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1.
A. I. Kafanov 《Oceanology》2008,48(2):233-238
Among the components of biological diversity such as taxonomic richness and ecological diversity (distribution of species with respect to their abundance), one should also distinguish the taxonomic diversity as a function of taxonomic richness and evenness of the distributions of taxa of lower ranks over those of higher rank. In the direction from the high latitudes to the tropics, the taxonomic richness and Shannon’s index of taxonomic diversity regularly increase, while the taxonomic evenness, on the contrary, decreases. Toward the north, a relatively small number of mass species become better manifested; they are approximately even over a few genera and families. In warm waters, more common is the existence of a relatively large number of families, which strongly differ in the number of species; each of the latter features low abundance parameters. The meridional asymmetry in the distribution of the taxonomic richness and diversity reveals itself in the higher values of these parameters in the northeast of the Pacific Ocean as compared to their values at the same latitudes in the northwest; in the former case, the trend of latitudinal variations in the taxonomic diversity is poorer expressed.  相似文献   

2.
Estimating diversity of modular organisms may be problematic due to actual difficulties in discriminating between 'individuals' and quantifying their abundances. Quantitative data, when available, are collected through methods that could preclude the application of classical diversity indices, making comparisons among studies difficult. Taxonomic distinctness indices, such as the 'Average Taxonomic Distinctness' (Δ+) and the 'Variation in Taxonomic Distinctness' (Λ+) may represent suitable tools in investigating diversity beyond the simple species number. The potential usefulness of such indices has been explored almost exclusively on unitary organisms, neglecting modular ones. In this study, we employed Δ+ and Λ+ to analyse patterns of diversity of epiphytic hydroid assemblages living on Cystoseira seaweeds at a hierarchy of spatial scales, along 800 km of rocky coast (SE Italy). ANOVA on species richness and Λ+ showed no significant difference in sample diversity at the investigated spatial scales. In contrast, there were significant differences at the scale of 10s of km in Δ+. Analyses based on simulations detected significant variations at all spatial scales in Δ+. Such findings underline the potential of Δ+ in highlighting relevant spatial scales of variation in patterns of hydroid diversity. Our results also suggest that the interplay between natural environmental variations and the complex ecological traits of modular organisms might affect taxonomic distinctness indices. We stress the need for further investigations focusing on modular organisms before any generalizations on the use of taxonomic relatedness measures in examining marine biodiversity can be made.  相似文献   

3.
During the SoJaBio expedition, the deep sea fauna of the north-western Sea of Japan was sampled in August–September 2010. From this study, 11 epibenthic sledge stations are analyzed, with a focus on species composition, diversity and distribution patterns of polychaetes. A total of 92 polychaete taxa belonging to 70 genera and 28 families and 3 indeterminate species were found. Twelve species and eight genera have not been reported from the Sea of Japan before, but were registered from other deep-sea basins. Calculation of diversity (Shannon–Wiener Index, Pielou's Evenness) showed that the upper bathyal of the Sea of Japan is an area of higher polychaete diversity than the abyssal plain. The increased richness and diversity here could possibly be explained by a zoogeographic overlapping with the shallower species' assemblages of the shelf. At a higher taxonomic level the polychaete fauna of the deep Sea of Japan does not seem to differ from that of other deep-sea regions world-wide. In depths below 2000 m about 30% polychaete species have wide distributional ranges.  相似文献   

4.
The results of a study performed before and after the discharge of an ocean outfall off Mõkapu Point, Oahu, Hawaii, from 1975 to 1979 show large increases in the number of species and abundance of fish around the outfall site. The construction of the outfall afforded new rock substratum for the attachment of algae and sessile animals which attracted large aggregations of fish. During a 112 year period there were no significant community structure 1–5 km from the outfall when measured by similarity indices. No significant changes in algal biomass occurred between samplings.Similarity indices may be better indicators of fish community change than diversity indices since the former reflect changes in the relative abundance of species in common and large temporal changes in a community structure may occur without changing the value of its diversity index. Morisita's (1959) similarity index has greater sensitivity to the changes in fish community structure than the index derived by Sorensen (1954) due to emphasis on the relative abundance of dominant species.  相似文献   

5.
黄河口鱼类群落分类学多样性的研究   总被引:2,自引:2,他引:0  
根据2013-2014年在黄河口水域进行的7个航次的鱼类资源底拖网调查数据,应用分类多样性指数分析了黄河口水域鱼类群落生物多样性的时空变化。黄河口鱼类主名录的平均分类差异指数△+理论平均值为75.5。2013-2014年调查中,该水域共捕获鱼类51种,隶属10目、27科、43属,以鲈形目、鲱形目和鲉形目鱼类居多。本次调查中鱼类群落分类多样性指数△和分类差异指数△*月变化范围分别为39.98~65.48和59.15~75.54,均表现为夏季月份最高,春、秋季月份次之,冬季最低。除春季月份外,其他月份鱼类平均分类差异指数△+均显著低于黄河口鱼类主名录的△+理论平均值。不同月份中平均分类差异指数△+也呈现一定的空间变化,夏、秋季月份绝大部分站位平均分类差异指数处于95%置信范围内,春、冬季月份个别站位的平均分类差异指数显著低于理论平均值,这与这些站位绝大多数种类集中在鲈形目有关。黄河口水域鱼类分类学多样性具有明显的时空变化,鱼类资源衰退,应加强该水域鱼类资源养护和多样性保护。  相似文献   

6.
渤海自由生活海洋线虫多样性的研究   总被引:5,自引:0,他引:5       下载免费PDF全文
首先介绍Warwick和Clarke(1995)提出的等级生物多样性指数以及在自由生活海洋线虫中的应用,重点以1998年9~10月和1999年4~5月两个航次的沉积物样品,用物种数目(S)、Margalef物种丰富度指数(d)、优势度Simpson指数(λ)、多样性Shannon-Wiener指数(H')、Pielou的均匀度指数(J)、Hill多样性系列指数(N1,N2和N+∞)、分类多样性指数(△)、分类差异指数(△*)、等级多样性指数(△+)对渤海自由生活海洋线虫的多样性进行了研究。结果表明,黄河输入的泥沙对渤海自由生活海洋线虫多样性的变化有一定的作用,距离黄河口越近的站位,群落具有的物种多样性和均匀性越低,优势度越高;距离黄河口越远的站位,群落具有的物种多样性和均匀性越高,优势度越低。  相似文献   

7.
Species diversity is generally considered one of the key factors of ecosystem resilience in response to anthropogenic pressures, including fishing. In this context, the spatial and temporal changes in demersal fish assemblages and species diversity were investigated in the northern Gulf of St. Lawrence (Canada), over a 20‐year period (1990–2010). Data were obtained from the summer research survey conducted by the Department of Fisheries and Oceans, and include commercial and non‐commercial species. The study covers the period of groundfish fishery collapse, the moratorium period, and the post‐moratorium period, and reflects various modifications in management. Multivariate statistical methods revealed two communities. A coastal community corresponds to strata located above 200 m depth and a deeper community located in the deep channels. Interannual differences in the composition of fish assemblages were observed and are mainly due to the changes in the relative biomass of some dominant species. Three diversity indices (Shannon–Wiener, Simpson's Index of Diversity and Motomura's constant) indicate a slight but significant increase of the diversity over time. This trend is due to the increase of the relative biomass of low‐rank species, which may have been favoured by the prohibition of groundfish trawling after 1997 in that region. The geographical distribution of the Shannon–Wiener index also shows temporal dynamics reflecting the biomass distribution of dominant demersal species.  相似文献   

8.
Taxonomic diversity of fish assemblages in the Changjiang Estuary(Yangtze River Estuary) and its adjacent waters was analyzed based on bottom trawl surveys carried out by R/V Beidou in June,August and October 2006.Four groups of fishes were identified for each survey by the twoway indicator species analysis(TWIA).Taxonomic distinctness,species richness and phylogenetic diversity were used to detect changes in taxonomic structure of fish assemblages.Most dominant fish species mainly belonged to Perciformes,Pleuronectiformes,Clupeiformes,Scorpaeniformes and Tetraodontiformes.Species richness in offshore waters of the survey area was higher than that in the other area.The average taxonomic distinctness value(AvTD,△+ ) of each fish assemblage was close to the average taxonomic distinctness of master list in the Changjiang Estuary(79.9),and had a positive correlation with species richness and Shannon diversity index(H ).A negative correlation was found between variations in taxonomic distinctness(VarTD,Λ + ) and traditional diversity indices,which were caused by fish species component that led to longer average path lengths among species.Taxonomic diversity index(△) had the similar results with species richness,H ,Simpson diversity index(D) and Pielou's evenness index(J ).VarTD also kept stable,which further certificated that fish assemblages and ecological environment were in equilibrium.Taxonomic distinctness index(△* ) was relatively stable in most of fish assemblages,and had higher values in some fish assemblages for a few absolutely dominant species.The present study showed that fish community formed new equilibrium stability in 2006 in the Changjiang Estuary when compared with those in the corresponding months of annual survey from 1985 to 1986.And AvTD in each sampling station was lower than AvTD of master list in the Changjiang Estuary,so some ecological niches were absent in each sampling stations from taxonomic or phylogenetic relationships,these would be helpful to stock natural resource and maintain ecological equilibruim of fish assemblages.  相似文献   

9.
To define the functional groups of fish and macroinvertebrates in the Eastern Tropical Pacific (ETP), visual censuses were performed in 18 areas of four biogeographic provinces: Cortés, Mexican, Panamic and Oceanic Islands. A total of 257 fish and macroinvertebrate species were recorded, and from them, 27 morpho‐functional groups (MFGs) were constructed on the basis of trophic level, maximum size, taxonomy and morphology. Biomass, richness, diversity and evenness of MFGs were calculated for each province and compared statistically; a regression analysis between taxonomic and functional diversity was conducted to observe the relationship between these two indicators. There were significant differences in all ecological indices (p < .002), highlighting the high biomass and richness of MFGs in the Cortés and Oceanic Islands provinces (>400 g/m2), associated with the influence of cold currents in the northernmost region and less fishing pressure in protected areas. A decreasing pattern of MFG richness towards the tropics was observed, which demonstrates that in the ETP, the relationship between habitat heterogeneity and species diversity has been translated into functional complexity. The Mexican province was the most functionally diverse (biomass well distributed in the MFG; H′ = 0.46 ± 0.009). Related to this, it is predicted that biomass is biased towards certain functional groups (i.e., large carnivores), which shows that the H′ index of the MFG is not a good indicator of the conservation status of ETP reefs. Finally, regression analysis suggests that functional diversity increases at low species diversity but eventually reaches an asymptote (almost all possible functions are represented).  相似文献   

10.
Nematode assemblages were investigated (in terms of size spectra, sex ratio, Shannon diversity, trophic structure and diversity, rarefaction statistics, maturity index, taxonomic diversity and taxonomic distinctness) at bathyal and hadal depths (from 1050 to 7800 m) in the deepest trench of the South Pacific Ocean: the Trench of Atacama. This area, characterised by very high concentrations of nutritionally-rich organic matter also at 7800-m depth, displayed characteristics typical of eutrophic systems and revealed high nematode densities (>6000 ind. 10 cm−2). Nematode assemblages from the Atacama Trench displayed a different composition than at bathyal depths. At bathyal depths 95 genera and 119 species were found (Comesomatidae, Cyatholaimidae, Microlaimidae, Desmodoridae and Xyalidae being dominant), whereas in the Atacama Trench only 29 genera and 37 species were encountered (dominated by Monhysteridae, Chromadoridae, Microlaimidae, Oxystominidae and Xyalidae). The genus Monhystera (24.4%) strongly dominated at hadal depths and Neochromadora, and Trileptium were observed only in the Atacama Trench, but not at bathyal depths. A reduction of the mean nematode size (by ca. 67%) was observed between bathyal and hadal depths. Since food availability was not a limiting factor in the Atacama Trench sediments, other causes are likely to be responsible for the reduction of nematode species richness and body size. The presence of a restricted number of families and genera in the Atacama Trench might indicate that hadal sediments limited nematode colonisation. Most of the genera reaching very high densities in Trench sediments (e.g., Monhystera) are opportunistic and were responsible for the significant decrease of the maturity index. The dominance of opportunists, which are known to be characterised by small sizes, might have contributed to the reduced nematode size at hadal depths. Shannon diversity and species richness decreased in hadal water depth and this pattern was more evident at genus than at species level. Epistrate feeders dominated and increased their relevance, determining a reduction of the index of trophic diversity at hadal depths. According to trophic diversity, taxonomic diversity and distinctness also decreased with depth. All diversity indices from the Atacama Slope and Trench were lower than in other equally deep areas world wide (e.g. Puerto Rico Trench). We suggest that such reduction was related to the high nutrient loading observed in this system (up to two orders of magnitude higher than in typical oligotrophic deep-sea sediments).  相似文献   

11.
Revillagigedo Archipelago and Clipperton atoll are oceanic islands located in the eastern tropical Pacific. The composition and trophic structure of their fish assemblages have never been characterized in detail, and the aim of this study was therefore to analyse the community structure and assess the similarity of fish assemblages between these two areas. Underwater visual fish censuses were conducted using band transects at two depth levels in three islands of the Revillagigedo Archipelago (Socorro, San Benedicto, and Roca Partida) and at Clipperton atoll. We calculated ecological indices (density, richness, Shannon diversity, Pielou evenness, and taxonomic distinctiveness) for each transect. These were compared in order to determine quantitative differences among the islands and bathymetric levels. Qualitative analyses (ANOSIM, nMDS, and SIMPER) were also performed. Finally, we analysed the trophic structure of the assemblages and the functional diversity of the four studied sites. The results showed significant differences for all indices, except diversity (H′). Roca Partida was the site with greatest abundance and richness due to the presence of pelagic species in large numbers. No significant differences were found in functional diversity, suggesting a high degree of ecological redundancy. Lower‐level carnivores with mean trophic level values of between 3.3 and 3.8 dominated both areas. The ordination analysis did not show a clear difference in fish composition among the islands, although Clipperton atoll has a particular fauna characterized by endemic species (Thalassoma robertsoni and Stegastes baldwini). In conclusion, the fish communities of the studied areas seem quite similar, but their assemblage structure differs. In particular, Clipperton is differentiated by the influence of endemic species.  相似文献   

12.
We examined tintinnid (loricate ciliate microzooplankton) diversity using data from 11 stations between the Moroccan upwelling system and the oligotrophic Eastern Mediterranean. Taxonomic and morphological diversity of tintinnids was compared to phytoplankton distribution and size-structure, to the abundance of competitors in the form of oligotrich ciliates, and predators as copepods. Tintinnid taxonomic diversity was estimated as numbers of species and the Shannon Index, H′; morphological diversity was quantified by substituting size classes of lorica dimensions for species. Total chlorophyll was partitioned into micro-, nano- and pico-fractions using pigment data and a size-diversity was estimated by considering the 3 size classes as 3 species. Along a west-to-east gradient, average water column concentrations of most organism groups declined approximately an order of magnitude yielding tight correlations. However, tintinnid diversity, both taxonomic and morphological, increased from the Atlantic upwelling station into the western basin of the Mediterranean, and declined slightly towards the Eastern Mediterranean, paralleling shifts in the chlorophyll size-diversity estimate. Diversity varied with absolute or relative abundance of oligotrich or copepods, but different diversity metrics were significantly correlated only with phytoplankton size-diversity. We conclude that tintinnid diversity more closely reflects resource diversity than competitive interactions or predation.  相似文献   

13.
The response of phytoplankton assemblage structure to terrestrial nutrient inputs was examined for the Gulf of Kalloni in the Northern Aegean Sea, a productive semi-enclosed coastal marine ecosystem. The study was focused on a typical annual cycle, and emphasis was placed on the comparative analysis between blooms developing after significant nutrient inputs from the watershed, and naturally occurring blooms. Baseline information was collected on a monthly basis from a network of stations located in the oligotrophic open sea and the interior and more productive part of the embayment. Intensive sampling was also carried out along a gradient in the vicinity of a river which was the most important source of freshwater and nutrient input for the Gulf. Phytoplankton assemblage structure was analyzed from 188 samples using diversity indices (Shannon and Average Taxonomic Distinctness), multivariate plotting methods (NMDS), multivariate statistics (PERMANOVA), and canonical correspondence analysis (CCA). Three characteristic assemblages were recognized: (1) an autumn assemblage developed under nutrient depleted conditions, having low diversity due to the dominance of two small diatoms, (2) a winter bloom of the potentially toxic species Pseudo-nitzschia calliantha occurring immediately after a nutrient peak and characterized by very low diversity, and (3) a naturally occurring early summer bloom of centric diatoms with relatively high diversity. The results of the study support the view that moderate nutrient inputs may have a beneficial effect on the functioning of coastal ecosystems, stimulating the taxonomic diversity through the growth of different taxonomic groups and taxa. On the other hand, a sudden pulse of high nutrient concentrations may greatly affect the natural succession of organisms, have a negative effect on the diversity through the dominance of a single species, and can increase the possibility of a harmful algal bloom development.  相似文献   

14.
Regular collections of fish were obtained from the surf-zone at King's Beach, Algoa Bay. A total of 3970 fish, representing 50 species was caught with a coarse net and 16 857 fish, representing 37 species, were caught with a fine net. Predominant species were the blacktail, Diplodus sargus; the sand steenbras, Lithognathus mormyrus; the mullet, Liza richardsoni; the gorrie, Pomadasys olivaceum; the white stumpnose, Rhabdosargus globiceps; the sandshark, Rhinobatos annulatus; and the streepie, Sarpa salpa. No seasonal trends were discernible in the overall abundance or species diversity. The species composition of the dominant component of the fish assemblage varied considerably. This indicated instability in the community structure and cast doubts on the applicability of a classic community concept and the use of diversity indices. Neither classification nor correspondence analysis were of any use in identifying a characteristic species component. Multiple regression analysis indicated that short-term variations in wind conditions might be a primary determinant of fluctuations in abundance. The lack of seasonality in the community parameters may reflect the fact that short-term variability masks seasonal perturbations.  相似文献   

15.
胶州湾底栖生物多样性初探   总被引:21,自引:5,他引:21  
毕洪生  冯卫 《海洋科学》1996,20(6):58-62
用1991-1995年胶州湾10个监测站的资料,采用Shannon-Weaver指数,Pielous均匀度指数,种的丰盛度指数以及Sanders稀疏曲线进行大型底栖生物多样性比较。结果表明,在胶州湾,粉砂-粘土混合型的沉积环境中大型底栖生物多样性高于匀质的粗砂底质。  相似文献   

16.
Short-term changes in the King's Beach surf-zone fish assemblage were investigated by means of two 24 h studies (January and August 1979), and by seine netting on three consecutive days at both high and low tide. They revealed considerable variation in the fish assemblage over a 24 h period. Four species: the gorrie, Pomadasys olivaceum; the mullet, Liza richardsoni; the streepie, Sarpa salpa; and the sand steenbras, Lithognathus mormyrus, dominated the catches of both 24 h studies. The number of species caught reached a peak just after twilight. Trends in other community parameters were not consistent for these two sets of data. Considerable variation in dominant species, numbers and species diversity indices was observed from day to as well as between high and low tide. Shortterm fluctuations in various community parameters often exceeded the long-term variability observed in a seasonal study of this fish assemblage.  相似文献   

17.
A brackish-water cold seep on the North Anatolian Fault (NAF) in the Marmara Sea was investigated with the Nautile submersible during the MarNaut cruise in 2007. This active zone has already been surveyed and revealed evidence of active seeping on the seafloor, such as bubble emissions, patches of reduced sediments, microbial mats and authigenic carbonate crusts. MarNaut was the first opportunity to sample benthic communities in the three most common microhabitats (bioturbated and reduced sediments, carbonate crust) and to examine their relationships with environmental conditions. To do so, faunal communities were sampled and chemical measurements were taken close to the organisms. According to diversity indices, the bioturbated microhabitat exhibited the highest taxonomic diversity and evenness despite a lower number of samples. Conversely, the reduced sediment microhabitat exhibited the lowest taxonomic diversity and evenness. The carbonate crust microhabitat was intermediate although it had the highest biomass. Multivariate analyses showed that (1) fauna were relatively similar within a single microhabitat; (2) faunal community structure varied greatly between the different microhabitats; (3) there was a link between faunal distribution and the type of substratum; and (4) chemical gradients (i.e. methane, oxygen and probably sulphides) may influence faunal distribution. The estimated fluid flow velocity (0.4–0.8 m/yr) confirmed the presence of fluid emission and provided evidence of seawater convection in the two soft-sediment microhabitats. Our results suggest that the reduced sediments may represent a harsher environment with high upward fluid flow, which restrains seawater from penetrating the sediments and inhibits sulphide production, whereas bioturbated sediments can be viewed as a bio-irrigated system with sulphide production occurring at greater depths. Therefore, the environmental conditions in reduced sediments appear to prevent the colonization of symbiont-bearing fauna, such as vesicomyid bivalves, which are more often found in bioturbated sediments. Fluid flow appears to control sulphide availability, which in turn influences the horizontal and vertical distribution patterns of fauna at small spatial scales as observed at other seep sites.  相似文献   

18.
19.
A study was carried out across the continental shelf and slope in the Western Indian Ocean along the Kenyan margin to unravel depth‐related species diversity patterns portrayed by different nematode families. Sediment samples were collected along four bathymetric transects at 20, 50, 200, 500, 1000 and 2000 m. Three nematode families were selected for species analysis, based on their general occurrence with relatively high numbers and diversity from shelf to slope. All three families exhibited a distinctly different shelf and slope nematode species community. However, all three families also had a significant proportion of species that displayed a eurybathic distribution. Coincidentally, Microlaimidae, the least species‐rich family had the most eurybathic species (75%) compared to Comesomatidae (39%) and Chromadoridae (32%). Total number of species per depth zone along the sampled area (gamma diversity) was three to four times the average number of species (alpha) per station. The difference was more pronounced at the slope than at the shelf. The species turnover was also higher at the slope than at the shelf stations. Each of the three families had a different diversity pattern: Comesomatidae showed a peak at mid‐depth, Chromadoridae increased with depth, and Microlaimidae showed no prominent change with depth. When the three families were combined, the shelf maintained a lower diversity compared to the slope (both local and regional). There was no consistency between genus and species diversity patterns with depth, indicating that genus diversity data may not be a proxy for species diversity. At the lowest taxonomic level (species), the slope showed a higher diversity than the shelf, whereas at higher (genus) taxonomic level the diversity was comparable between the slope and the shelf. The number of species encountered increased with the number of samples analysed and did not reach asymptote, meaning that the area was still under‐sampled. This situation points to the possibility of an even higher regional diversity.  相似文献   

20.
Differences in macroinvertebrate community structure and composition were examined from April 1980 to March 1981 in three potholes that had been ditched for mosquito control and three natural (i.e. unditched) potholes, which are located in a San Francisco Bay, California, U.S.A. salt marsh. Measurements of incipient tidal flooding into potholes (i.e. pothole inundation threshold) indicated that these sites comprise a gradient of tidal influences. Exponential decreases in the frequency and duration of tidal inundation corresponded to linear increases in inundation threshold. Since ditched study sites had low thresholds they tended to be more uniformly and regularly influenced by tides, were less saline, had less variable temperature regimens, and supported less filamentous algae than natural potholes. Habitat conditions were generally more similar among ditched than unditched potholes, but environmental conditions were most severe at natural sites near the upper limit of the inundation threshold gradient, where some potholes desiccate during the dry season each year.Differences in macroinvertebrate communities corresponded to differences in habitat conditions. Species richness and diversity (Simpson's Index) were generally highest near the middle of the inundation threshold gradient, which is a pattern predicted by the Intermediate Disturbance Hypothesis. Analysis of faunal composition using discriminant functions indicated more similarity among potholes located at the lowest positions of the inundation gradient than among potholes with intermediate thresholds. Since ditching lowers the inundation thresholds of potholes, it reduces species richness and diversity, while increasing faunal similarity. As a result, extensive ditching to control salt marsh mosquitoes can reduce the overall complexity of lentic macroinvertebrate communities.  相似文献   

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