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1.
A review of the tetrapod (amphibian and amniote) record across the Permo-Triassic boundary (PTB) indicates a global evolutionary turnover of tetrapods close to the PTB. There is also a within-Guadalupian tetrapod extinction here called the dinocephalian extinction event, probably of global extent. The dinocephalian extinction event is a late Wordian or early Capitanian extinction based on biostratigraphic data and magnetostratigraphy (the extinction precedes the Illawara reversal), so it is not synchronous with the end-Guadalupian marine extinction. The Russian PTB section documents two tetrapod extinction events, one just before the dinocephalian extinction event and the other at the base of the Lystrosaurus assemblage. However, generic diversity across the latter extinction remains essentially the same despite a total evolutionary turnover of tetrapod genera. The Chinese and South African sections document the stratigraphic overlap of Dicynodon and Lystrosaurus. In the Karoo basin, the lowest occurrence of Lystrosaurus is in a stratigraphic interval of reversed magnetic polarity, which indicates it predates the marine-defined PTB, so, as previously suggested by some workers, the lowest occurrence of Lystrosaurus cannot be used to identify the PTB in nonmarine strata. Correlation of the marine PTB section at Meishan, southern China, to the Karoo basin based primarily on magnetostratigraphy indicates that the main marine extinction preceded the PTB tetrapod extinction event. The ecological severity of the PTB tetrapod extinction event has generally been overstated, and the major change in tetrapod assemblages that took place across the PTB was the prolonged and complex “replacement” of therapsids by archosaurs that began before the end of the Permian and was not complete until well into the Triassic. The tetrapod extinctions are not synchronous with the major marine extinctions at the end of the Guadalupian and just before the end of the Permian, so the idea of catastrophic causes of synchronous PTB extinctions on land and sea should be reconsidered.  相似文献   

2.
The fossil record of mid to late Permian terrestrial vertebrates in the South African Karoo Basin is regarded as the most abundant and diverse in the world. Despite the extensive research on body fossils, to-date the vertebrate footprint sites have not been subjected to an anatomy-consistent ichnotaxonomic investigation. Here we present a comprehensive ichnotaxonomic revision of Permian-Early Triassic tracksites in the main Karoo Basin of South Africa. Furthermore, a track-trackmaker correlation for all Permian synapsid groups is provided for the first time, based on the analysis of the functional morphology of potential producers. The following ichnotaxa and their proposed trackmakers are recognized: Brontopus giganteus (dinocephalians), cf. Capitosauroides isp. (therocephalians), cf. Dicynodontipus isp. (cynodonts), Dolomitipes accordii (small dicynodonts), Dolomitipes icelsi n. comb. (large dicynodonts), Karoopes gansfonteinensis n. igen. n. isp. (gorgonopsids), Procolophonichnium nopcsai (procolophonids) and Rhynchosauroides isp. (non-archosauriform diapsids). Three different footprint assemblages (FA I–III) are proposed for footprint biostratigraphy: FA I (lower Tapinocephalus AZ), a Guadalupian assemblage dominated by dinocephalian tracks; FA II (topmost Tapinocephalus-Cistecephalus AZ), a latest Guadalupian-Wuchiapingian assemblage dominated by gorgonopsid and dicynodont tracks in association with subordinate therocephalian tracks and FA III (lower Lystrosaurus AZ), an Induan assemblage with dicynodont, cynodont, procolophonid and diapsid tracks. The lower FA II includes the earliest ichnofauna with Lopingian affinity all over the world (topmost Tapinocephalus-Pristerognathus AZ, ~260–259 Ma) and could indicate an early recovery phase after the end-Guadalupian mass extinction, because of the high abundance of large gorgonopsid tracks and absence of dinocephalian tracks. This footprint record may also predate the body fossil record, suggesting an earlier gorgonopsid radiation. FA III represents the earliest and most complete post end-Permian extinction ichnofauna, which includes an early phase of abundant small dicynodont tracks, potentially indicating a stressed post-event community. Nevertheless, this ichnofauna looks very similar to pre-extinction ichnofaunas from Europe, in agreement with the skeletal record at the Daptocephalus-Lystrosaurus AZ transition.  相似文献   

3.
A review of the tetrapod (amphibian and amniote) record across the Permo-Triassic boundary (PTB) indicates a global evolutionary turnover of tetrapods close to the PTB. There is also a within-Guadalupian tetrapod extinction here called the dinocephalian extinction event, probably of global extent. The dinocephalian extinction event is a late Wordian or early Capitanian extinction based on biostratigraphic data and magnetostratigraphy (the extinction precedes the Illawara reversal), so it is not synchronous with the end-Guadalupian marine extinction. The Russian PTB section documents two tetrapod extinction events, one just before the dinocephalian extinction event and the other at the base of the Lystrosaurus assemblage. However, generic diversity across the latter extinction remains essentially the same despite a total evolutionary turnover of tetrapod genera. The Chinese and South African sections document the stratigraphic overlap of Dicynodon and Lystrosaurus. In the Karoo basin, the lowest occurrence of Lystrosaurus is in a stratigraphic interval of reversed magnetic polarity, which indicates it predates the marine-defined PTB, so, as previously suggested by some workers, the lowest occurrence of Lystrosaurus cannot be used to identify the PTB in nonmarine strata. Correlation of the marine PTB section at Meishan, southern China, to the Karoo basin based primarily on magnetostratigraphy indicates that the main marine extinction preceded the PTB tetrapod extinction event. The ecological severity of the PTB tetrapod extinction event has generally been overstated, and the major change in tetrapod assemblages that took place across the PTB was the prolonged and complex “replacement” of therapsids by archosaurs that began before the end of the Permian and was not complete until well into the Triassic. The tetrapod extinctions are not synchronous with the major marine extinctions at the end of the Guadalupian and just before the end of the Permian, so the idea of catastrophic causes of synchronous PTB extinctions on land and sea should be reconsidered.  相似文献   

4.
Two new tetrapod burrow casts from the Naobaogou Formation (Middle or Late Permian) of Nei Mongol,China are described.It marks the first pre-Cenozoic tetrapod burrow from China,and one of the earliest records of tetrapod burrows.Comparison to other Permian and Triassic burrows suggests that these burrows were created by tetrapod slightly smaller than Lystrosaurus.Deduced from the morphology and sizes of two burrows and known tetrapods of the Naobaogou Formation,the burrow should be the production of a therapsid,most likely a dicynodon.These burrows indicate a seasonal climate and this area was semiarid or arid during that time.  相似文献   

5.
白垩纪四足动物足印的生物地层学、生物年代学与遗迹相   总被引:3,自引:2,他引:1  
从全球范围来看,白垩纪四足动物的足印多数是非鸟恐龙与鸟类留下的痕迹;少量足印来自翼龙、鳄鱼、龟、哺乳动物和其他四足动物。白垩纪的足迹化石以东亚(尤其是中国和朝鲜)和北美西部的最为人所知。南美(主要是阿根廷和巴西)也有一定数量广泛分布的足迹化石,欧洲、非洲与澳大利亚的白垩纪足迹组合则鲜为人知。以白垩纪四足动物的足印记录为基础,我们对两个全球足印生物年代重新进行了检查。早白垩世生物年代以蜥脚类与鸟脚类的足迹为特征。晚白垩世生物年代中的蜥脚类足迹较少,但是鸭嘴龙、暴龙和角龙的足迹增多了。另外,白垩纪足印化石的记录中记载了许多重要的生物地层学信息,如北美白垩纪中期蜥脚类恐龙的消失,以及白垩纪末恐龙的绝灭。越来越多来自东亚的白垩纪足印记录使我们对更精细的地方性白垩纪足印生物年代学有了初步印象。因此,以地方性四足恐龙(包括鸟类)遗迹属的地层分布为基础,可以识别出三个或四个足印生物年代。种类丰富并具有地方性特色的东亚的白垩纪鸟类动物的遗迹群,可能指示白垩纪时东亚存在着一个独特而繁盛的鸟类动物群。以足印化石为基础的这一假说有待进一步的验证。  相似文献   

6.
Two ichnogenus of “large-sized” ornithopods are found from the Lower Cretaceous (Aptian) Kitadani Formation in central Japan. Caririchnium isp. is characterized by the longer pes print than its width with strong mesaxony. Amblydactylus isp. is characterized by the wider pes print than its length with weak mesaxony. In the northern hemisphere, Caririchnium-type footprints are known from the Berriasian–Cenomanian strata, whereas Amblydactylus-type footprints are known from the Barremian–Maastrichtian strata. It is consistent with the temporal and geographic distribution of non-hadrosauroid iguanodontians and basal hadrosauroids. It suggests that footprint length-width ratio and mesaxony are important factors to indicate trackmakers (basal iguanodontian or hadrosauroid). Two “large-sized” ornithopod ichnogenus from the Kitadani Formation is also consistent with two iguanodontians from same site. It indicates a high diversity of the Kitadani ichnofauna and its importance to elucidate the ecosystem of the Kitadani Formation.  相似文献   

7.
Terrestrial Middle Triassic strata occur throughout continental Africa and are particularly well exposed in South Africa, Tanzania, Zambia, and Namibia. The youngest age for all these African deposits is widely accepted as early Middle Triassic (Anisian). Fossils collected recently from the uppermost strata of the upper Omingonde Formation in Namibia highlighted the presence of Chiniquodon, a carnivorous cynodont previously only found in Ladinian-Carnian aged rocks of South America. In addition, work in progress indicates that a large archosaur, originally reported as Erythrosuchus, also discovered from levels close to the top of this unit, is in fact a rauisuchian, a group of archosaurs well known from Ladinian-Carnian beds of southern South America. Here we present the first record of the tuskless dicynodont Stahleckeria potens from the top of the upper Omingonde Formation in central Namibia. This taxon was up until now only known from the Ladinian Dinodontosaurus Assemblage Zone of the Santa Maria Formation in southern Brazil. Thus, compelling evidence for a Ladinian age for the upper levels of the upper Omingonde Formation is provided by two therapsid and one archosaur taxa. The tetrapod fauna of the upper Omingonde Formation partially fills the gap of the well-documented hiatus (Ladinian gap), prevalent throughout the Karoo basins of south and central Africa. The presence of the same therapsid taxa in the Namibian Waterberg Basin and the Paraná Basin of Brazil during Middle Triassic suggests that these basins were biogeographically linked through a series of interconnecting lowlands, with no major ecological, climatic and/or physical barriers.  相似文献   

8.
In the first ever systematic study of trace fossils from the Badhaura Formation, the authors described a nesting burrow, which they ascribed to a stomatopod. The purpose of this paper is two-fold: primarily, to document ichnofauna from (post-glacial marine late Palaeozoic rocks of peninsular India) the Badhaura Formation (Sterlitmakian) representing marine rocks deposited following the Late Palaeozoic glaciation and secondly to contribute to the data on post-glacial ichnofauna from constituent continents of the Gondwanaland. Trace fossils described here are from the Harbans Bed, the topmost lithounit of the Badhaura Formation. The ichnofauna includes Arenicolites tenuis, Beaconites isp., Curvolithus isp., Cylindrichnus concentricus, Didymaulichnus lyelli, Ophiomorpha isp., Palaeophycus tubularis, Planolites beverleyensis, P. montanus, Rosselia chonoides, R. socialis, Skolithos linearis, Taenidium cameronensis, Thalassinoides paradoxicus, Thalassinoides isp. and a flask-shaped brood chamber assigned to a stomatopod crustacean. This mixed assemblage is assigned to distal Skolithos ichnofacies and is suggestive of a period of relatively quiet, shallow water conditions of deposition. The ichnofauna, when viewed in context of peri-gondwanic ichnofaunas, mainly consisting of simple tracks and trails, from late Palaeozoic post-glacial deposits of other Gondwanan continents, is interesting due to dominance of domichnia. Profusion of brood chambers along with Thalassinoides in the Badhaura Formation validates the concept of pre-Mesozoic Thalassinoides being non-decapod in origin and suggestive of adaptive convergence.  相似文献   

9.
New finds reveal that cases of larval caddisflies (indusifauna) are widespread aquatic domichnia in Eurasian non-marine deposits. 16 new ichnospecies are described in 6 ichnogenera from the Wealden Supergroup (Lower Cretaceous) of southeast England, mainly from the Ashdown Formation (Valanginian), but some also from the Weald Clay Group (Hauterivian–Barremian). New ispp. are: Terrindusia valdensis isp. nov., T. anomala isp. nov.; Secrindusia sarahae isp. nov.; Conchindusia dianae isp. nov., C. elderi isp. nov., C. goodmani isp. nov.; Pelindusia duprati isp. nov.; ?Ostracindusia mixta isp. nov.; Folindusia stouti isp. nov., F. bipennis isp. nov., F. ruffordi isp. nov., F. chiasma isp. nov., F. woodhamsi isp. nov., F. boothi isp. nov., F. avancnae isp. nov., and F. whitei isp. nov. Two new forms, a variety and aberration of Conchindusia (an igen. with no modern analogue) are also recognised. The first Chinese isp. (Conchindusia sinensis isp. nov.) is described from the Yixian Formation (Aptian) and 31 morphotypes are listed from eight Lower Cretaceous Chinese formations.The aquatic palaeoenvironment and palaeoecology are discussed. The UK indusifauna is dominated by ConchindusiaFolindusia in contrast to a Terrindusia dominance in China; furthermore, the UK EPT (Ephemeroptera–Plecoptera–Trichoptera) richness is skewed towards Trichoptera. This reflects differences in the fluvio-lagoonal and fluvio-lacustrine settings, respectively.  相似文献   

10.
Paleogene sediments of the inner fold belt, Naga hills, have very well preserved ichnofossils. 16 ichnospecies have been documented among 13 ichnogenera such as Arenicolites isp., Chondrites targionii, Cylindrichnus isp., Diplocraterion parallelum, Gyrochorte isp., Ophiomorpha annulata, O. irregulaire, O. nodosa, O. rudis, Palaeophycus tubularis, Planolites beverleyensis, Scolicia palaeobullia, Skolithos linearis, Trypinites weisei, Thalassinoides horizontalis and Zoophycos isp. The ichnofossil assemblages comprise mostly domichnia and fodinichnia benthos of the Skolithos and Cruziana ichnofacies. A shallow marine nearshore to offshore marine environment with fluctuating energy condition has been envisaged.  相似文献   

11.
The Marwar Supergroup of the Bikaner-Nagaur Basin is composed of sediments deposited from the late Neoproterozoic (Ediacaran) to Upper Cambrian. The Nagaur Sandstone Formation of the Nagaur Group (uppermost division of the Marwar Supergroup) preserves trace fossils significant for establishing Early Cambrian biostratigraphic zones and depositional facies. Fifteen ichnospecies (and eight ichnogenera) identified in the Nagaur Sandstone Formation include “Treptichnus” pedum, Cruziana cf. tenella, Cruziana isp., Diplichnites ispp. A, B, and C, Gyrophyllites isp., Lockeia isp., Merostomichnites isp., Monomorphichnus gregarius isp. nov., Monomorphichnus isp., Planolites isp., Psammichnites isp., Rusophycus bikanerus isp. nov., Rusophycus cf. carbonarius, Rusophycus isp. and radial trace fossils.These trace fossils belong to ethological categories pascichnia, repichnia, cubichnia, and fodinichnia and represent arthropod and worm-like burrowing biota. The assemblage and a regional comparison with contemporaneous trace fossils in the eastern Gondwanan realm suggest that the sequence in the study area belongs to the Cruziana tenella Ichnozone and to Stage 2 (upper part of Terreneuvian), however the Middle Cambrian is not excluded. The trace fossil assemblage belongs to the archetypal Cruziana ichnofacies. Cross bedded sandstone, mud cracks and rainprints in the ichniferous strata of the Nagaur Sandstone Formation indicate deposition in an intertidal sand flat with channels that was exposed episodically.  相似文献   

12.
The Middle Triassic fossil reptile localities near Bayreuth (Bavaria, southern Germany) consist of shallow marine autochthonous glauconitic marls and terebratulid-rich tempestite carbonates of the newly defined Bindlach and Hegnabrunn formations. Single bones and incomplete skeletons of marine reptiles have been recorded in bone beds within in the Illyrian and Fassanian stages. These include the remains of the sauropterygians Neusticosaurus sp., Lariosaurus cf. buzzii [1], Nothosaurus mirabilis [2], Paranothosaurus giganteus [2], Placodus gigas [3], Cyamodus rostratus [4], Cyamodus münsteri [5], Pistosaurus longaevus [6], and ichthyosaursOmphalosaurus sp., and Shastasaurus sp. or proterosaur Tanystrophaeus conspicuus [7]. New skeletal reconstructions are based on the osteological analysis of three dimensionally preserved bones and skeletal remains. The large number of marine endemic placodont macroalgae feeders (P. gigas) in the Bayreuth sites coincides with the presence of invertebrate palaeocommunities that are characteristic of macroalgae meadow paleoenvironments. Most of the reptile species and genera from the Bayreuth localities also occur in beds of similar ages from the Monte San Giorgio (Switzerland/Italy) or Perledo (Italy) lagoonal areas. Ichthyosaurs and pistosaurs were adapted for open marine conditions, and may have migrated from the Panthalassa Oceans into the shallow marine Germanic Basin to reproduce, whereas placodonts and many other sauropterygians seem to have lived permanently in those shallow marine habitats, with large squamates and thecodont or smaller archosaurs in coastal areas.  相似文献   

13.
Abundant Trypanites isp. associated with Gastrochaenolites isp. and Petroxestes pera are evidenced in the first found hardground of the late Albian marine transgressive series, overlying the Continental Intercalaire in the Ouled Nail Range, eastern Saharan Atlas. This hardground is encrusted by oyster shells belonging to Ostrea falco Coquand. The studied hardground developed in warm environment and displays a lateral extension of at least 6 km. It is interpreted as an erosional transgressive surface. The ichnogenus Petroxestes is reported for the first time from Africa. This ichnoassemblage is attributed to the Trypanites ichnofacies.  相似文献   

14.
The Upper Emsian to Frasnian Ia-Ib strata of the Marhouma area (or “km 30” outcrop), exposed in the Ougarta Range (SW Algeria) belong to the Chefar El Ahmar Formation. On the basis of distinct lithological and palaeontological features, this formation is subdivided into three members (Lower Marly Limestones Member, Middle Marly Limestones Member, and Upper Marly Limestones Member). The studied beds show low to moderate diversity of trace fossil assemblage which contains thirteen ichnotaxa: Chondrites intricatus, Chondrites isp., Chondrites cf. targionii, Circulichnis cf. montanus, Cochlichnus isp., Neonereites biserialis, Neonereites multiserialis, Nereites isp., Palaeophycus isp., Planolites isp., Thalassinoides isp., Zoophycos aff. cauda-galli, and Zoophycos isp. A. The two latter ichnotaxa are the most common trace fossils in the assemblage and occur at three different levels showing different bioturbation intensities. The first Zoophycos-bearing level (Zl 1) is characterised by an overall high bioturbation intensity reflecting a very high oxygenation rate and nutrient supply, allowing the development of large and dense Zoophycos specimens. The second Zoophycos-bearing level (Zl 2) has a considerable reduction of bioturbation intensity as compared to the previous level, with an abundance of Chondrites, which is probably due to radical palaeoecological changes that suggests dysoxic and stressful conditions. The third Zoophycos-bearing level (Zl 3) is characterised by an overall moderate bioturbation intensity. The distribution of trace fossils was influenced by lithology, sedimentation rate, energy level (storm events), bottom oxygenation, and nutrient supply. The lithofacies and trace fossils of the Chefar El Ahmar Formation both indicate a depositional environment fluctuating from the lower shoreface to lower offshore zone.  相似文献   

15.
The Early Cretaceous (latest Hauterivian) Faraoni Oceanic Anoxic Event (F-OAE) in the Río Argos section (Caravaca region, southern Spain), the candidate for the Global Boundary Stratotype Section and Point (GSSP) of the Hauterivian–Barremian transition, has been studied. Its ichnological bed-by-bed analysis allows for interpretation of oxygenation changes through the Faraoni Level interval and improvement upon previous characterization of oxygen conditions prior, during and after the F-OAE. The trace fossil assemblage belongs to the Zoophycos ichnofacies and it includes Chondrites intricatus, Chondrites targionii, Halimedides isp., Palaeophycus isp., ?Patagonichnus isp., Planolites isp., Rhizocorallium isp., Taenidium isp., Thalassinoides isp., Trichichnus linearis, Zavitokichnus fusiformis, and Zoophycos isp. Their diversity in particular beds fluctuates. Beds with four to six ichnotaxa reflect a multi-tiered macrobenthic trace maker assemblage living in good oxic and trophic conditions. In one bed below and one bed above the Faraoni Level (both marls without primary lamination), there are only two or three, mostly opportunistic ichnotaxa (Trichichnus, Chondrites, Planolites). They record dysoxic conditions. At the base of the Faraoni Level, one thicker and two thinner beds of marly mudstones (21.2 and 3.5 cm thick, respectively) are characterized by primary lamination. At the top and in the basal part of the thicker bed and in the thinner beds some trace fossils are present. These beds were deposited under anoxic conditions and later colonized by trace makers either from overlying beds deposited under oxic conditions or from the level of a greenish lamina in the lower part of the thicker bed, recording short episodes of dysoxic conditions. The thinner anoxic beds are separated by marls deposited under dysoxic conditions.  相似文献   

16.
《Gondwana Research》2014,26(4):1348-1356
The Early Toarcian Oceanic Anoxic Event (OAE) in the Early Jurassic Period is associated with a major negative carbon isotope excursion (CIE), mass extinction, marine transgression and global warming. The Toarcian OAE is thought to have been caused by flood basalt magmatism, and may have been a trigger for mass extinction. However, these proposed causes of the Toarcian OAE and associated biotic crisis are not adequately resolved by a precise chronology. The duration of the Toarcian OAE has been estimated to be anywhere from ~ 0.12 to ~ 0.9 Myr, most recently 0.74 to 3.26 Myr from U–Pb dating. The CIE associated with the Toarcian OAE has a similar pattern at numerous localities, and there is evidence that the marine carbon isotope variations recorded astronomical forcing signals. Here we estimate a duration of ~ 620 kyr for the main negative CIE, ~ 860 kyr for the polymorphum zone and > 1.58 Myr for the levisoni zone based on 405-kyr astronomical eccentricity tuning of the marine section at Peniche (Portugal). This 405-kyr tuned series provides a ~ 2.5 Myr continuous high-resolution chronology through the Early Toarcian. There are 6, or possibly 7 short eccentricity cycles in the main CIE interval at Peniche. To confirm this astronomically based estimate, we analyzed three other sections at Yorkshire (UK), Dotternhausen (Germany), and Valdorbia (Italy) from marine carbon isotopic series. These four stratigraphic sections from Early Jurassic western Tethys record the Toarcian OAE with ~ 6 prominent carbon isotope cycles in the CIE that span a 600 ± 100 kyr duration. The Peniche 405-kyr tuned series indicates that the pre- and post-CIE intervals experienced strong precession–eccentricity-forced climate change, whereas the CIE interval is marked by dominant obliquity forcing. These dramatic and abrupt changes in astronomical response in the carbon isotopes point to fundamental shifting in the Early Toarcian paleoclimate system that was directly linked to the global carbon cycle.  相似文献   

17.
The 18 ichnospecies of the vertically oriented, helical marine to marginal marine trace fossil Gyrolithes Saporta, 1884 known so far are revised. A new ichnospecies, G. lorcaensis isp. n. (Miocene, SE Spain), is introduced, and Conispiron Vialov, 1969 and Spirocircus Mikulá? and Pek, 1994 are included in Gyrolithes. The ichnotaxobases of the Gyrolithes ichnospecies are based on the character of the burrow margin (presence or absence of a wall or ornamentation) and/or on morphometric parameters (burrow width versus radius of the whorls). As a result of synonymization, 13 ichnospecies are recommended. The morphometric parameters of Gyrolithes show two distinct lineages (large ratio of whorl radius to burrow width and medium ratio of whorl radius to burrow width; 10–12 and 1–1.6, respectively). The third lineage (small ratio of whorl radius to burrow width; up to 1) is less distinct.  相似文献   

18.
The end-Permian mass extinction is now robustly dated at 252.6 ± 0.2 Ma (U–Pb) and the Permian–Triassic (P–T) GSSP level is dated by interpolation at 252.5 Ma. An isotopic geochronological timescale for the Late Permian–Early Triassic, based on recent accurate high-precision U–Pb single zircon dating of volcanic ashes, together with calibrated conodont zonation schemes, is presented. The duration of the Early Triassic (Induan + Olenekian stages) is estimated at only 5.5 million years. The duration of the Induan Stage (Griesbachian + Dienerian sub-stages) is estimated at ca. one million years and the early Olenekian (Smithian sub-stage) at 0.7 million years duration. Considering this timescale, the “delayed” recovery following the end-Permian mass extinction may not in fact have been particularly protracted, in the light of the severity of the extinction. Conodonts evolved rapidly in the first 1 million years following the mass extinction leading to recognition of high-resolution conodont zones. Continued episodic global environmental and climatic stress following the extinction is recognized by multiple carbon isotope excursions, further faunal turnover and peculiar sedimentary and biotic facies (e.g. microbialites). The end-Permian mass extinction is interpreted to be synchronous globally and between marine and non-marine environments. The nature of the double-phased Late Permian extinction (at the Guadalupian–Lopingian boundary and the P–T boundary), linked to large igneous provinces, suggests a primary role for superplume activity that involved geomagnetic polarity change and massive volcanism.  相似文献   

19.
上二叠统记录着地质历史上最大的生物灭绝事件和最深刻的环境变化。长期以来由于世界各主要沉积区晚二叠世地层的同期性未能确定 ,对这一事件及其过程难以全面了解。随着乐平统成为全球上二叠统的标准 ,我们选用若干生物地层、磁性地层和层序地层等的联接点 ,提出乐平世地层世界对比的试用方案  相似文献   

20.
The Almería-Níjar Basin is a Neogene, intermontane depression marginal to the Mediterranean in southern Spain in the vicinity of El Argamasón, Andalucia. The Pliocene Cuevas Formation rests unconformably on the Upper Messinian rock succession in the Carboneras Fault Zone. The Cuevas Formation is a coarse-grained, bioclastic-rich, calcarenite to calcirudite shoreface deposit. Oysters, namely Saccostrea cucullata (Born), are locally common and preserve a moderate diversity of borings: Caulostrepsis taeniola Clarke; Entobia isp.; Gastrochaenolites isp. aff. G. lapidicus Kelly and Bromley; Maeandropolydora isp. cf. M. sulcans Voigt; Oichnus paraboloides Bromley; and Talpina isp. aff. T. hirsuta Voigt. All represent domiciles except for the predatory O. paraboloides trace. This suite of ichnotaxa is assigned to the Entobia ichnofacies sensu Bromley and Asgaard; they are comparable, particularly, with the Boulder Assemblage of the Pliocene of Rhodes, Greece. Physical disturbance is an important parameter in favouring this pattern of infestation, whether the bored clasts are boulders or oyster valves.  相似文献   

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