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1.
Dissolved and particulate Mn concentrations were investigated on a seasonal scale in surface waters of the NW German Wadden Sea (Spiekeroog Island) in 2002 and 2003. As the Wadden Sea forms the transition zone between the terrestrial and marine realms, Mn was analysed in coastal freshwater tributaries and in the adjoining German Bight as well. Additionally, sediments and porewaters of the tidal flat sediments were analysed for Mn partitioning and microbial activity.Dissolved Mn concentrations show strong tidal and seasonal variation with elevated concentrations during summer at low tide. Summer values in the Wadden Sea (av. 0.7 μM) are distinctly higher than in the central areas of the German Bight (av. 0.02 μM), suggesting a possible impact of the Wadden Sea environment on the Mn budget of the North Sea. Seasonality is also observed for particulate Mn in the Wadden Sea (winter av. 800 mg kg− 1; summer av. 1360 mg kg− 1). Although particles are relatively Mn-poor during winter, the high SPM load during this season causes elevated excess concentrations of particulate Mn, which in part exceed those of the dissolved phase. Therefore, winter values cannot be ignored in balance calculations for the Wadden Sea system.Porewater Mn concentrations differ depending on sediment type and season. Maximum concentrations are found in surface sediments at a mixed flat site (190 μM) during summer, while winter values are distinctly lower. This indicates that enhanced microbial activity owing to higher temperature during summer leads to increased reduction of Mn-oxides in surface sediments and enhances the corresponding diffusive and advective Mn flux across the sediment-water interface. Draining of Mn-rich porewaters from sediments is also documented by analyses of tidal creek waters, which are highly enriched in Mn during summer.Furthermore, an important Mn source is freshwater discharged into the Wadden Sea via a flood-gate. The concentration of dissolved Mn in freshwater was highly variable during the sampling campaigns in 2002 and 2003, averaging 4 μM. In contrast, particulate Mn displayed a seasonal behaviour with increasing contents during summer. On the basis of salinity variations in the Wadden Sea, the total amount of Mn contributed to the Wadden Sea via freshwater was estimated. This balance shows the importance of the freshwater environment for the Mn inventory of the Wadden Sea. During winter the total Mn inventory of the Wadden Sea water column may be explained almost completely by freshwater discharge, whereas in summer the porewater system forms the dominating source.  相似文献   

2.
《Journal of Sea Research》1999,41(3):233-244
Harbour seals in the Wadden Sea (The Netherlands, Germany and Denmark) have been monitored by aerial surveys since the 1980s. Annual maximum figures reported in the various regions (38 separate tidal basins) and the number of pups counted were used to estimate vital population parameters and to quantify the net migration of harbour seals within the Wadden Sea before (1980–1987) and after an epizootic (1990–1994). The total pre-whelping population increased at a mean annual rate of 9 and 14%, respectively, and varied considerably among the various regions of the Wadden Sea. The overall mean annual survival was found to increase from 0.88 during the pre-epizootic period to 0.93 during the post-epizootic period. The distribution of pups and of older seals over the 38 separate sub-areas was highly uneven. Based on between-year population changes and the number of pups reported, net dispersal fluxes among the four regions of the Wadden Sea were estimated for the two time intervals. During the pre-epizootic period, only Schleswig-Holstein, Germany, produced a surplus of animals which counter-balanced the low recruitment of the other regions. During the post-epizootic period, the net dispersal fluxes were at a much lower level. Nevertheless, in The Netherlands, a continuous influx of seals was required because the recruitment was insufficient to explain the observed numerical increase. At the level of the 38 separate tidal basins, the actual pattern of dispersal showed that the majority of sub-areas were not self-supporting with respect to the local pup production and thus dependent on the influx of animals. More than 65% of all immigrants originated from only 7 sub-areas, which are considered `key areas' of vital importance for the Wadden Sea harbour seal population.  相似文献   

3.
The Wadden Sea fronting the North Sea coast of Germany is a unique, virtually untouched, natural landscape. It is a very important dynamic site for special plants and animals dependent on this habitat. Therefore the Lower Saxony Wadden Sea and the islands have the status of a wetland of international importance, agreed by the Ramsar-Convention, in 1976. It has also been declared as an area of the European Bird Directive and a region with priority for nature. As a consequence, the total Wadden Sea (without the Dollart) of 240 000 ha was constituted as the national park ‘Niedersächsisches Wattenmeer’ on 1 January 1986. It is sectioned into three zones with different states of preservation and human use. The national park administration in Wilhelmshaven protects and tries to develop the park. Therefore it is necessary to stop or reduce all kinds of human activities, for instance farming, hunting, fishing, coast protection and tourism.  相似文献   

4.
Daily observations of the salinity of the Marsdiep tidal inlet, which connects the Dutch western Wadden Sea with the North Sea, already started over 140 years ago, in 1860. Since the year 2000 the sampling frequency has increased because of the use of electronic sensors. Analysis of these salinity data have revealed variations on time scales from tidal (~ 12 hour), seasonal, inter-annual, and multi-decadal, to centennial. The contributions of the salinity variations in the Marsdiep for these different spectral bands or time scales are all of the order of a standard deviation of 0.5 to 1. The centennial variation, which can be expressed as a 140 year long salinity trend, is related to engineering works on the rivers Rhine and IJssel, which already started in the early 18-th century, and more than doubled the magnitude of the freshwater content of the western Wadden Sea since then. In contrast with this anthropogenic salinity trend, the climatic variability of the precipitation over western Europe, and the connected changes in the Rhine discharge, are mainly responsible for the inter-annual variations in the salinity and/or freshwater content of the western Wadden Sea. Since variations in salinity and freshwater content also reflect variations in the terrigeneous and river influence on the Wadden ecosystem, e.g. via the nutrient content, it can be expected that the ecology of the Wadden Sea also experienced changes on centennial time scales.  相似文献   

5.
The Wadden Sea is a shallow coastal region, with a large area of sedimentary tidal flats that extends from The Netherlands to Denmark and has been declared a site of international importance in the Dutch and German parts (Ramsar status and UNESCO World Heritage Site). Benthic macrofauna are central to the ecosystem functioning of this area, as they recycle nutrients, decompose organic matter and are an important food source for many secondary consumers, like fish and waterbirds. Due to the environmental gradients characteristic of estuarine systems, it is expected that changes in assemblage composition will be observed across the physical and environmental gradients of the Wadden Sea. First, we explored the spatial variation in assemblage composition of benthic macrofauna across the intertidal part of the Dutch Wadden Sea using 3 years of biomass data. Then, we identified the relative importance of six environmental variables for explaining and predicting changes in assemblage composition across the intertidal areas of the Wadden Sea using generalised dissimilarity modelling (GDM). In accordance with the environmental gradients across this system, the biomass contributed by a few common species differed from west to east and were distinct in the Dollard. In the west, bivalves Mya arenaria, Cerastoderma edule and Ensis directus contributed a relatively large and equal share of the total biomass, whereas C. edule contributed the sole largest share of the total biomass towards the east. The polychaete Alitta succinea became a large share of the total biomass in the upper Ems and in the Dollard estuary, but contributed little elsewhere. Similar to the observed differences in species composition, the spatial patterns in assemblage composition, as predicted by the GDM models, identified the Dollard as distinct and that the prevalence of assemblage types in the west differed to the east. Median grain size, followed by microphytobenthic biomass, and exposure time were the most important variables describing differences in assemblage composition. That the Wadden Sea forms a heterogeneous landscape where assemblage composition varies across multiple gradients has repercussions for management and monitoring.  相似文献   

6.
The new diatom species Mediopyxis helysia was described to science from clones found in 2003 in the North Sea, northern Wadden Sea, and the Gulf of Maine. Seven years after its first occurrence, we observed Mediopyxis to contribute up to almost 50% of the biovolume of the diatoms during a diatom spring bloom in the western Wadden Sea. Grazing experiments based on the dilution technique could not detect any microzooplankton grazing impact on the bloom community. Mediopyxis is now also well established in the western Wadden Sea and does have the potential to become a dominant species. The reasons for its success remain largely unresolved but avoidance of being grazed might be one factor. Future research on this new species is needed to understand the success and forecast the ecological footprint of this large diatom species arriving in the western European Seas.  相似文献   

7.
《Journal of Sea Research》2009,61(4):227-234
Daily observations of the sea surface temperature in the Marsdiep tidal inlet, which connects the shallow Dutch western Wadden Sea with the deeper North Sea, already started in the summer of 1860, over 140 years ago. Since the year 2000 the sampling frequency has strongly increased because of the use of electronic sensors and data logging by computer. Analysis of these temperature data has revealed variations with time scales from tidal, daily, seasonal, inter-annual, to centennial. The tidal temperature variations are generated by advection of the seasonally varying temperature gradient between Wadden Sea and North Sea, while the daily variations are mainly caused by the daily variation of solar radiation. The seasonal variation in sea surface temperature only lags a few days behind the coastal surface air temperature, contrary to the sea surface temperature in the deeper nearby North Sea, which is delayed with about 1 month. The North Atlantic Oscillation index has been used as large-scale proxy for the atmospheric forcing of the Wadden Sea temperature. Only for the winter and spring a significant correlation is found between temperature and the winter index. However, this correlation is so strong that also the annual mean temperature is correlated significantly with the North Atlantic Oscillation. At longer time scales, from decadal to centennial, also large temperature variations are observed, of the order of 1.5 °C. However, these are not related to long-term changes of the North Atlantic oscillation. These long-term temperature changes involve a cooling of about 1.5 °C in the first 30 years of the record and a similar warming in the last 25 years. In between, these long-term changes were smaller and more irregular. Similar conclusions can also be applied to individual seasons as well as to the date of the onset of spring.  相似文献   

8.
Rüdiger  Berghahn 《Marine Ecology》1996,17(1-3):251-260
Abstract. The brown shrimp Crangon crungon is considered to be a key species in the coastal waters of the North Sea and in particular in the Wadden Sea, since it occurs in masses and acts as both a highly efficient predator and important prey. The Wadden Sea is the nursery ground for brown shrimp. In 1990, a mass invasion of O-group whiting ( Merlangius merlangus ) reduced Crangon numbers to almost zero. Similar events, which had previously been reported in the 19th century, were observed in 1959 (whiting), 1970 (cod, Gadus morhua ), and 1983 (cod and whiting).
The 1990 invasion and the decline of the shrimp stock are documented on the basis of surveys that are compared with the years preceding and following. Consequences for the food web in the Wadden Sea and the reasons for the complete recovery of the shrimp stock in each case within one year after its collapse are discussed with regard to the Crangon crungon behaviour and reproductive potential. Sampling error and the impact of fisheries are also considered.  相似文献   

9.
Daily observations of the sea surface temperature in the Marsdiep tidal inlet, which connects the shallow Dutch western Wadden Sea with the deeper North Sea, already started in the summer of 1860, over 140 years ago. Since the year 2000 the sampling frequency has strongly increased because of the use of electronic sensors and data logging by computer. Analysis of these temperature data has revealed variations with time scales from tidal, daily, seasonal, inter-annual, to centennial. The tidal temperature variations are generated by advection of the seasonally varying temperature gradient between Wadden Sea and North Sea, while the daily variations are mainly caused by the daily variation of solar radiation. The seasonal variation in sea surface temperature only lags a few days behind the coastal surface air temperature, contrary to the sea surface temperature in the deeper nearby North Sea, which is delayed with about 1 month. The North Atlantic Oscillation index has been used as large-scale proxy for the atmospheric forcing of the Wadden Sea temperature. Only for the winter and spring a significant correlation is found between temperature and the winter index. However, this correlation is so strong that also the annual mean temperature is correlated significantly with the North Atlantic Oscillation. At longer time scales, from decadal to centennial, also large temperature variations are observed, of the order of 1.5 °C. However, these are not related to long-term changes of the North Atlantic oscillation. These long-term temperature changes involve a cooling of about 1.5 °C in the first 30 years of the record and a similar warming in the last 25 years. In between, these long-term changes were smaller and more irregular. Similar conclusions can also be applied to individual seasons as well as to the date of the onset of spring.  相似文献   

10.
As a result of aquaculture activities Pacific oysters Crassostrea gigas (Thunberg, 1793) have invaded the European Wadden Sea. Using a variable noncoding mitochondrial marker, we show that the invaded range is the result of two independent invasions. Haplotype frequencies point towards two separate groups, one in the southern and the other in the northern Wadden Sea. We found virtually no genetic differentiation throughout the southern range and the putative source from British Columbia, Canada, suggesting that the Southern region can be considered as a closed population. In the North, mismatch distributions, haplotype ordination and isolation-by-distance analysis suggest a stronger, persistent impact of aquaculture on invasive populations. Due to the ongoing supply of new genetic material from hatchery production the northern invasive populations can therefore be considered as an open population highlighting the importance of aquaculture practice on the genetics of this keystone invader in the Wadden Sea.  相似文献   

11.
In 1995 the seasonal development of concentrations of both phytoplankton and larvae of the bivalve Macoma balthica was studied in the coastal zone behind the back-barrier island of Spiekeroog (German Wadden Sea). In July/August larvae reached maximum concentrations of about 1000 to 4200 ind. m−3 (depending on the sampling site and tidal period), probably in relation to a phytoplankton bloom in July. This observation of an unusually late maximum of Macoma larvae is discussed in detail in connection with the recent literature available about spawning of M. balthica in the Wadden Sea.  相似文献   

12.
The Wadden Sea (North Sea, Europe) is a shallow coastal sea with high benthic and pelagic primary production rates. To date, no studies have been carried out in the Wadden Sea that were specifically designed to study the relation between pelagic respiration and production by comparable methods. Because previous studies have suggested that the import of primary-produced pelagic organic matter is important for benthic Wadden Sea carbon budgets, we hypothesised that on an annual average the northern Wadden Sea water column is autotrophic. To test this hypothesis, we studied annual dynamics of primary production and respiration at a pelagic station in a shallow tidal basin (List Tidal Basin, northern Wadden Sea). Since water depth strongly influences production estimates, we calculated primary production rates per unit area in two ways: on the basis of the mean water depth (2.7 m) and on the basis of 1 m depth intervals and their respective spatial extent in the List Tidal Basin. The latter more precise estimate yielded an annual primary production of 146 g C m− 2 y− 1. Estimates based on the mean water depth resulted in a 40% higher annual rate of 204 g C m− 2 y− 1. The total annual pelagic respiration was 50 g C m− 2 y− 1. The P/R ratio varied between seasons: from February to October the water column was autotrophic, with the highest P/R ratio of 4–5 during the diatom spring bloom in April/May. In autumn and winter the water column was heterotrophic. On an annual average, the water column of the List Tidal Basin was autotrophic (P/R 3). We suggest that a large fraction of the pelagic produced organic matter was respired locally in the sediment.  相似文献   

13.
In this paper , we present a novel Kalman filter approach to combine a hydrodynamic model-derived lowest astronomical tide (LAT) surface with tide gauge record-derived LAT values. In the approach, tidal water levels are assimilated into the model. As such, the combination is guided by the model physics. When validating the obtained “Kalman-filtered LAT realization” at all tide gauges, we obtained an overall root-mean-square (RMS) difference of 15.1 cm. At the tide gauges not used in the data assimilation, the RMS is 17.9 cm. We found that the assimilation reduces the overall RMS difference by ～ 31% and ～ 22%, respectively. In the Dutch North Sea and Wadden Sea, the RMS differences are 6.6 and 14.8 cm (all tide gauges), respectively. Furthermore, we address the problem of LAT realization in intertidal waters where LAT is not defined. We propose to replace LAT by pseudo-LAT, which we suggest to realize similarly as LAT except that all water level boundary conditions and assimilated tidal water levels have to be enlarged by a constant value that is removed afterward. Using this approach, we obtained a smooth reference surface for the Dutch Wadden Sea that fits LAT at the North Sea boundary within a few centimeters.  相似文献   

14.
In the Dutch Wadden Sea a whelk fishery existed from 1900 to 1970. The high post-war catches, following a period without fishery, and the subsequent decrease in catches when fishery was reopened, allowed us to estimate fishery mortality and recruitment. Information on natural mortality and growth, which were required in the estimation procedure, was obtained from previous studies elsewhere. Natural mortality could be calculated from an experiment with marked whelks in the Thames estuary. Age-length relations could be used from French investigations on whelks near the Channel Islands. The size of a potential whelk population could be estimated. Effects of fishery and possible causes of the disappearance of the whelk from the Dutch Wadden Sea are discussed.  相似文献   

15.
Mass mortalities of common eiders Somateria mollissima have been ascribed to high parasite loads. However, the actual role of parasites in mortalities is disputed as in the case of a mass mortality of eiders in the Wadden Sea in the winter of 1999/2000. A critical evaluation of the role of parasites in eider mass mortalities is hampered by (1) a lack of data on actual parasite loads of the birds involved, (2) missing regional data for comparison, and (3) a lack of unbiased samples: investigations are often based on dead beached individuals, which are presumably the more heavily infected birds of a population and thus more likely to die and be washed ashore. Although published data on parasite loads in birds of the winter 1999/2000 mortality are available, no data on background parasitism in eiders from the Wadden Sea exist, making an evaluation of the potential role of parasites in this mortality event difficult. By investigating endoparasites of 102 eiders affected by an oil spill in the northern Wadden Sea in winter 1998/1999, we provide a data set of background parasitism in wintering eiders from the Wadden Sea. We found 13 different parasite taxa with high prevalence values (% infected birds) in the acanthocephalan Profilicollis botulus, the nematode Amidostomum acutum, cestodes and trematodes. In some taxa we observed pronounced differences in prevalence values between juvenile eiders and adults, as well as between adult sexes. The parasite composition shows that bivalves, crabs (Carcinus maenas) and other crustaceans are important sources of infections by being intermediate hosts. This is partly mirrored in the food content of eider stomachs where bivalves and crabs were predominantly found. Intensities of the acanthocephalan P. botulus, suspected of causing eider mortalities, were especially high in juveniles (1112 ± 416 ind per infected host), but lower in adult males (40 ± 7) and adult females (81 ± 18). However, no extraordinary mortality event was observed in the winter of 1998/1999, indicating no or a very weak effect of the parasites on host condition. A comparison with the parasite loads of eiders from the mass mortality in the winter of 1999/2000 shows that parasite numbers were by no means exceptional for birds from the area. Hence, parasites alone are unlikely to have caused this mortality. Regional background parasite loads are important to differentiate between primary and secondary roles of parasites in anomalous mortality events.  相似文献   

16.
In this paper a contribution is made to the ongoing debate on which brown shrimp generation mostly sustains the autumn peak in coastal North Sea commercial fisheries: the generation born in summer, or the winter one. Since the two perspectives are based on different considerations on the growth timeframe from settlement till commercial size, the Dynamic Energy Budget (DEB) theory was applied to predict maximum possible growth under natural conditions. First, the parameters of the standard DEB model for Crangon crangon L. were estimated using available data sets. These were insufficient to allow a direct estimation, requiring a special protocol to achieve consistency between parameters. Next, the DEB model was validated by comparing simulations with published experimental data on shrimp growth in relation to water temperatures. Finally, the DEB model was applied to simulate growth under optimal food conditions using the prevailing water temperature conditions in the Wadden Sea. Results show clear differences between males and females whereby the fastest growth rates were observed in females. DEB model simulations of maximum growth in the Wadden Sea suggest that it is not the summer brood from the current year as Boddeke claimed, nor the previous winter generation as Kuipers and Dapper suggested, but more likely the summer generation from the previous year which contributes to the bulk of the fisheries recruits in autumn.  相似文献   

17.
In different parts of the Dutch Wadden Sea, relationships between intertidal level and abundance of marine macrozoobenthos were similar. Numerical densities, biomass and species richness increased from values close to 0 at the high-water level to maximum values around mean-tide level (numbers) or halfway between this level and low-tide level (biomass). Species richness hardly declined below mean-tide level, whereas mean weight per individual continued to increase from high- to low-water level.Biomass was about 45 g ash-free dry weight per m2 at its maximum and declined in an approximately linear way to values close to 0 at the high-water level and to about 7 g per m2 at the low-water level. These two linear relationships were used to predict biomass changes on intertidal flats of the Wadden Sea at various scenarios of sea-level rise and bottom subsidence. Net sea-level rise is expected to result in increased amounts of intertidal zoobenthos in areas with predominantly high tidal flats, but in declines in lower areas. However, such changes will occur only if sea-level rise proceeds too fast to be compensated by extra sedimentation. Bottom subsidence as a consequence of gas extraction is expected to be too small to cause any measurable change in the benthic fauna.  相似文献   

18.
In terms of grain size, surficial sediment distribution patterns in back-barrier tidal basins (e.g., the East Frisian Wadden Sea, Germany) often show a landward fining trend from the sea boundary to the mainland shore. In addition to the cross-shore patterns, there are lateral grain-size trends toward the watersheds of the basins and toward the watersheds of tidal flats bordered by tidal channels on either side. In the present study, interrelationships between morphological evolution and grain-size trends in the back-barrier tidal basins of the East Frisian Wadden Sea were simulated for a period of 60 years by a process-based forward modeling approach using the Delft3D system. The model outputs show that grain size displays a shoreward fining trend within the basin area, which is consistent with in situ observations; such a trend can be interpreted by the shoreward decrease in the cross-shore maximum velocity. Moreover, the model predicts lateral grain-size trends similar to those observed in the tidal basins: coarser sediment remains in the inlets and channels, while finer sediment settles at the tidal watersheds and on the tidal flats between channels. The spatial patterns of tidal flat sediment grain size within the tidal basins are thus related to the distance from the sea boundary and from the tidal channels. The modeling exercise also indicates that the development of the grain-size pattern observed in the East Frisian Wadden Sea is accomplished within a few decades, and that the time periods required to reach equilibrium are much shorter for grain size than for bed elevation. Evidently, spatial grain-size information can be used to assess sediment transport and morphological adaptation processes as, for example, attempted in sediment trend analysis procedures.  相似文献   

19.
Suction dredging for cockles removes large cockles from tidal flats and may also cause mortality of non-target fauna and make the habitat less suitable for some species. This study examines whether suction dredging for cockles on tidal flats of the Dutch Wadden Sea had affected densities of non-target fauna, directly after fishing and one year later. Densities of non-target fauna in two randomly chosen undredged locations were compared to densities at the surrounding heavily commercially dredged area. A significant negative effect of cockle dredging on densities of 0-group Macoma balthica was observed and this effect persisted one year after dredging. The dredged area appeared to be less suitable for settlement of mussels Mytilus edulis. No significant effects of dredging on the mudsnail Hydrobia ulvae and on 0 and 1-group C. edule were found. For the mobile young Macoma balthica it seems unlikely that the effect found after one year was still due to the mortality caused by dredging and this suggests that the habitat was less suitable as a consequence of dredging. Thus, even in the highly dynamic ecosystem of the Wadden Sea, effects of bottom disturbance by cockle dredging may persist after one year.  相似文献   

20.
The concept of ecological speciation implies that habitat differences may split a species by strong selection and rapid adaptation even under sympatric conditions. Studies on the polychaete Scoloplos armiger in the Wadden Sea (North Sea) indicate sibling species existing in sympatry: the intertidal ‘Type I’ with holobenthic development out of egg cocoons and the subtidal ‘Type S’ producing pelagic larvae. In the current study, Types I and S are compared in habitat-related traits of reproductive timing and physiological response to hypoxia and sulphide. Spawnings of Type I and Type S recorded over six years overlap in spring and both appear to be triggered by a rise in seawater temperature above 5 °C. Type S exhibits an additional autumn spawning (at seawater temperatures around 10 °C) which was previously unknown and is absent in Type I. The overall abundance of pelagic larvae in the Wadden Sea is higher in spring than in autumn. Tolerance of both sulphide and hypoxia was lower in Type S than in Type I. This correlates with a 5 to 10-fold lower sulphide concentration in the subtidal compared to the intertidal habitat. Physiological tolerance and divergence in developmental mode appear as traits which may have led to reproductive isolation between Type I and Type S. Their role in allopatric and sympatric speciation scenarios in S. armiger is discussed. Since the pelagic dispersal mode has been neglected so far, a reassessment of population dynamics models for S. armiger is suggested.  相似文献   

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