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1.
Basing upon the total of 1,081 samples collected by a large plankton net (160 cm in mouth diameter) in the Pacific Ocean, the geographical and vertical distribution ofGonostoma gracile were studied. The species is distributed in the water masses of the Kuroshio, the Kuroshio Extension, the North Pacific Current, the Oyashio, the North Pacific Subarctic Water and the western North Pacific Central Water. The center of distribution lies in the Kuroshio area off Japan. Vertically, the species occurs between the depths of 200 and 1,000 m, mainly 300–700 m, both during daytime and at night. A part of population might come up to 0–200 m at night, although its biomass is negligibly small in comparison to that remaining in 300–700 m layer. Postlarvae are found in 300–700 m layer, mainly at 300–500 m, and apparently do not undertake diurnal vertical migration.  相似文献   

2.
在IONESS系统中浮游介形类的昼夜垂直移动   总被引：1，自引：0，他引：1  
根据IONESS系统对浮游动物的分层采样结果,研究浮游介形动物的昼夜垂直移动.浮游介形类存在着整个类群的昼夜垂直移动,白天其丰度最大值记录于200～300m层,并往上层水域递减,而夜间则明显往上层密集,丰度最大值提升至50～100m层;就物种多样性而言,由于工作水层(0～1000m)以深水层的众多种类于夜间上升至1000m以浅水层,导致工作水层内总种数由白天的49种剧增至夜间的78种,增幅达60%,各分层的种类数都有较明显的增加,750～1000m层的增幅最大,达150%,其次为50～100m层(增幅达125%);各种浮游介形类有不同的昼夜垂直移动模式,但绝大多数种类属于夜间上升型,少数种类属于不敏感类型,极少数种类却具夜间下降的移动特点.浮游介形类虽具明显的昼夜垂直移动,但这一变动往往局限于一定水深范围,从而反映出介形类的层状分布特点,可划分为广深分布型、上层分布型、中层分布型和下层分布型.  相似文献   

3.
We describe the distribution and abundance of the midwater fish community, between 200 m and 500 m, in the North Pacific. The main area of interest was the Subarctic Pacific gyres, but we include species from the Bering Sea and the Sea of Okhotsk. There were 196 species identified in each gyre, 38 of which were common to both gyres. The most abundant species belong to the family Myctophidae and the most ecologically important myctophid probably is Stenobrachius leucopsarus. This species could have a biomass of approximately 21 million tonnes (t) in the Subarctic Pacific (including the Bering Sea and Sea of Okhotsk). S. leucopsarus is a small (about 8 cm maximum length) fish that lives up to 7 years. It is prey for a variety of other fishes, birds and mammals and may migrate into the mixing layer each evening where it feeds mainly on euphausiids and copepods.The total abundance of midwater fishes appears to be large relative to total catches of other fish in the same areas. The vertical migratory behaviour of some of the residents provides a mechanism to transfer production out of the mixing layer. The movement into the surface layer by some fishes at night indicates that dynamic changes occur in the midwater community between the day and night, and the ecosystem dynamics in the surface layer are different in the day and in the night. This behaviour and the huge biomass relative to commercial species means that the dynamics of fish communities in the Subarctic Pacific are complex and need to be studied over a 24 hour period. The large biomasses may eventually attract commercial interest, thus it is important to establish international, cooperative programs now to learn more about the dynamics of these populations and the relationships with other species.  相似文献   

4.
南沙群岛海区毛颚动物的昼夜垂直移动   总被引：10，自引：2，他引：10  
对1997年11月和1999年4月、7月在南沙群岛海区3个昼夜连续测站采集的毛颚动物样品进行了研究。结果表明，这些水域的毛颚动物的昼夜垂直移动可以划分为3个类型：种类作显著移动，不作显著移动和仅在夜晚移动。作为昼夜垂直移动显著的种类，又可以把它们划分为：（1）种类在整个水柱移动；（2）种类在上层（0－60m）移动；（3）种类在中层（60－100m）移动。作为昼夜垂直移动不明显的种类，又可以把它们分为：（1）种类分布在于上层（0－60m）。（2）种类分布于底层（100－200m）。种类在白天下降或附着在海底物体上，仅在夜晚移动。光照，水温，食物和种类特征是影响本水域毛颚动物昼夜垂直移动的重要因素。  相似文献   

5.
A study was undertaken to investigate the spatial, diurnal and interannual dynamics of the pelagic component of Namibian hake stocks. Data were derived from trawl surveys conducted on board the R.V. Dr Fridtjof Nansen during the period 1991–1999. In addition to trawl surveys, acoustic methods were applied concurrently to quantify fish biomass off the bottom (above the headline). The pelagic component of hake was variable in both time and space – it was denser in the north and at depths of 200–500 m. Pelagic densities also correlated well with the density of adult fish (≥30 cm) at the bottom, but not with that of young hake (>18 and <30 cm) at the bottom. Pelagic densities were lowest at midday and highest at night, early morning and late afternoon densities being intermediate between the two. Incorporation of acoustic methods allows for at least partial account to be taken of diurnal variation in determining catchability. The fraction of the biomass off the bottom was not the same each year, thus affecting assessment and management advice for the stock, even if the survey results are treated only as relative indices of abundance. More field studies are needed to investigate the size range and species of hake active in vertical migration.  相似文献   

6.
The vertical distribution of microphytobenthos (MPB) was investigated in intertidal flats of the Nakdong River estuary over different temporal periods. The MPB biomass were measured as chlorophyll a concentration in the sediment, and monitored at two different depth scales. From August 2006 to August 2007, monthly sampling was carried out at four sites to a depth of 10 cm. The vertical distribution exhibited an exponential decline with depth and the slope was closely related to sediment composition and MPB biomass in the surface sediment. The two slightly muddy sand (5–25% mud, <63 μm) sites presented similar trends of seasonal variation to a sediment depth of 10 cm, and no statistical difference was observed in their initial values and the slope of the exponential curves applied to each data set. The site with higher mud content also had higher surface biomass, and the fitted exponential curve had a larger slope than sites with more sand content. The seasonal variations of phaeo-pigments compared to chlorophyll a revealed higher grazing stress on the MPB in summer at the three slightly muddy sand sites, and low grazing stress all year around at the sand (<5% mud) site. In January, April and October 2007, and January 2008, diurnal sampling was conducted at hourly intervals at one slightly muddy sand site. The investigation in January 2007 which was conducted at 1 cm intervals to a depth of 10 cm sediment showed the migration mainly existed in the upper 1 cm. The other diurnal sampling was undertaken to a 1 cm depth at 0.5 mm intervals. An evident migratory rhythm was present, with migration up to the surface during daytime emersion and down into deeper sediment during night and high tide submersion. Migrating cells could reach sediments as deep as 6 or even 10 mm. When the visible thick biofilms formed, the migratory rhythm changed by keeping higher biomass at the surface even at night and during high tide submersion. Generally, the MPB surface biomass was an important factor in structuring the MPB vertical distribution. The vertical distribution to depths of 10 cm was closely related to the sediment composition over the long term, while the MPB migration influenced the vertical distribution over the short term in the top 1 cm of the sediment.  相似文献   

7.
Zooplankton in the coastal upwelling region off northern Chile may play a significant biogeochemical role by promoting carbon flux into the subsurface OMZ (oxygen minimum zone). This work identifies the dominant zooplankton species inhabiting the area influenced by the OMZ in March 2000 off Iquique (20°S, northern Chile). Abundance and vertical distribution studies revealed 17 copepod and 9 euphausiid species distributed between the surface and 600 m at four stations sampled both by day and by night. Some abundant species remained in the well-oxygenated upper layer (30 m), with no evidence of diel vertical migration, apparently restricted by a shallow (40–60 m) oxycline. Other species, however, were found closely associated with the OMZ. The large-sized copepod Eucalanus inermis was found below the oxycline and performed diel vertical migrations into the OMZ, whereas the very abundant Euphausia mucronata performed extensive diel vertical migrations between the surface waters and the core of the OMZ (200 m), even crossing it. A complete assessment of copepods and euphausiids revealed that the whole sampled water column (0–600 m) is occupied by distinct species having well-defined habitats, some of them within the OMZ. Ontogenetic migrations were evident in Eucalanidae and E. mucronata. Estimates of species biomass showed a substantial (>75% of total zooplankton biomass) daily exchange of C between the photic layer and the OMZ. Both E. inermis and E. mucronata can actively exchange about 37.8 g C m−2 d−1 between the upper well-oxygenated (0–60 m) layer and the deeper (60–600 m) OMZ layer. This migrant biomass may contribute about 7.2 g C m−2 d−1 to the OMZ system through respiration, mortality, and production of fecal pellets within the OMZ. This movement of zooplankton in and out of the OMZ, mainly as a result of the migratory behavior of E. mucronata, suggests a very efficient mechanism for introducing large amounts of freshly produced carbon into the OMZ system and should, therefore, be considered when establishing C budgets for coastal upwelling systems.  相似文献   

8.
Species compositions of deep-sea demersal fishes were compared during the day and at night in two cruises of KT-01-8 and KT-02-5. A total of 742 individuals belonging to 28 species were collected in four day and four night hauls in KT-01-8, and in KT-02- 5 128 individuals belonging to 33 species were sampled in three night and four day hauls. Principal Component Analysis (PCA) revealed that there were two distinct groups corresponding to sampling time in both cruises, although in KT-02-5 the species composition from the first haul sampled after sunrise was similar to those found in night hauls. This suggests that the species composition of deep-sea demersal fishes changes between day and night. The existence of a time lag between the sea surface and the sea floor was also suggested, lag being 2–4 hr at 500 m depths in the study area. There may be two possible explanations for the change in deep-sea demersal fish composition between daytime and nighttime: the artificial effect of successive sampling design, and actual fish migration. Since the artificial effect would not be sufficient to explain the change of demersal fish composition, the vertical or horizontal movements of deep-sea demersal fishes would contribute to the faunal change between daytime and nighttime in the study areas.  相似文献   

9.
The diurnal vertical migration of planktonic crustaceans may be divided into two types: Type 1, species with marked diurnal vertical migration; Type 2, species without marked diurnal vertical migration. The former may be further grouped into three patterns: Pattern 1, sinking by day and rising by night; Pattern 2, rising by dawn and evening and sinking by day and midnight; Pattern 3, rising by day and sinking by night. The latter may be further grouped into the following three patterns: Pattern 1, surface layer distribution; Pattern 2, bottom layer distribution; Pattern 3, even distribution between the surface and bottom layers. Among these patterns the two most common ones appear to be the sinking by day and rising by night' and the surface layer distribution. Most of the females are no more marked in the diurnal vertical migration than males in the Xiamen Harbour. The diurnal vertical migration of the animal become more and more conspicuous with the increase of the ages and varies markedly with season  相似文献   

10.
Vertical changes in abundance, biomass and community structure of copepods down to 3000 m depth were studied at a single station of the Aleutian Basin of the Bering Sea (53°28′N, 177°00′W, depth 3779 m) on the 14th June 2006. Both abundance and biomass of copepods were greatest near the surface layer and decreased with increase in depth. Abundance and biomass of copepods integrated over 0–3000 m were 1,390,000 inds. m?2 and 5056 mg C m?2, respectively. Copepod carcasses occurred throughout the layer, and the carcass:living specimen ratio was the greatest in the oxygen minimum layer (750–100 m, the ratio was 2.3). A total of 72 calanoid copepod species belonging to 34 genera and 15 families occurred in the 0–3000 m water column (Cyclopoida, Harpacticoida and Poecilostomatoida were not identified to species level). Cluster analysis separated calanoid copepod communities into 5 groups (A–E). Each group was separated by depth, and the depth range of each group was at 0–75 m (A), 75–500 m (B), 500–750 m (C), 750–1500 m (D) and 1500–3000 m (E). Copepods were divided into four types based on the feeding pattern: suspension feeders, suspension feeders in diapause, detritivores and carnivores. In terms of abundance the most dominant group was suspension feeders (mainly Cyclopoida) in the epipelagic zone, and detritivores (mainly Poecilostomatoida) were dominant in the meso- and bathypelagic zones. In terms of biomass, suspension feeders in diapause (calanoid copepods Neocalanus spp. and Eucalanus bungii) were the major component (ca. 10–45%), especially in the 250–3000 m depth. These results are compared with the previous studies in the same region and that down to greater depths in the worldwide oceans.  相似文献   

11.
A large database representing the bathymetric distribution of 117 species of halocyprid ostracods has been compiled from seven stations forming a transect from the equator to 60°N along 20°W, plus an additional station at 32°N, 65°W. This data base is analysed to examine the latitudinal and bathymetric changes in species composition and diversity of assemblages of this important, yet neglected, holoplanktonic group. At each station stratified sampling of the complete water column from the surface down mostly to 2000 m was carried out both day and night. Each sample resulted from the filtration of at least 2500 m3 of water and was analysed using a consistent protocol. The differences between the day and night profiles are attributable to diel vertical migrations, to local-scale heterogeneity, and possibly to a degree of net avoidance. There is a gradient of increasing species richness and diversity from high to low latitudes. By day, halocyprids are either infrequent or absent from the upper 50 m of the water column, but at night after diel vertical migration they become quite abundant in the epipelagic zone, particularly at low latitudes. Bathymetric profiles show ostracod abundances increase rapidly below the thermocline, reaching maxima at 200-400 m and then declining by at least an order of magnitude at 2000 m. Diversity (both species richness, H′ and evenness, J) also increases below the thermocline and thereafter is either maintained or declines only slightly to 2000 m. There are no relationships among diversity, abundance and productivity, but analysis of the whole database shows that the changes in community structure are consistent with Longhurst’s [Longhurst, A.R., 1998. Ecological Geography of the Sea. Academic Press, San Diego, pp. xiv, 398.] biogeochemical provinces.  相似文献   

12.
The mesozooplankton in both epipelagic and mesopelagic zones is essentially important for the study of ecosystem and biological carbon pump. Previous studies showed that the diel vertical migration(DVM) pattern of mesozooplankton varied among ecosystems. However, that pattern was largely unknown in the Western Pacific Warm Pool(WPWP). The vertical distribution, DVM and community structure of mesozooplankton from the surface to 1 000 m were compared at Stas JL7K(WPWP) and MA(North Pacific Subtropical Gyre, NPSG). Two sites showed similarly low biomass in both epipelagic and mesopelagic zones, which were in accordance with oligotrophic conditions of these two ecosystems. Stronger DVM(night/day ratio) was found at JL7K(1.31) than that at MA(1.09) on surface 0–100 m, and an obvious night increase of mesopelagic biomass was observed at JL7K, which was probably due to migrators from bathypelagic zone. Active carbon flux by DVM of zooplankton was estimated to be 0.23 mmol/(m2·d) at JL7K and 0.16 mmol/(m~2·d) at MA. The community structure analysis showed that calanoid copepods, cnidarians and appendicularians were the main contributors to DVM of mesozooplankton at both sites. We also compared the present result with previous studies of the two ecosystems,and suggested that the DVM of mesozooplankton was more homogeneous within the WPWP and more variable within the NPSG, though both ecosystems showed typically extremely oligotrophic conditions. The different diel vertical migration strength of mesozooplankton between NPSG and WPWP implied different efficiency of carbon pump in these two ecosystems.  相似文献   

13.
Zooplankton biomass, gut fluorescence and electron transfer system (ETS) activity were measured in vertical profiles (0–900 m) in two different size classes (<1 and >1 mm) in Canary Island waters. Both size fractions displayed a typical pattern of distribution with higher biomass, gut fluorescence and ETS in the shallower layers at night. By day, however, the vertical distribution varied between the size fractions, with higher biomass of the small fraction in the 0–200 m and a layer of large organisms at depth (∼500 m). For both size fractions, average ETS activity was higher by day than at night at depths between 200 and 600 m. Similarly, gut fluorescence was slightly higher by day below 200 m. The downward export of respiratory carbon was 1.92 and 4.29 mg C m−2 d−1 for samples obtained southwest of Gran Canaria Island and west of Tenerife Island respectively, being 2.68 mg C m−2 d−1 for the whole area. These values represented 16–45% (22–28% for the area) of the calculated passive particulate export production resulting from primary production. The estimated “gut flux” accounted for 0.35 (western zone) and 2.37 mg C m−2 d−1 (southwest of Gran Canaria), being 1.28 mg C m−2 d−1 for the whole area and represented between 3 and 25% (11–14% for the whole area) of the estimated passive particle export flux. These results agree with previous estimates and suggest that diel-migrant zooplankton can play an important role in the downward flux of carbon.  相似文献   

14.
Visual observations were made in September 1997 during the 39 cruise of R/V “Akademik Mstislav Keldysh” with 2 deep-sea manned submersibles “Mir” aboard. During 4 dives the following plankton countings were made: 3 vertical throughout the water column during the day, 2 vertical in the upper 1000 m at night, and 1 oblique in the plume area during the day. Biomass profiles are represented for each dive for all abundant animal groups: copepods, euphausiids+decapods+mysids, chaetognaths, medusae, ctenophores, siphonophores, cyclothones, myctophides, radiolarians, and the total zooplankton. Plankton distribution shows 2 aggregations, one within the main pycnocline and the other near the plume; Gelatinous animals and radiolarians dominate in both aggregations by biomass and make a significant contribution to the plankton biomass throughout the water column. Oblique counting indicates the presence of aggregations of animals near the upper and lower borders of the plume and biomass depletion within the plume core.  相似文献   

15.
To identify seasonal patterns of change in zooplankton communities, an optical plankton counter (OPC) and microscopic analysis were utilised to characterise zooplankton samples collected from 0 to 150 m and 0 to 500 m in the Oyashio region every one to three months from 2002 to 2007. Based on the OPC measurements, the abundance and biomass of zooplankton peaked in June (0–150 m) or August (150–500 m), depending on the depth stratum. The peak periods of the copepod species that were dominant in terms of abundance and biomass indicated species-specific patterns. Three Neocalanus species (Neocalanus cristatus, Neocalanus flemingeri and Neocalanus plumchrus) exhibited abundance peaks that occurred before their biomass peaks, whereas Eucalanus bungii and Metridia pacifica experienced biomass peaks before their abundance peaks. The abundance peaks corresponded to the recruitment periods of early copepodid stages, whereas the biomass peaks corresponded to the periods when the dominant populations reached the late copepodid stages (C5 or C6). Because the reproduction of Neocalanus spp. occurred in the deep layer (>500 m), their biomass peaks were observed when the major populations reached stage C5 after the abundance peaks of the early copepodid stages. The reproduction of E. bungii and M. pacifica occurred near the surface layer. These species first formed biomass peaks of C6 and later developed abundance peaks of newly recruited early copepodid stages. From the comparison between OPC measurements and microscopic analyses, seasonal changes in zooplankton biomass at depths of 0–150 m were governed primarily by E. bungii and M. pacifica, whereas those at depths of 150–500 m were primarily caused by the three Neocalanus species.  相似文献   

16.
The flux and composition of wax esters and triacylglycerols were measured in particulate matter collected by free-drifting sediment traps in the Peru upwelling area. The purpose was to assess diurnal variations in sources and vertical transport of these lipids. Traps were deployed for periods of 8–12 h during both day and night at the base of the euphotic zone (10–14m) and at about 50m. Although the flux of total particulate organic carbon varied 4×, the flux of wax esters varied by 84× and triacylglycerols by 30×. POC, wax esters, and triacylglycerols also showed different temporal patterns. The highest wax ester flux was measured in shallow traps at night and is attributed to increased inputs from zooplankton which feed near the surface at night. Triacylglycerol flux was high in both shallow and deep nocturnal traps, consistent with inputs of both phytoplankton and zooplankton lipids. The composition of these lipids was also quite variable and attests to the complex nature of the lipid sources.  相似文献   

17.
The biomass, species and chemical composition of the mesozooplankton and their impact on lower food levels were estimated along a transect across the Arctic Ocean. Mesozooplankton biomass in the upper 200 m of the water column was significantly higher (19–42 mg DW m-3) than has previously been reported for the Arctic Ocean, and it reached a maximum at ca. 87°N in the Amundsen Basin. The lowest values were recorded in the Chukchi Sea and Nansen Basin, where ice cover was lower (50–80%) than in the central Arctic Ocean. In the deeper strata (200–500 m) of the Canadian and Eurasian Basins, the biomass was always much lower (4.35–16.44 mg DW m-3). The C/N (g/g) ratio for the mesozooplankton population was high (6.5–8.5) but within the documented range. These high values (when compared to 4.5 at lower latitudes) may be explained by the high lipid content. Mesozooplankton accounted for approximately 40% of the total particulate organic carbon in the upper 100 m of the water column. Mesozooplankton species composition was homogeneous along the transect, consisting mainly of copepods (70–90% of the total number). It was dominated by four large copepod species (Calanus hyperboreus, C. glacialis, C. finmarchicus and Metridia longa), which together accounted for more than 80% of the total biomass. According to measurements of gut pigment and gut turnover rates, the mesozooplankton on average ingested between 6 and 30% of their body carbon per day as phytoplankton. Microzooplankton may have provided an additional source of energy for the mesozooplankton community. These data emphasize the importance of mesozooplankton in the arctic food web and reinforce the idea that the Arctic Ocean should no longer be considered to be a “biological desert”.  相似文献   

18.
The migratory response of intertidal microphytobenthos to changes in irradiance was studied on undisturbed estuarine sediments. Two non-destructive optical techniques were used to trace variations in vivo of surface biomass: PAM fluorometry, for measuring the minimum fluorescence level (Fo); and spectral reflectance analysis, for quantifying the normalized difference vegetation index (NDVI). Following the formation of a dense biofilm at the surface, replicated sediment samples were simultaneously exposed to six different irradiance levels, ranging from 50 to 1500 μmol m−2 s−1, during a period of 120 min. The migratory photoresponse of the biofilms was characterised by constructing biomass vs. light curves (BLC), relating the accumulation of microalgal biomass after that period (estimated by Fo or NDVI) to the irradiance level incident on the surface. BLCs allow characterising the main features of the migratory photoresponse of intact biofilms. Typical BLC showed a clear biphasic pattern, with an increase in microalgal accumulation under irradiances below 100 μmol m−2 s−1, maximum values under 100–250 μmol m−2 s−1, and a gradual decrease of surface biomass under higher irradiances, indicating a strong photophobic downward migratory response. Similar BLC patterns were obtained when measuring Fo or NDVI. The construction of BLCs for biofilms from intertidal sites with distinctive sediment characteristics and diatom taxonomic composition allowed to detected significant differences in the migratory photoresponse. Biofilms from a muddy sediment exhibited considerably larger amplitude in the migratory photoresponse than the biofilms from a sandy mud site, especially under high irradiances. The photophobic migratory response to high light was found to vary among diatom species, particularly in the case of the biofilms from the muddy sediments.  相似文献   

19.
The biomass, abundance, and vertical distribution of micronekton, including enidarians, mysids, euphausiids, decapods, thaliaceans, and fishes, were studied on the basis of samples collected with an 8-m2 opening-closing rectangular midwater trawl (RMT-8, mesh size: 4.5 mm) at three stations in the subarctic Pacific (the western subarctic gyre, the central Subarctic, and the Gulf of Alaska) and one station in the oceanic Bering Sea. The total biomass in the 0–1000 m water column ranged from 2.9 to 5.1 gDW m–2. Except for primary consumers that showed highly variable biomass (thaliaceans and euphausiids), biomass was highest in the oceanic Bering Sea followed by the central (boundary between eastern and western gyres), western gyre, and eastern Gulf of Alaska. The biomass compositions by higher taxa were basically similar between regions: fishes were most dominant, followed by enidarians at all stations, except for the marked predominance of thaliaceans in the Gulf of Alaska. High biomasses of gelatinous animals (31% of overall dry weight), occasionally comparable to those of fishes and crustaceans, suggest their potential importance in the subarctic Pacific. Characteristics in vertical patterns of micronekton biomass common in all stations were: (1) a mesopelagic peak around 500–600 m both day and night, (2) a layer of low biomass in the cold intermediate water and/or in the upper mesopelagic zone, (3) a nighttime shift of biomass to upper layers, and (4) an highly variable biomass of epipelagic/interzonal migrants (euphausiids and thaliaceans).  相似文献   

20.
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