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1.
Choristoderes are a group of extinct diapsid reptiles that once occupied the freshwater systems in the Northern Hemisphere from the Middle Jurassic through the Miocene. The Early Cretaceous monjurosuchid Philydrosaurus from western Liaoning, China, represents a transitional morphotype between a broad-snouted (crocodile-like) and a narrow-snouted (gavial-like) skull during the evolution of Choristodera. New specimens of the taxon from the Early Cretaceous Jiufotang Formation include an incomplete braincase and a nearly complete juvenile skeleton; the study of these specimens provides new information on the braincase and ontogenetic change of the skeleton of this transitional monjurosuchid. In the braincase floor, the foramen internus canalis caroticus externus opens in a groove lateral to the constricted lateral aspect of the parasphenoid, and ventral exposure of this foramen differs from Champsosaurus, in which parasphenoid/pterygoid fusion completely encloses the canal. On the occiput, the vagus nerve foramen penetrates the basioccipital, while the two foramina for the hypoglossal nerve open at the exoccipital/basioccipital suture. Comparative study of the new juvenile specimen with adult Philydrosaurus reveals previously unknown developmental changes of the cranial and postcranial skeleton of this monjurosuchid. Ontogenetically, Philydrosaurus underwent a dramatic change of skull proportions, including elongation of the antorbital and postorbital regions, and elongation of the jaws, with a great increase of the number of marginal teeth. The lower temporal fenestra is entirely closed in early ontogeny, as in large, fully-grown adults. 相似文献
2.
Michael E. Burns Philip J. CurrieRobin L. Sissons Victoria M. Arbour 《Cretaceous Research》2011,32(2):174-186
Four juvenile specimens referable to Pinacosaurus grangeri (Ankylosauria: Dinosauria) are described from the Campanian (Upper Cretaceous) locality Bayan Mandahu in northern Inner Mongolia Autonomous Region (People’s Republic of China). All the specimens preserve the skulls as well as, in some cases, mandibles, postcrania, and osteoderm. They are not taphonomically deformed by expanding matrix distortion, unlike many Gobi specimens, including the holotype of P. grangeri. Bayan Mandahu is also the type locality for Pinacosaurus mephistocephalus. The proximity in space and time of these two closely related species warrants a generic and specific revision for Pinacosaurus. The distinction of the two species is based on characters of the squamosal dermal elaborations, cranial roof posterior to the orbits, premaxillary notch, and distal margin of the ilium. Although a relatively well-represented ankylosaur taxon, the phylogenetic position of Pinacosaurus has not been unequivocally resolved. A new analysis recovers Pinacosaurus as the most basal member of the Ankylosaurinae. 相似文献
3.
The anatomy of an articulated juvenile specimen of the saurolophine hadrosaurid dinosaur Edmontosaurus annectens, LACM 23504, is described in detail. This individual consists of a partial skull and nearly complete articulated postcranium, collected from upper Maastrichtian strata of the Hell Creek Formation of Montana, USA. This study aims to help understand the ontogenetic morphological changes occurring in the cranium and postcranium of saurolophine hadrosaurids using E. annectens as a case study. Emphasis is placed on those morphological attributes that are ontogenetically variable via comparative osteology between the juvenile and the available adult specimens. It is observed that much of the cranial ontogenetic variation relates to the elongation of the skull and mandible. In the postcranium, most of the ontogenetic variation concentrates in the pectoral and pelvic girdles and the stylopodia. Not all the identified patterns of ontogenetic variation may be generalized to all hadrosaurids. The impact of ontogenetic variation on phylogenetically informative characters of saurolophine hadrosaurids is evaluated. It is concluded that, at least for Edmontosaurus annectens (and perhaps other saurolophine hadrosaurids), most characters used in phylogenetic inference of these animals are not affected by ontogeny. Thus, juvenile specimens are still a source of substantial character data suitable for use in phylogenetic analyses of saurolophine relationships. Nevertheless, it is recommended that ontogenetically variable characters are left as missing data in a character–taxon matrix when only juvenile material is available for a given saurolophine taxon. Scoring those characters based solely on juveniles is likely to decrease the accuracy of the phylogenetic inference. 相似文献
4.
A Corythosaurus skull (UALVP 13) was collected in 1920 from what is now Dinosaur Provincial Park in southern Alberta, Canada and was designated the holotype of Corythosaurus excavatus, a new hadrosaur species by Gilmore (1923). In 1992, a previously uncovered, weathered, headless skeleton was found. This was collected in 2012 by the University of Alberta as there was potential it could belong to the holotype skull. In addition, an isolated hadrosaur dentary had been found in 1992 close to the articulated postcranial skeleton and may be one of the missing jaws of Corythosaurus excavatus. The hypothesis that it may be the skeleton of the holotype of Corythosaurus excavatus is tested using anatomical information and statistical analyses. Statistical comparisons suggest it is possible that the skull and dentary belong to the same specimen. Furthermore, bivariate plots and percentage prediction errors also indicate that the postcranial material could belong to the UALVP 13 skull. Because many large vertebrate fossil specimens (even types) are not always collected in their entirety, this method may be used as a line of evidence to determine whether independently collected specimens potentially belong to the same individual. The problems described herein highlight the need to collect specimens in their entirety, and for good field documentation, including the spatial and stratigraphic context of all finds. 相似文献
5.
Verónica Díez Díaz Géraldine Garcia Fabien Knoll Xabier Pereda Suberbiola Xavier Valentin 《Proceedings of the Geologists' Association. Geologists' Association》2012,123(4):626-637
The Fox-Amphoux Syncline of the Aix Basin (SE France) has yielded a diverse Late Cretaceous vertebrate assemblage, including several taxa of dinosaurs. Here, we report on cranial material of titanosaurian sauropods, which consist of a partial braincase, two fragmentary skull roofs, and nine teeth, from the Métisson locality (Var Department). The braincase differs from those found previously in Europe (i.e., Ampelosaurus atacis, Lirainosaurus astibiae, and an unnamed juvenile skull from Romania) as well as from other titanosaurian braincases in having a groove that extends along the ventral surface of the occipital condyle neck (this feature may be autapomorphic). One of the fragmentary skull roofs may belong to the same taxon, whereas the other suggests the presence of a second titanosaur at Métisson very close to Ampelosaurus. Two dental morphologies are present in the sample; one of them includes teeth of different sizes. We suggest that this could be accounted for by age differences within a single taxon, or be due to different positions in the tooth row. The presence of a new, still unnamed titanosaurian taxon in the Ibero-Armorican Island supports previous works indicating a high titanosaurian diversity during the Campanian-Maastrichtian in southern Europe. 相似文献
6.
Yabeinosaurus was the first lizard genus described from the Early Cretaceous Jehol Biota of China. The holotype of the type species, Y. tenuis, is an immature skeleton but it has been lost for decades. A second species, Y. youngi, was erected based on another immature skeleton and is distinguished by its longer, more gracile limbs. In 2001, a juvenile skeleton from the Jingangshan locality of Liaoning was designated as the neotype for Y. tenuis. Subsequently, several further specimens from other localities, juvenile and adult, have been attributed to this species. Here we describe a second lizard specimen from the neotype locality. In anterior tooth shape, finger-like mandibular angular process, cranial sculpture, maxilla shape, and short, robust humerus, the new specimen resembles adult and subadult material currently attributed to Y. tenuis, but it differs in having bicuspid posterior teeth and a straight rather than a hooked angular process of the mandible. We propose a new species (Y. bicuspidens sp. nov.) for this specimen which represents the first occurrence of bicuspid teeth in an Early Cretaceous lizard from China. However, the recognition of a second species at the neotype locality raises a taxonomic problem. Due to the immaturity of the designated holotype and neotype of Y. tenuis, they cannot be coded for the jaw and dental characters that distinguish the two robust-limbed species. This renders Yabeinosaurus tenuis a nomen dubium. Here we propose that Yabeinosaurus tenuis should be treated as a historic taxon, permitting retention of the generic name under ICZN rules, and erect a second new species (Y. robustus sp. nov.) for the principal robust-limbed morphotype in the Jehol Biota with monocuspid posterior teeth and a hooked angular process. 相似文献
7.
The taxonomic status of specimens referred to Tarchia kielanae, T. gigantea, and Minotaurasaurus ramachandrani has been contested. The holotype of T. kielanae, Zaklad Paleobiologii (Institute of Paleobiology)-Polish Academy of Sciences (ZPAL) MgD I/111, is redescribed. It has features common to both Palaeontological Institute-Russian Academy of Sciences (PIN) 3142/250, long considered Tarchia, and to M. ramachandrani but which are lacking in Saichania chulsanensis. The specimen PIN 3142/250 is not referable to Saichania but instead represents a new species, Tarchia teresae sp. nov. Similarly, the holotype skull of M. ramachandrani is not referable to T. kielanae, so Minotaurasaurus is a valid taxon. Tarchia is more derived than either Saichania or Minotaurasaurus while sharing features of both. 相似文献
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Mark P. Witton 《Cretaceous Research》2009,30(5):1293-1300
A new species of the sail-crested pterosaur Tupuxuara is described from the Santana Formation of Brazil, Tupuxuara deliradamus sp. nov. The holotype, a partial skull, and a larger, partial skull referred to the same taxon differs from Tupuxuara leonardii by having a nasoantorbital fenestra with an acutely-angled posterior border with a long, straight posterodorsal margin, a reclined cranium, and an orbit situated entirely in the ventral half of the nasoantorbitral fenestra. Unfortunately, neither specimen is comparable with the fragmentary rostrum representing Tupuxuara longicristatus. In addition, resolution of a recent nomenclatural problem over the correct name for the clade containing Tupuxuara and its sister taxon, Thalassodromeus, is provided. Both genera are used by different authors as the nomenclatural basis for the group, but “Tupuxuaridae” has never been explicitly erected as a new taxon, and therefore fails to meet ICZN criteria that new taxa are only valid if authors clearly indicate their intention to establish new names. By contrast, “Thalassodrominae” was explicitly erected as a name for the Thalassodromeus + Tupuxuara clade, thereby fulfilling all ICZN requirements for naming of a new taxon and making Thalassodromeus stand as the type genus for this group. 相似文献
10.
New Middle Turonian mosasauroid remains were discovered in the same large-sized nodule that yielded a specimen referred to Tethysaurus nopcsai Bardet et al., 2003, from the Goulmima region (southern Morocco). They comprise isolated, fragmentary cranial elements (skull and mandible) and some vertebrae. Their very small size suggests a juvenile condition, an observation supported also by anatomical (spongious nature of numerous bone parts), micro-anatomical (loose inner spongiosa) and histological (numerous remains of calcified cartilage deep into the centrum; radial vascular canals) data. These bones belonged to a basal mosasauroid that cannot be distinguished from Tethysaurus nopcsai to which taxon we tentatively assign the material. 相似文献
11.
Jaco Weinstock Beth Shapiro Alfredo Prieto Juan Carlos Marín Benito A. González M. Thomas P. Gilbert Eske Willerslev 《Quaternary Science Reviews》2009,28(15-16):1369-1373
We present the results of a palaeogenetic analysis of two Late Pleistocene camelids originating in southern Chile. Our analysis of two mitochondrial DNA fragments (control region and cytochrome b gene) reveals that these specimens do not belong to an extinct taxon, but rather to extant vicuña (Vicugna vicugna), whose modern distribution is restricted to the extreme elevations of the Andes, more than 3500 km to the north of where these specimens originated. Our results also suggest fossil specimens from Patagonia that are currently assigned to the extinct taxon Lama gracilis, may actually belong to V. vicugna, implying a continuous distribution of the latter from the southern tip of South America to the Andes during the Final Pleistocene. The haplotypes of both specimens are not present in modern populations, suggesting a loss of genetic diversity concomitant with the contraction of the vicuña geographical distribution during the Final Pleistocene or early Holocene. 相似文献
12.
WANG Xuri Jingmai K. O’Connor ZHAO Bo Luis M. Chiappe Gao Chunling Cheng Xiaodong 《《地质学报》英文版》2010,84(2):247-256
<正>We report on a new species of enantiornithine bird from the Lower Cretaceous Qiaotou Formation of northern Hebei,China.The new taxon,Shenqiornis mengi gen.et sp.nov.,possesses several enantiornithine synapomorphies but is unique from other known species.The specimen has a well-preserved skull that reveals new information about enantiornithine cranial morphology.The new taxon possesses a large postorbital with a long tapering jugal process indicating that some enantiornithines may have had a fully diapsid skull,as in Confuciusornis.The tooth morphology of the specimen is unique and likely represents a previously unknown trophic specialization within Enantiornithes. 相似文献
13.
新铺龙是产于我国西南地区的海龙目的一个属,最早发现于贵州省关岭晚三叠世卡尼期小凹组地层。虽然新铺龙材料众多,但其未成年个体骨骼学特征及个体发育模式并未被详细地记述。本文报道了一个保存完好的孙氏新铺龙未成年标本,是目前发现最小的完整的孙氏新铺龙个体,标本产自贵州省关岭县新铺乡。依据其体长小于正型标本的一半,头骨长度和躯干比值大,头骨骨缝清晰等特征,判断该标本为孙氏新铺龙的幼年个体。新标本补充了孙氏新铺龙幼年个体的形态学信息,如上颞骨内侧突短于外侧突,翼骨无牙齿发育,神经弓与椎体未融合,尺骨两端微凸,五边形坐骨等。此外,结合已发表的新铺龙标本数据进行统计分析发现,该物种吻部相对头部呈负异速生长,证明了吻部占头骨比例的大小属于新铺龙种内个体发育差异,而并非之前所认为的新铺龙种间差异。 相似文献
14.
新铺龙是产于我国西南地区的海龙目的一个属,最早发现于贵州省关岭晚三叠世卡尼期小凹组地层。虽然新铺龙材料众多,但其未成年个体骨骼学特征及个体发育模式并未被详细地记述。笔者等报道了一个保存完好的孙氏新铺龙未成年标本,是目前发现最小的完整的孙氏新铺龙个体,标本产自贵州省关岭县新铺乡。依据其体长小于正型标本的一半,头骨长度和躯干比值大,头骨骨缝清晰等特征,判断该标本为孙氏新铺龙的幼年个体。新标本补充了孙氏新铺龙幼年个体的形态学信息,如上颞骨内侧突短于外侧突,翼骨无牙齿发育,神经弓与椎体未融合,尺骨两端微凸,五边形坐骨等。此外,结合已发表的新铺龙标本数据进行统计分析发现,该物种吻部相对头部呈负异速生长,证明了吻部占头骨比例的大小属于新铺龙种内个体发育差异,而并非之前所认为的新铺龙种间差异。 相似文献
15.
There has been considerable debate about whether the controversial tyrannosauroid dinosaur ‘Nanotyrannus lancensis’ from the uppermost Cretaceous of North America is a valid taxon or a juvenile of the contemporaneous Tyrannosaurus rex. In a recent Cretaceous Research article, Schmerge and Rothschild (2016) brought a new piece of evidence to this discussion: the morphology of the dentary groove, a depression on the lateral surface of the dentary that houses neurovascular foramina. They argued that an alleged ‘Nanotyrannus’ specimen, which possesses a groove, cannot be referable to Tyrannosaurus rex, which they considered as lacking the groove, and they hypothesized that ‘Nanotyrannus’ is closely related to albertosaurine tyrannosauroids, which also are said to possess the groove. However, we show that the groove is a widespread feature of tyrannosauroids that is present in T. rex and many other specimens, and that it is an ontogenetically variable feature that changes from a sharp, deeply-impressed groove to a shallower sulcus as an individual matures. As a result, the presence or absence of a dentary groove does not clarify the validity of ‘Nanotyrannus’ or its phylogenetic position among tyrannosauroids. We consider it most parsimonious that ‘Nanotyrannus’ specimens belong to juvenile T. rex. 相似文献
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18.
Luc G. Bulot Jean-Louis Latil Alex Hairabian Arnaud Fournillon 《Proceedings of the Geologists' Association. Geologists' Association》2014
Nolaniceras nolani ( Seunes, 1887) has been widely quoted in the Upper Aptian literature over the years. Re-examination of the holotype of the species shows that it has always been misidentified and that this taxon is represented by a single specimen, the holotype, of uncertain age. As a consequence, its use as an index species for the Upper Aptian should be reconsidered and abandoned. 相似文献
19.
Here we describe the well-preserved skull of a juvenile specimen of Sapeornis, STM 16-18. Only the tail and hindlimb plumage of this specimen have been previously described. It preserves what we consider to be the complete dentition of Sapeornis with four premaxillary teeth, three maxillary teeth and two tiny dentary teeth on each side, the latter feature being previously unknown for this taxon. Based on a study of 71 specimens of Sapeornis preserving dentition, we infer the absence of dentary teeth in previously described specimens of Sapeornis is taphonomic, and consider the true dental formula of Sapeornis to be 4-3-2 (premaxillary teeth number – maxillary teeth number – dentary teeth number). Rostral to the dentary teeth, STM 16-18 preserves three shallow, empty aveoli. We hypothesize that this morphology captures an early stage in dental reduction and indicates that in at least some avian lineages, the loss of teeth was gradual. 相似文献
20.
The fluvio-lacustrine deposits of the ?Cenomanian-Turonian Candeleros Formation in the northwestern fringe of Patagonia have yielded numerous remains of vertebrates, including anurans. A new, partially articulated immature specimen of a small pipimorph pipoid from the upper part of this unit is described herein. Although incompletely preserved, the morphology and proportions of the skeleton are strongly reminiscent of corresponding elements of the holotype of Avitabatrachus uliana from the lower section of the same formation at a nearby locality, thus suggesting that these specimens might represent the same, or a closely related, taxon. The new specimen is a metamorphosing individual: it has a zygapophyseal articulation between the sacrum and the neural arch of a postsacral vertebra that bears conspicuous transverse processes, and four pairs of ribs. These traits are not present in the available material of A. uliana, but they might have occurred within the normal range of variation of a single species. Although the generic and specific allocation of the new material is uncertain, a parsimony analysis performed to clarify the relationships of A. uliana supports a phylogenetic placement of this taxon outside the crown Pipidae. The evidence provided by the Candeleros anurans confirms the presence of stem pipids in Patagonia in the early Late Cretaceous. 相似文献