjuniperus, as Agaresuchus subjuniperus, new combination.  相似文献   

15.
Here we define a new morphospecies of planktonic foraminifera Globigerinoides eoconglobatus n. sp., identified from International Ocean Discovery Program (IODP) expedition 359 samples from drift deposits of the Maldives, Inner Sea. Through biostratigraphic analysis, we infer it may be the direct ancestor of Globigerinoides conglobatus evolving from Globigerinoides obliquus in the Late Miocene (Subzone M13a). Globigerinoides eoconglobatus n. sp. can be distinguished from G. conglobatus through both its morphological traits and stable isotopic signature (δ18O and δ13C) in pre-adult and adult specimens. The most defining characteristic being its aperture height (AH). A variance in adult stable isotopic signals shows a possible difference in life strategies, possibly related to symbionts (presence/absence or concentration) and/or depth habitat. This work also tentatively shows G. eoconglobatus n. sp. and G. conglobatus abundances are linked to glacial–interglacial stages. Its low abundances and similarities to its descendent Globigerinoides conglobatus has likely accounted for it being unreported, until present, in both modern and fossil studies.  相似文献   

16.
A new soft-shelled turtle (“Trionyxjixiensis sp. nov.) from the Lower Cretaceous Chengzihe Formation, Jixi city, Heilongjiang Province, China is described on the basis of a nearly complete carapace. The new species is diagnosed by the absence of suprascapular fontanelles and absence of a preneural; eight neurals, tetragonal fifth neural; and eight pairs of costals, with the large eighth costals meeting after the eighth neural. Due to the incompleteness of the specimen and confused classification of the genera of trionychids, the new species cannot be included in any genus of Trionychinae and is temporarily assigned to “Trionyx” (sensu lato). “Trionyxjixiensis is one of the earliest trionychids, its discovery indicates that the family was already diversified during the Early Cretaceous in Asia.  相似文献   

17.
From the Kruja Zone of Albania, shallow-water carbonates assigned to the lower Campanian are described and grouped into six microfacies types of shallow subtidal to intertidal depositional settings. The limestones display internal layering suggestive of microbial fabrics with abundance of nubeculariid foraminifera, incertae sedis Thaumatoporella Pia, and subordinate calcimicrobes of possible cyanobacterial origin: Gahkumella Zaninetti, Girvanella sp., and Decastronema kotori (Radoičić). The nubeculariid morphotypes with ornamented tests and microbial coatings reveal some kind of mutualistic relationship comparable to Late Palaeozoic–Mesozoic Tubiphytes Maslov and Crescentiella Senowbari-Daryan et al. The present study expands our knowledge on the micropalaeontological characteristics of the Late Cretaceous “DecastronemaThaumatoporella association” widespread in carbonate platforms of the peri-Mediterranean region. Our findings indicate that nubeculariids may have played an important binding role in Late Cretaceous peritidal laminated limestones.  相似文献   

18.
Perissocytheridea Stephenson is characteristic of brackish water facies. In 57 samples from the Upper Oligocene to Lower Miocene, from five localities, Pirabas Formation, Pará State, Brazil eleven species have been identified. Among these species, four are new reports: Perissocytheridea punctoreticulata n. sp., Perissocytheridea largulateralis n. sp., Perissocytheridea colini n. sp. and Perissocytheridea pirabensis n. sp.; five species in open nomenclature: Perissocytheridea sp. 1, P. sp. 2, P. sp. 3, P. sp. 4, and P. sp. 5 and two species left in “aff.” abbreviation: Perissocytheridea aff. Perissocytheridea pumila and Perissocytheridea aff. Perissocytheridea brachyforma subsp. excavata. The distributional pattern of the Perissocytheridea combined with the occurrence of foraminifera Elphidium and Ammonia in the studied sections supports the presence of the brackish water facies to the respective layers. Their quantitative variation through the studied sections indicate more than one phase of salinity reduction (about >5 and <30 ppm) and/or the decrease of the sea level, forming lagoon along the Pirabas Formation. This genus has a wide paleobiogeographical occurrence and stratigraphic distribution ranging from the Cretaceous to Recent, and already been recorded in the northern of South America, especially in the Neogene of Solimões Basin, but this is the first report of a neotropical genus to the Oligo-Miocene deposits of Pirabas Formation, northern Brazilian Coast, Pará State.  相似文献   

19.
A recently discovered fish assemblage from the “Schistes à Meletta” facies (Lower Oligocene) of the Glières Plateau, Bornes Massif, Haute–Savoie, eastern France is described. The assemblage, comprising specimens ranging from fully-articulated skeletons to isolated scattered ossifications, is composed of Anenchelum cf. glarisianum, Pristigenys sp., Fistularia sp., Caranx cf. glarisianus alongside indeterminate teleosts. This new assemblage greatly increases the fish diversity previously known from the “Schistes à Meletta” of this area. The fauna shows biogeographic affinities with assemblages from the Peritethys domain, in particular from the Helvetic molassic basin and from the Paratethys. The genera identified at the Glières locality are represented today by species living in tropical nearshore environments together with species from the open sea. A possible explanation is that the environment of deposition was rather deep, and that shallow-water fishes were brought in by turbidity currents.  相似文献   

20.
The laminated marine mudstones of the Late Jurassic of Kimmeridge, southern England, yield two exceptionally well-preserved partial skeletons of a previously unrecognised species of early batoid. These are described as a new genus and species, Kimmerobatis etchesi gen. et sp. nov. which has a general “guitarfish” bauplan as in all other batoids known from the Jurassic. This species possesses a combination of primitive characters such as centra present within the majority of the synarcual and antorbital cartilages that fail to reach the pectoral skeleton along with more derived characters, such as the lack of fin spines. Until now, little study has been carried out on the affinities of Jurassic batoids, despite their key role in understanding batoid evolution. Results from parsimony and likelihood phylogenetic reconstruction indicates that the whole-bodied Jurassic batoids Spathobatis, Belemnobatis, and Kimmerobatis gen. nov. form their own clade, Spathobatidae, and do not lend support to a monophyletic “Rhinobatidae”. Among Jurassic batoids, Kimmerobatis gen. nov. is most derived, but with derived characters being independently acquired compared to modern batoids (e.g. presence of a postpelvic process). The inclusion of whole bodied Jurassic fossils have generated a more resolved hypothesis of batoid evolution throughout the Cretaceous and into the Cenozoic.  相似文献   

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