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1.
For a long time, “spelaeoid” (cave-bear-like) bears, Ursus (Spelearctos) spp., were believed to be almost purely European animals. Their geographic range has recently been extended to the east, in southern Siberia, Transbaikalia, Kirghizia, Mongolia and Korea. Two unexpected new findings, presented here in detail, significantly change existing views on the distribution of cave bears; both were found in North-Eastern Siberia, far beyond the Arctic Circle, more than 1500 km to the north-east of the previously accepted range.One of the fossils is a mandible, found near the town of Cherskiy at 68.73°N, 161.38°E. The analysis of local geology and accompanying mammal fossils suggests that it comes from the Olyorian Fauna (Early to early Middle Pleistocene). Morphologically, the Cherskiy mandible is closest to Ursus savini, a small middle Pleistocene cave bear from the British Cromer Forest-bed Formation, but differs in having a slightly more advanced dentition, and thus it is described as a new subspecies Ursus savini nordostensis. Another newly recognized fossil of the “spelaeoid” bear is an astragalus found at the Oskhordokh site at 67.54°N, 135.67°E, on a large gravel bar on the right bank of the Adycha River. This specimen is attributed to Ursus cf. deningeri.The paper also presents an interesting example of the interaction between classical and “molecular” palaeontology.The new finds significantly change existing ideas on the ecology and evolution of cave bears, some of the most remarkable members of the extinct Pleistocene megafauna.  相似文献   

2.
An isotopic investigation of upper Pleistocene mammal bones and teeth from Scladina cave (Sclayn, Belgium) demonstrated the very good quality of collagen preservation. A preliminary screening of the samples used the amount of nitrogen in whole bone and dentine in order to estimate the preserved amount of collagen before starting the extraction process. The isotopic abundances of fossil specimens from still-extant species are consistent with their trophic position. Moreover, the15N isotopic abundance is higher in dentine than in bone in bears and hyenas, a phenomenon already observed in modern specimens. These results demonstrate that the isotopic compositions of samples from Scladina cave can be interpreted in ecological terms. Mammoths exhibit a high15N isotopic abundance relative to other herbivores, as was the case in Siberian and Alaskan samples. These results suggest distinctive dietary adaptations in herbivores living in the mammoth steppe. Cave bears are clearly isotopically different from coeval brown bears, suggesting an ecological separation between species, with a pure vegetarian diet for cave bear and an omnivorous diet for brown bear.  相似文献   

3.
The skeleton of a young prime adult cave bear, Ursus spelaeus, was found in Chiostraccio Cave (Siena, Tuscany, central Italy), only slightly buried under rock falls. The specimen was dated yielding a conventional age of 24,030 ± 100 14C yr BP (29,200–28,550 cal yr BP), which makes it the latest known representative of the species in Italy. The skeleton was accompanied by the remains of wolf (Canis lupus), wild boar (Sus scrofa), aurochs (Bos primigenius), red deer (Cervus elaphus), roe deer (Capreolus capreolus), bat (Vespertinus murinus), and crow (Corvus monedula). The site seems confirming that the latest Italian U. spelaeus populations shared the risk of intrusion. The association of the cave bear with other animals suggests that the assemblage is an attritional palimpsest of remains of different species not originally associated in life. Cave bears were probably more vegetarian than brown bears and possibly became extinct when plant productivity dropped at the onset of MIS 2. Central and southern Italy may have offered isolated and sheltered refugia for cave bears.  相似文献   

4.
Univariate and multivariate statistics were applied to analyse the morphometrical variability of 4920 upper cheek teeth (P4, M1 and M2) of cave bears from 123 geographical sites (180 samples) of different Pliocene – Pleistocene ages. The analysed specimens included those belonging to the big cave bears Ursus kudarensis, U. deningeri, U. spelaeus (three subspecies) and U. kanivetz (including U. ingressus), as well as the small cave bear U. rossicus. The information‐theoretical parameters (Shannon entropy and orderliness (Von Foerster, 1960: On self‐organizing systems and their environments. In Self‐Organizing Systems, 31–50. Pergamon Press, London) were used to estimate tooth diversity in different teeth, different taxa and in selected local chrono‐populations. Multivariate allometry coefficients (Klingenberg, 1996: Multivariate allometry. In Advances in Morphometrics, 23‐49. Plenum Press, New York) were used to describe the relationships of different ‘parts’ of a tooth and to compare allometric patterns amongst species or selected local samples. A multivariate analysis showed a significant overlap of the size/shape parameter ranges in deningeroid and spelaeoid bears within morphological spaces. Within the cave bear lineage, the Deninger's bear has the greatest morphological diversity index (entropy) of all the teeth overall, and the lowest diversity is observed in the final taxon of this lineage – U. kanivetz (=ingressus). The P4 and M2 diversity showed multidirectional correlations with elevation above sea level amongst several ‘local’ populations of Late Pleistocene cave bears. The morphological disparities between the studied taxa are in close agreement with the distances in the available schemes of genetic differentiation based on ancient mitochondrial DNA. The split of U. kudarensis and U. deningeri has a good bootstrap support, which corresponds to the hypothesis about their parallel evolution. The small cave bear U. rossicus is placed between U. arctos and U. deningeri. The phylogenetic signal is more pronounced in the variability of teeth in comparison with other skeletal remains of cave bears (cranium, mandible, or metapodial bones).  相似文献   

5.
Tens of thousands of palaeontological and archaeological remains were collected by William Pengelly during 19th century excavations of Kents Cavern, but are now widely dispersed between museums. This has previously precluded spatial analysis. We have now assembled available museum records into a single database, and, using our previously-reconstructed Pengelly excavation map as a base, we have been able to exploit the unique Pengelly location code to set up a GIS mapping system. This allows, for the first time, the analysis of spatial patterns. In addition, the GIS serves to highlight potential problems of recording or curation in the original data. Here we report on the construction of the GIS system and its first use in the analysis of spatial distribution of bear remains. The maps demonstrate that Ursus deningeri entered the cave through a now-sealed High Level Chamber entrance at the back of the cave in the middle Pleistocene, whereas Ursus arctos accessed the cave in the late Pleistocene through the now-sealed Northeast Gallery entrance. The denning areas are reconstructed as Labyrinth/Bear's Den for U. deningeri and Vestibule/Great Chamber for U. arctos. Considerable post-mortem re-distribution of the remains of both species is indicated.  相似文献   

6.
Competition between taxa related to climate changes has been proposed as a possible factor in Pleistocene megafaunal extinctions, and here we present isotope evidence of the diets of three co-existing bear species [black bear (Ursus americanus), brown bear (Ursus arctos), and the now extinct short-faced bear (Arctodus simus)] from a locale in western North America dating to the Late (Terminal) Pleistocene (~14.5–11.7 ka). The three bear species were found at several sites on Vancouver Island, on the western coast of Canada. To examine the chronological overlap and niche partitioning between these species of bear, we used direct radiocarbon dating, stable isotope analysis and ZooMS proteomic identification methods. Here we present new radiocarbon evidence from Terminal Pleistocene U. americanus, U. arctos and A. simus from several sites on the island, along with both bulk collagen and compound-specific isotope data for these species. Radiocarbon dates confirm the chronological overlap of Arctodus and both Ursus species in the montane regions of the island at the end of the Pleistocene. Stable isotope data reveal niche differentiation between these species, with U. americanus occupying a distinctly lower trophic position than the other two taxa.  相似文献   

7.
Paleobiologists generally agree that within the past 10,000 yr North American black bears (Ursus americanus) have decreased in body and tooth size. Some researchers infer that diminution was gradual and continuous; thus, one might infer that a specimen is old if it is larger than an average-size modern bear. Ursid remains recovered in the 1950s from Lawson Cave, Missouri, that are larger than some modern bears have been reported to date to the late Pleistocene, but association with modern taxa, taphonomic considerations, and a radiocarbon date of 200 yr B.P. indicate that they are modern. Modern specimens from Lawson Cave and other parts of the American Midwest are relatively large compared to modern North American black bears from other areas, suggesting that many supposed late Pleistocene bears from the area might be modern also.  相似文献   

8.
Cave bears (Ursus spelaeus) are an iconic component of the European late Quaternary Ice Age megafauna. Recent demographic analyses based on cave bear mtDNA sequences and refined radiocarbon dating indicate that cave bear population size and genetic diversity started to decline some 50 kilo years ago (kya). Hence, neither the coldest phase of the last glaciation (started some 24 kya), nor the colonization of Europe by Palaeolithic hunters (started some 45 kya) coincides with the beginning of population decline. Here, we reconstructed cave bear climatic niche evolution through time. Then, we performed spatially explicit population viability analyses to assess cave bear demographics through time in response to climatic changes, human effects on bear survival and their combination. We found that climate change was responsible for a 10‐fold decrease in cave bear population size after 40 kya. However, climate change on its own could not explain U. spelaeus extinction at 24 kya. Additional negative effects consistent with human population expansion are required to explain both U. spelaeus' retreat from eastern Europe since 40 kya and its final extinction.  相似文献   

9.
Norwegian caves contain stratigraphical information regarding the historical faunal composition valuable in the assessment of postglacial colonization and phylogeographical structure. In some of these limestone caves, brown bear ( Ursus arctos L.) remains have been excavated and radiometrically dated. We present osteology (95 traits defined) from 29 brown bears found within 22 Norwegian caves above 658N. Our data span the time interval 6210–420 14C yr BP. The sex of individuals was based on dimorphic canines, while age was determined by cementum analysis and body size estimated from regressions between morphological traits compared to a contemporary reference collection. Five females and 14 males were recognized, while the remaining bears could not be sexed because of their small size, low age or lack of canines. The ages of 26 bears (tooth cementum age and estimates) ranged from juveniles (<0.5 year) to adults (23 years), the majority being old animals. Both sexes have used these caves for over-wintering lairs, as seen in several caves, where denning is also suggested based on the observations of adult females and cubs. There were no signs of predator or human transported bear remains, suggesting natural mortality. The age of the bears suggests that young or old bears may have died from insufficient nutrient storage, environmental harshness, or may have drowned in spring or autumnal flooding. We conclude that Norwegian brown bears have used these caves sporadically for thousands of years, which suggests that caves are not a preferred site for the brown bear during hibernation or denning.  相似文献   

10.
The mandible of a polar bear (Ursus maritimus Phipps) found in about 1920 at Kjul Å, North Jutland, and described by Nordmann & Degerbol in 1930. has been l4C dated to 11.100 ± 160 B.P. It is so far the only find of polar bear in Denmark. Comparison with recent 14C datings of Swedish and Norwegian polar bears shows that the Danish specimen was a member of a southern Scandinavian Late Weichselian population. The contemporaneous Zirphaeu sea deposits can be regarded as the boreal-arctic shallow water equivalent of the arctic Upper Saxicava sand deposits from northern Jutland. The polar bear mandible, however, was deposited on land, as was the metacarpal bone of a brown bear ( Ursus arctos ) from the nearby Nr. Lyngby locality of Allerød age. The overall picture of the Late Weichselian mammal fauna in Denmark shows a mixed composition of different ecotypes. Their sympatric occurrence points at a unique environment not comparable to any now existing, and probably related to the very low latitude of the Weichselian ice sheet.  相似文献   

11.
Recent investigations of a limestone solution cave on the Queen Charlotte Islands (Haida Gwaii) have yielded skeletal remains of fauna including late Pleistocene and early Holocene bears, one specimen of which dates to ca. 14,400 14C yr B.P. This new fossil evidence sheds light on early postglacial environmental conditions in this archipelago, with implications for the timing of early human migration into the Americas.  相似文献   

12.
Two hypotheses have been proposed to explain the origin of marine isotope stage (MIS) 11 deposits in small Bermudian caves at +21 m above modern sea level: (1) a +21 m MIS 11 eustatic sea-level highstand, and (2) a MIS 11 mega-tsunami event. Importantly, the foraminifera reported in these caves have yet to be critically evaluated within a framework of coastal cave environments. After statistically comparing foraminifera in modern Bermudian littoral caves and the MIS 11 Calonectris Pocket A (+21 m cave) to the largest available database of Bermudian coastal foraminifera, the assemblages found in modern littoral caves – and Calonectris Pocket A – cannot be statistically differentiated from lagoons. This observation is expected considering littoral caves are simply sheltered extensions of a lagoon environment in the littoral zone, where typical coastal processes (waves, storms) homogenize and rework lagoonal, reefal, and occasional planktic taxa. Fossil protoconchs of the Bermudian cave stygobite Caecum caverna were also associated with the foraminifera. These results indicate that the MIS 11 Bermudian caves are fossil littoral caves (breached flank margin caves), where the total MIS 11 microfossil assemblage is preserving a signature of coeval sea level at +21 m. Brackish foraminifera (Polysaccammina, Pseudothurammina) and anchialine gastropods (95%, >300 individuals) indicate a brackish anchialine habitat developed in the elevated caves after the prolonged littoral environmental phase. The onset of sea-level regression following the +21 m highstand would first lower the ancient brackish Ghyben-Herzberg lens (<0.5 m) and flood the cave with brackish water, followed by drainage of the cave to create a permanent vadose environment. These interpretations of the MIS 11 microfossils (considering both taphonomy and paleoecology) are congruent with the micropaleontological, hydrogeological and physical mechanisms influencing modern Bermudian coastal cave environments. In conclusion, we reject the mega-tsunami hypothesis, concur with the +21 m MIS 11 eustatic sea-level hypothesis, and reiterate the need to resolve the disparity between global marine isotopic records and the physical geologic evidence for sea level during MIS 11.  相似文献   

13.
Sur‐les‐Creux rockshelter is located in the Prealps of southwestern Switzerland. The sequence of deposits in the rockshelter is 80 cm thick and consists of weathered gravels in a phosphate‐rich matrix. A few Middle Palaeolithic artifacts and the bones of cave bear (Ursus spelaeus) were recorded in the fill. We present the results of sedimentological, geochemical, and micromorphological analyses of the rockshelter sediment. All analyses suggest an endokarstic origin of the sediments. The alteration cortices of the gravels imply in situ weathering over a long period. The phosphates are essentially biogenic and have an apatitelike nature. Phosphatization and intense mixing of the sediment are attributed to cave bear (digging of lairs, input of excrements, and carcasses). Only rare carnivore coprolites (lynx) were preserved in the cave deposits. © 2004 Wiley Periodicals, Inc.  相似文献   

14.
Doklady Earth Sciences - New finds of brown bear (Ursus arctos L., 1758) fossil remains from the territory of Yakutia, namely, skulls and mandibular bones, have been investigated. The new finds are...  相似文献   

15.
The cave bear was a prominent member of the Upper Pleistocene fauna in Eurasia. While breakthroughs were recently achieved with respect to its phylogeny using ancient DNA techniques, it is still challenging to date cave bear fossils beyond the radiocarbon age range. Without an accurate and precise chronological framework, however, key questions regarding the palaeoecology cannot be addressed, such as the extent to which large climate swings during the last glacial affected the habitat and possibly even conditioned the final extinction of this mammal. Key to constraining the age of cave bear fossils older than the lower limit of radiocarbon dating is to date interlayered speleothems using 230Th/U. Here we report new results from one such site in the Eastern European Alps (Schwabenreith Cave), which yielded the highest density of bones of cave bear (Ursus spelaeus eremus). Although dating of the flowstones overlying this fossiliferous succession was partly compromised by diagenetic alteration, the 230Th/U dates indicate that the bear hibernated in this cave after about 113 ka and before about 109 ka. This time interval coincides with the equivalent of Greenland Stadial 25, suggesting possible climate control on the cave bear's habitat and behaviour. © 2019 The Authors. Journal of Quaternary Science Published by John Wiley & Sons Ltd  相似文献   

16.
This review provides an up-to-date synthesis of the matrilineal phylogeography of a uniquely well-studied Holarctic mammal, the brown bear. We extend current knowledge by presenting a DNA sequence derived from one of the earliest known fossils of a polar bear (dated to 115 000 years before present), a species that shares a paraphyletic mitochondrial association with brown bears. A molecular clock analysis of 140 mitochondrial DNA sequences, including our new polar bear sequence, provides novel insights into the times of origin for different brown bear clades. We propose a number of regional biogeographic scenarios based on genetic data, divergence time estimates and paleontological records. The case of the brown bear provides an example for researchers working with less well-studied taxa: it shows clearly that phylogeographic models based on patterns of modern genetic variation alone can be substantially improved by including data on historical patterns of genetic diversity in the form of ancient DNA sequences derived from accurately dated samples and by using an approach to divergence-time estimation that suits the data under analysis. Using such approaches it has been possible to (i) establish that the processes shaping modern genetic diversity in brown bears acted recently, within the last three glacial cycles; (ii) distinguish among hypotheses concerning species’ responses to climatic oscillations in accordance with the lack of phylogeographic structure that existed in brown bears prior to the last glacial maximum (LGM); (iii) reassess theories linking monophyletic brown bear populations to particular LGM refuge areas; and (iv) identify vicariance events and track analogous patterns of migration by brown bears out of Eurasia to North America and Japan.  相似文献   

17.
Specimens of fossil wood preserved lignified in Pliocene brown coal and identified as Pinus armandii Francher come from an opencast coalmine at Longling in western Yunnan Province, China. Phytochemical investigation of the fossil wood isolated using liquid column chromatography seven compounds （1-7） including a new fluorene derivative named 11,11-dimethyl-11H-benzo[b]fluorene. A further 28 volatiles were detected by gas chromatography-mass spectrometry （GC-MS）. Spectroscopic investigation methods, including MS and 1D and 2D-NMR techniques elucidated the structure of the seven compounds. Two types of natural products, isopimara and stilbene commonly occuring in extant and Pliocene fossil P. armandii indicate phytochemical fidelity during burial under certain circumstances in sediments. Discovery of stilbenes that can inhibit the activities of wood-destroying fungi in the Pliocene P. armandii prompts the assumption that the chemical preservation of this Pliocene fossil wood of P. armandii in brown coal might contribute to the presence of inner natural inhibitors against wood-destroying fungi.  相似文献   

18.
The investigation of a limestone cave on the Ryonggok-ri of Sangwon County, North Hwanghae Province, DPRK, has yielded many faunal remains including Late Pleistocene deer, brown bears and horses. Uranium series dating of fossil teeth yielded dates of ~72ka at the base of Layer 2 – the bone-bearing unit – and ~44ka at the top. The rich and diverse mammal remains of Ryonggok Cave No. 1 are indicative of warm and humid temperate conditions during the Late Pleistocene, and reflect the presence of both dense forests and open grassland. In addition, pollen and spores from the site include two families and 24 genera of trees, six families and a genus of grasses and herbs, and two families and six genera of ferns, corroborating the environments inferred from the faunal remains.  相似文献   

19.
The cave bear ( Ursus spelaeus ) was one of several spectacular megafaunal species that became extinct in northern Eurasia during the late Quaternary. Vast numbers of their remains have been recovered from many cave sites, almost certainly representing animals that died during winter hibernation. On the evidence of skull anatomy and low δ15N values of bone collagen, cave bears appear to have been predominantly vegetarian. The diet probably included substantial high quality herbaceous vegetation. In order to address the reasons for the extinction of the cave bear, we have constructed a chronology using only radiocarbon dates produced directly on cave bear material. The date list is largely drawn from the literature, and as far as possible the dates have been audited (screened) for reliability. We also present new dates from our own research, including results from the Urals. U. spelaeus probably disappeared from the Alps and adjacent areas – currently the only region for which there is fairly good evidence – c . 24 000 radiocarbon years BP ( c . 27 800 cal. yr BP), approximately coincident with the start of Greenland Stadial 3 ( c . 27 500 cal. yr BP). Climatic cooling and inferred decreased vegetational productivity were probably responsible for its disappearance from this region. We are investigating the possibility that cave bear survived significantly later elsewhere, for example in southern or eastern Europe.  相似文献   

20.
There is little fossil evidence for cave arthropods. Small size, disfunctional wings, extremely long antenna, dense minute setation, palidity and reduced eyes are typical adaptations to life in caves shown by the cockroach Mulleriblattina bowangi gen. et sp. n., found together with epigeic Crenocticola svadba sp. n., both from Myanmar amber and belonging to the cavernicolous cockroach family Nocticolidae. These lineages of earliest still living cavernicoles suggest large, numerous caverns, lava tubes or caves within the source area. They provide the first unequivocal evidence for the Mesozoic origin of any living troglomorphic organism, and explain the “long branches” in DNA analyses. Phylogenetic trees show little hierarchical structure and place Latindiinae and Myrmecoblatta within the explosively radiating Nocticolidae. Biogeography indicates a common cosmopolitan Early Cretaceous ancestor except for 8 (of 49) species of true Nocticolidae, which diverged during the Late Cretaceous breakup of Gondwana. A review of all troglofauna documents no other unequivocal pre-Cenozoic cave biotas (including vertebrates). Stable environments and small populations result in a short-time (˂3 Ma) origination of bizarre forms and long term extinctions (˃30 Ma).  相似文献   

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