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1.
Pollen in Quaternary deposits from the subtropical Hanjiang Delta records three major phases in the local vegetation and climate history during the last 55,000 yr: (1) a prevalent cool-to-temperate and humid climate at ca. 24,000 14C yr B.P. is indicated by abundant pollen of temperate trees including conifers; (2) between 20,000 and 15,000 14C yr B.P., a cold, dry environment was associated with low sea level during the last glaciation, leading to subaerial exposure, weathering, and interruption of sedimentation, as well as departure from the region of Dacrydium and Sonneratia; (3) a short-term expansion of grassland at ca. 10,300 14C yr B.P. reduced the predominant Lauraceae-Fagaceae evergreen forest, possibly corresponding to the Younger Dryas cooling. The combined data indicate a maximum sea-level rise in the mid-Holocene (7500–4000 14C yr B.P.) and a marine influence in the late Pleistocene at 45,000–20,000 14C yr B.P. The Holocene warming, however, did not bring back moisture-sensitive taxa, indicating high seasonal aridity probably caused by renewed monsoon conditions.  相似文献   

2.
Seismic stratigraphy, sedimentary facies, pollen stratigraphy, diatom-inferred salinity, stable isotope (δ18O and δ13C), and chemical composition (Sr/Ca and Mg/Ca) of authigenic carbonates from Moon Lake cores provide a congruent Holocene record of effective moisture for the eastern Northern Great Plains. Between 11,700 and 950014C yr B.P., the climate was cool and moist. A gradual decrease in effective moisture occurred between 9500 and 710014C yr B.P. A change at about 710014C yr B.P. inaugurated the most arid period during the Holocene. Between 7100 and 400014C yr B.P., three arid phases occurred at 6600–620014C yr B.P., 5400–520014C yr B.P., and 4800–460014C yr B.P. Effective moisture generally increased after 400014C yr B.P., but periods of low effective moisture occurred between 2900–280014C yr B.P. and 1200–80014C yr B.P. The data also suggest high climatic variability during the last few centuries. Despite the overall congruence, the biological (diatom), sedimentological, isotopic, and chemical proxies were occassionally out of phase. At these times the evaporative process was not the only control of lake-water chemical and isotopic composition.  相似文献   

3.
Holocene environments have been reconstructed by multiproxy studies of an 850-cm-long core from Rio Curuá dating to >8000 14C yr B.P. The low-energy river lies in the eastern Amazon rain forest in the Caxiuanã National Forest Reserve, 350 km west of Belém in northern Brazil. Sedimentological, mineralogical, and geochemical dates demonstrate that the deposits correspond to two different environments, sediments of an active river before 8000 14C yr B.P. and later a passive river system. The pollen analytical results indicate four different local and regional Holocene paleoenvironmental periods: (1) a transition to a passive fluvial system and a well-drained terra firme (unflooded upland) Amazon rain forest with very limited development of inundated forests (várzea and igapó) (>7990–7030 14C yr B.P.); (2) a sluggish river with a local Mauritia palm-swamp and similar regional vegetation, as before (7030–5970 14C yr B.P.); (3) a passive river, forming shallow lake conditions and with still-abundant terra firme forest in the study region (5970–2470 14C yr B.P.); and (4) a blocked river with high water levels and marked increase of inundated forests during the last 2470 14C yr B.P. Increased charcoal during this last period suggests the first strong presence of humans in this region. The Atlantic sea level rise was probably the major factor in paleoenvironmental changes, but high water stands might also be due to greater annual rainfall during the late Holocene.  相似文献   

4.
Benthonic foraminiferal faunal and isotopic data from Champlain Sea sediments (approximately 12,500 to 10,000 yr B.P. in age) in two piston cores from Lake Champlain provide a detailed, apparently continuous record of the evolution of the Champlain Sea. Cassidulina reniforme and Islandiella helenae are the dominant benthonic foraminifera during the initial phase of the Champlain Sea, and are replaced by Elphidium excavatum forma clavatum and Protelphidium orbiculare as the dominant species during the remainder of the sea's history. The oxygen-isotopic data show a gradual decrease in δ18O between approximately 12,500 and 10,900 yr B.P., followed by a more rapid decrease during the interval 10,900 to 10,000 yr B.P. The δ13C data have a similar trend as δ18O, with generally decreasing values up the section. The isotopic and faunal data suggest that nearly marine conditions were present in the initial plase of the Champlain Sea, followed by gradually decreasing salinities and increasing temperatures as the sea evolved. The beginning of the rapid isotopic decrease at approximately 10,900 yr B.P. marks the onset of the largest environmental change in the history of the Champlain Sea, probably reflecting a major pulse of meltwater from the Laurentide Ice Sheet.  相似文献   

5.
A sediment core from Smorodinovoye Lake (SML), northeastern Siberia (area to the east of the Verkhoyansk Range) spanning the last 24,000 14C yr indicates that vegetational and climatic changes in the upper Indigirka basin resemble those in eastern Siberia (Lena basin and westward). For example, maximum postglacial summer temperatures at SML probably occurred 6000–4000 14C yr B.P., an age more in accordance with eastern than northeastern records. Larix arrived near the lake by 9600 14C yr B.P., approximately when forests expanded in the east but ca. 1500 14C yr later than forests were established in the neighboring upper Kolyma basin. Paleobotanical data further suggest that Larix possibly migrated southward from populations in the arctic lowlands of eastern Siberia and did not originate from interior refugia of the upper Kolyma basin. Although a Younger Dryas cooling has been noted in eastern Siberia, SML provides the first evidence from the northeast for a similar climatic reversal. Climatic variations seemingly have persisted between the Indigirka and Kolyma basins over at least the last 11,000 14C yr, despite the proximity of the two drainages and the occurrence of major changes in boundary conditions (e.g., seasonal insolation, sea levels) that have influenced other regional climatic patterns.  相似文献   

6.
The organic deposits derived from the mangrove swamps form reliable stratigraphic markers within the Late Quaternary sequence of Kerala–Konkan Basin. Three generations of such deposits have been identified. The older one is dated to around 43,000–40,000 14C yr B.P., with a few dates beyond the range of radiocarbon. The younger ones date from the Middle Holocene to latest Pleistocene (10,760–4540 14C yr B.P.) and the Late Holocene (<4000 14C yr B.P.). Pollen analyses confirm that the deposits are mostly derived from the mangrove vegetation. Peat accumulation during the period 40,000–28,000 14C yr B.P. can be correlated with the excess rainfall, 40–100% greater than modern values, of the Asian summer monsoon. The low occurrence of mangrove between 22,000 and 18,000 14C yr B.P. can be attributed to the prevailing aridity and/or reduced precipitation associated worldwide with Last Glacial Maximum, because exposure surfaces and ferruginous layers are commonly found in intervals representing this period. The high rainfall of 11,000–4000 14C yr B.P. is found to be the most significant as the mangrove reached an optimum growth around 11,000 14C yr B.P. but with periods of punctuated weaker monsoons. From the present and previous studies, it has been observed that after about 5000 or 4000 14C yr B.P., the monsoons became gradually reduced leading to drying up of many of the marginal marine mangrove ecosystems. A case study of Hadi profile provided an insight to the relevance of magnetic susceptibility (χ) to record the ecological shift in Late Holocene.  相似文献   

7.
New cross sections and dates from along the Pomme de Terre River clarify the complex local history of valley development and floodplain sedimentation. The observed history begins with a series of ancient bedrock strath terraces that record past bedrock valley positions at 15.5 to more than 58 m above the modern bedrock floor. Each strath is capped by 1–2 m of channel gravel and sand permeated by red clay. Sometime previous to ca. 140,000 yr B.P., a much lower bedrock valley only about 5–6 m above the modern level was excavated. By 140,000 yr B.P., accumulation of red and gray mottled silty clay had commenced, and had reached to 8.5 m above the modern floodplain before 48,900 ± 900 14C yr B.P. Sometime between ca. 49,000 and 45,000 14C yr B.P., erosion caused abandonment of an oxbow meander, and lowered the bedrock valley to about its present depth. Younger yellowish-red and gray mottled silty clay alluvium then began accumulating. This mid-Wisconsinan fill reached to 2.5 m above the modern floodplain sometime before 31,800 ± 1340 14C yr B.P., at which time another erosional phase was in progress. A late Wisconsinan olive clay accumulated between 27,480 ± 1950 and ca. 23,000 14C yr B.P., followed by approximate stability until 13,550 ± 400 14C yr B.P. After stability, an erosional episode began, but by 10,200 ± 330 14C yr B.P., deposition of a distinctive brown clayey silt was underway. This early Holocene fill reached to about the same level as the mid-Wisconsinan fill by 8100 ± 140 14C yr B.P. Erosion occurred between this date and 7490 ± 170 14C yr B.P., but the former floodplain level was rapidly reattained, and was apparently stable until ca. 5000 14C yr B.P. Finally, erosional unconformities and 17 dates from the brown clayey silt, and from younger grayish-brown silty sand underlying the modern floodplain, record subsequent episodes of floodplain erosion at ca. 5000, 2900, 1500 and 350 14C yr B.P. The timing of Pomme de Terre floodplain sedimentary regimes, characterized by net aggradation, erosion, or stability, may have been controlled by climate. In particular, both periods of stability appear to have been coeval to times of strongly zonal upper atmospheric circulation. Intensified zonal circulation would have resulted in less frequent large floods and an increased dominance by floods of small to moderate size. In contrast, there are no obvious parallels to be drawn between this local alluvial history and sea level or glacial outwash induced baselevel changes.  相似文献   

8.
Diatom assemblages and organic carbon records from two sediment cores located within an estuarian bay of the inner Kara Sea trace changes in Yenisei River runoff and postglacial depositional environments. Paleosalinity and sea-ice reconstructions are based on modern relationships of local diatom assemblages and summer surface-water salinity. Approximately 15,500 cal yr B.P., rivers and bogs characterized the study area. When sea level reached the 38- to 40-m paleo-isobath approximately 9300 cal yr B.P., the coring site was flooded. From 9300–9100 cal yr B.P., estuarine conditions occurred proximal to the depocenter of fluvially derived material, and salinity was <7–8. Paleosalinity increased to 11–13 by 7500 cal yr B.P., following postglacial sea-level rise and the southward shift of the Siberian coast. Sharp decreases in diatom accumulation rates, total sediment, and organic carbon also occurred, suggesting the presence of brackish conditions and greater distance between the coast and study site. Maximum paleosalinity (up to 13) was recorded between 7500 and 6000 cal yr B.P., which was likely caused by the enhanced penetration of Atlantic waters to the Kara Sea. Stepwise decreases to modern salinity levels happened over the last 6000 cal yr.  相似文献   

9.
Lipid extracts from a 61.7-cm-long subtropical stalagmite in southern China, spanning the period of ca. 10,000–21,000 yr ago as constrained by U–Th dating, were analyzed using gas chromatography–mass spectrometry. The higher plants and microorganisms in the overlying soils contribute a proportion of n-alkanes identified in the stalagmite. The occurrence of LMW (lower molecular weight) n-alkanols and n-alkan-2-ones in the stalagmite was mainly related to the soil microorganisms. We suggest that HMW (higher molecular weight) n-alkanols and n-alkan-2-ones identified in the stalagmite originate from soil organics and reflect input from contemporary vegetation. Shifts in the ratio of LMW to HMW n-alkanols or n-alkan-2-ones indicative of the variation of soil ecosystems (e.g., microbial degradation of organic matter and/or the relative abundance of soil microorganisms to higher plants) are comparable with the subtropical alkenone-SST (sea surface temperature) record of the same period. The similar trends seen in the δ13C data and the lipid parameters in this stalagmite imply that the overlying soil ecosystem response to climate might be responsible for the variation of δ13C values.  相似文献   

10.
Archaeological investigations in Camels Back Cave, western Utah, recovered a series of small-mammal bone assemblages from stratified deposits dating between ca. 12,000 and 500 14C yr B.P. The cave's early Holocene fauna includes a number of species adapted to montane or mesic habitats containing grasses and/or sagebrush (e.g., Lepus townsendii, Marmota flaviventris, Reithrodontomys megalotis, and Brachylagus idahoensis) which suggest that the region was relatively cool and moist until after 8800 14C yr B.P. Between ca. 8600 and 8100 14C yr B.P. these mammals became locally extinct, taxonomic diversity declined, and there was an increase in species well-adapted to xeric, low-elevation habitats, including ground squirrels, Lepus californicus and Neotoma lepida. The early small-mammal record from Camels Back Cave is similar to the 11,300–6000 14C yr B.P. mammalian sequence from Homestead Cave, northwestern Utah, and provides corroborative data on Bonneville Basin paleoenvironments and mammalian responses to middle Holocene desertification.  相似文献   

11.
High-resolution paleomonsoon proxy records from peat and eolian sand–paleosol sequences at the desert–loess transition zone in China denote a rapid oscillation from cold–dry conditions (11,200–10,60014C yr B.P.) to cool–humid conditions (10,600–10,20014C yr B.P.), followed by a return to cold–dry climate (10,200–10,00014C yr B.P.). Variations in precipitation proxies suggest that significant climatic variability occurred in monsoonal eastern Asia during the Younger Dryas interval. Late-glacial climate in the Chinese desert–loess belt that lies downwind from Europe was strongly influenced by cold air from high latitudes and from the North Atlantic via the westerlies. The inferred precipitation variations were likely caused by variations in the strength of the Siberian high, which influenced the pressure gradient between land and ocean and therefore influenced the position of the East Asian monsoon front.  相似文献   

12.
Basal sediments of Lake Torfadalsvatn, northern Iceland, record changes in terrestrial and limnic environments in the period 11,300-9000 14C yr B.P. These changes were probably forced by climate and connected with displacements of the marine polar front and sea-ice margin. Pollen, spores, green algae (Pediastrum), saturation isothermal remanent magnetization, and carbon content of the basal sediments provide the first detailed biostratigraphic record of the last glacial-interglacial transition in Iceland. During the first pioneer phase, beginning at ca. 11,300 14C yr B.P., grasses and fell-field herbs became established, and lake productivity was very low. At ca. 10,900 14C yr B.P., climatic and soil conditions became favorable for shrubs and dwarf shrubs. This change, together with increased limnic productivity, clearly indicates long seasons without ice-cover in the sea immediately north of Iceland. A return to a colder climate (Younger Dryas), probably in connection with a southward displacement of the marine polar front, occurred by 10,600 14C yr B.P. Shrub and dwarf-shrub vegetation disappeared, and limnic productivity diminished. A second pioneer vegetation phase, dominated by Oxyria/Rumex and grasses, was initiated by a change to longer seasons without sea ice at ca. 9900 14C yr B.P. This warming is also evident as a contemporaneous increase in lake productivity. After ca. 9400 14C yr B.P. the reestablishment of dwarf-shrub heaths and very high limnic productivity indicate further warming.  相似文献   

13.
In the southern Argentine Andes, ten advances of valley glaciers were used to reconstruct the late-glacial and Holocene glacier history. The accumulation areas of these glaciers lie in the Precordillera and are thus independent of fluctuations of the South Patagonian Icefield. Like the Viedma outlet glacier, the valley glaciers advanced three times during late-glacial time (14,000–10,000 yr B.P.). The youngest advance correlates with the Younger Dryas Stade, based on two minimum AMS14C dates of 9588 and 9482 yr B.P. The second oldest advance occurred before 11,800 yr B.P. During the first half of the Holocene, (ca. 10,000–5000 yr B.P.), advances culminated about 8500, 8000–7500, and 5800–5500 yr B.P. During the second half of the Holocene, advances occurred between ca. 4500 and 4200 yr B.P., as well as between 3600 and 3300 yr B.P. In the Río Cóndor valley three subsequent advances have been identified.  相似文献   

14.
A new packrat midden chronology from Playas Valley, southwestern New Mexico, is the first installment of an ongoing effort to reconstruct paleovegetation and paleoclimate in the U.S.A.–Mexico Borderlands. Playas Valley and neighboring basins supported pluvial lakes during full and/or late glacial times. Plant macrofossil and pollen assemblages from nine middens in the Playas Valley allow comparisons of two time intervals: 16,000–10,000 and 4000–0 14C yr B.P. Vegetation along pluvial lake margins consisted of open pinyon–juniper communities dominated by Pinus edulis, Juniperus scopulorum, Juniperus cf. coahuilensis, and a rich understory of C4 annuals and grasses. This summer-flowering understory is also characteristic of modern desert grassland in the Borderlands and indicates at least moderate summer precipitation. P. edulis and J. scopulorum disappeared or were rare in the midden record by 10,670 14C yr B.P. The late Holocene is marked by the arrival of Chihuahuan desert scrub elements and few departures as the vegetation gradually became modern in character. Larrea tridentata appears as late as 2190 14C yr B.P. based on macrofossils, but may have been present as early as 4095 14C yr B.P. based on pollen. Fouquieria splendens, one of the dominant desert species present at the site today, makes its first appearance only in the last millennium. The midden pollen assemblages are difficult to interpret; they lack modern analogs in surface pollen assemblages from stock tanks at different elevations in the Borderlands.  相似文献   

15.
The landscapes of northern New England and adjacent areas of Canada changed greatly between 14,000 and 9000 yr B.P.: deglaciation occurred, sea levels and shorelines shifted, and a vegetational transition from tundra to closed forest took place. Data from 51 14C-dated sites from a range of elevations were used to map ice and sea positions, physiognomic vegetational zones, and the spread of individual tree taxa in the region. A continuum of tundra-woodland-forest passed northeastward and northward without major hesitation or reversal. An increased rate of progression from 11,000 to 10,000 yr B.P. suggests a more rapid warming than in the prior 2000–3000 yr. Elevational gradients controlled the patterns of deglaciation and vegetational change. The earliest spread of tree taxa was via the lowlands of southern Vermont and New Hampshire, and along a coastal corridor in Maine. Only after 12,000 yr B.P. did the taxa spread northward through the rest of the area. Different tree species entered the southern part of the area at different times and continued their spread at different rates. The approximate order of arrival follows: poplars (13,000–12,000 yr B.P. in the south), spruces, paper birch, and jack pine, followed by balsam fir and larch, and possibly ironwood, ash, and elm, and somewhat later by oak, maple, white pine, and finally hemlock (10,000–9000 yr B.P. in the south).  相似文献   

16.
Large glaciers descended western valleys of the Olympic Mountains six times during the last (Wisconsin) glaciation, terminating in the Pacific coastal lowlands. The glaciers constructed extensive landforms and thick stratigraphic sequences, which commonly contain wood and other organic detritus. The organic material, coupled with stratigraphic data, provides a detailed radiocarbon chronology of late Pleistocene ice-margin fluctuations. The early Wisconsin Lyman Rapids advance, which terminated prior to ca. 54,000 14C yr B.P., represented the most extensive ice cover. Subsequent glacier expansions included the Hoh Oxbow 1 advance, which commenced between ca. 42,000 and 35,000 14C yr B.P.; the Hoh Oxbow 2 advance, ca. 30,800 to 26,300 14C yr B.P.; the Hoh Oxbow 3 advance, ca. 22,000–19,300 14C yr B.P.; the Twin Creeks 1 advance, 19,100–18,300 14C yr B.P.; and the subsequent, undated Twin Creeks 2 advance. The Hoh Oxbow 2 advance represents the greatest ice extent of the last 50,000 yr, with the glacier extending 22 km further downvalley than during the Twin Creeks 1 advance, which is correlative with the global last glacial maximum. Local pollen data indicate intensified summer cooling during successive stadial events. Because ice extent was diminished during colder stadial events, precipitation—not summer temperature—influenced the magnitude of glaciation most strongly. Regional aridity, independently documented by extensive pollen evidence, limited ice extent during the last glacial maximum. The timing of glacier advances suggests causal links with North Atlantic Bond cycles and Heinrich events.  相似文献   

17.
Despite growing evidence for environmental oscillations during the last glacial–interglacial transition from high latitude, terrestrial sites of the North Pacific rim, oxygen-isotopic records of these oscillations remain sparse. The lack of data is due partially to the paucity of lakes that contain carbonate sediment suitable for oxygen-isotopic analysis. We report here the first record of oxygen-isotopic composition in diatom silica (δ18OSi) from a lake in that region. δ18OSi increases gradually from 19.0 to 23.5‰ between 12,340 and 11,000 14C yr B.P., reflecting marked climatic warming at the end of the last glaciation. Around 11,000 14C yr B.P., δ18OSi decreases by 1.7‰, suggesting a temperature decrease of 3.5–8.9 °C at the onset of the Younger Dryas (YD) in southwestern Alaska. Climatic recovery began ca. 10,740 14C yr B.P., as inferred from the increase of δ18OSi to a maximum of 23.9‰ near the end of the YD. Our data reveal that a YD climatic reversal in southwestern coastal areas of Alaska occurred, but the YD climate did not return to full-glacial conditions.  相似文献   

18.
Paleolake sediment, constrained by tephrochronology, from Onepoto basin volcanic crater in Auckland, Northern New Zealand (36° 48′S), provides one of the few uninterrupted records of paleovegetation for marine oxygen isotope stages (MIS) 4 and 3 (76,000–26,000 yr B.P.) in the region. This period was characterized by cool temperate conifer-hardwood forest that lacked some of the warmer taxa typical of the Holocene. The period 64,400–60,500 yr B.P. was marked by opening of forest canopy and expansion of small trees and shrubs, and correlates to the thermal minima of MIS 4. However, the landscape was never as open as the forest-shrubland mosaic of the MIS 2. The beginning of MIS 3 (60,500–50,500 yr B.P.) was marked by the dramatic expansion and then decline of conifer-hardwood forest dominated by Dacrydium cupressinum, a species that prefers wetter conditions. This forest was succeeded by the typically montane Nothofagus at 50,500 yr B.P., corresponding to a thermal decline. Thus, MIS 3 began with an abrupt change to moist cool conditions that lasted about 5000 yr, followed by gradual cooling and dryer conditions. This supports some interpretations from other parts of the southwest Pacific region, that MIS 3 was a period of increased precipitation. The widespread and stratigraphically important Rotoehu tephra, erupted from Okataina Volcanic Centre, has been variously dated at 45,000–65,000 yr B.P. At Onepoto, sedimentation rate and paleovegetation reconstruction imply an age of c. 44,300 yr B.P. The tephra provides a correlation horizon in the marine and terrestrial realms during a period (MIS 3) difficult to date by radiometric methods.  相似文献   

19.
Coupled measurements of δ18O and accelerator mass spectrometry (AMS) 14C in a particular species of planktonic foraminifera may be used to calculate sea-level estimates for the last deglaciation. Of critical importance for this type of study is a knowledge of the seasonality of foraminiferal growth, which can be provided by δ18O measurements of modern shells (core tops, plankton tows). Isotopic (δ18O, AMS-14C dating) and faunal records (transfer function sea surface temperature) were obtained from two cores in the North Atlantic at about 37°N. The locations were chosen to obtain high sedimentation rate records removed from the major ice-melt discharge areas of the last deglaciation. Based upon Globigerina bulloides data, four δ18O-based sea-level estimates were calculated: −67 ± 7 m at 12,200 yr B.P. and −24 ± 8 m at about 8200 yr B.P. for core SU 81-18; −83 ± 10 m at 12,200 yr B.P. and −13 ± 11 m at about 8500 yr B.P. for core SU 81-14. Using a second working hypothesis concerning the seasonability of G. bulloides growth, it is suggested that the sea-level rose by about 40 m during the millennium which followed 14,500 yr B.P.  相似文献   

20.
A high-resolution pollen record from a 5-m-long sediment core from the closed-lake basin Laguna Piusbi in the southern Colombian Pacific lowlands of Chocó, dated by 11 AMS14C dates that range from ca. 7670 to 22014C yr B.P., represents the first Holocene record from the Chocó rain forest area. The interval between 7600 and 610014C yr B.P. (500–265 cm), composed of sandy clays that accumulated during the initial phase of lake formation, is almost barren of pollen. Fungal spores and the presence of herbs and disturbance taxa suggest the basin was at least temporarily inundated and the vegetation was open. The closed lake basin might have formed during an earthquake, probably about 440014C yr B.P. From the interval of about 600014C yr B.P. onwards, 200 different pollen and spore types were identified in the core, illustrating a diverse floristic composition of the local rain forest. Main taxa are Moraceae/Urticaceae,Cecropia,Melastomataceae/Combretaceae,Acalypha, Alchornea,Fabaceae,Mimosa, Piper, Protium, Sloanea, Euterpe/Geonoma, Socratea,andWettinia.Little change took place during that time interval. Compared to the pollen records from the rain forests of the Colombian Amazon basin and adjacent savannas, the Chocó rain forest ecosystem has been very stable during the late Holocene. Paleoindians probably lived there at least since 346014C yr B.P. Evidence of agricultural activity, shown by cultivation ofZea maissurrounding the lake, spans the last 1710 yr. Past and present very moist climate and little human influence are important factors in maintaining the stable ecosystem and high biodiversity of the Chocó rain forest.  相似文献   

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