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The Staphylinine group of rove beetle subfamilies is a significant animal radiation, and one subordinate monophyletic clade – the ‘Euaesthetine subgroup’ – includes around 3000 species in subfamilies Euaesthetinae and Steninae and has a fossil record dating to the Early Cretaceous. Detailed morphological study of a new well-preserved Cretaceous Burmese amber fossil revealed strong evidence consistent with its taxonomic placement in the euaesthetine genus Octavius. We thus describe Octavius electrospinosus sp. nov., the first Cretaceous record of the genus and of the tribe Euaesthetini. Previously, the oldest records of Octavius and Euaesthetini were from the Eocene (Baltic amber) and discovery of O. electrospinosus sp. nov. therefore nearly doubles the minimum lineage age of Octavius, increasing it by 50 million years. We also briefly review the known Euaesthetine subgroup fossil record and tabulate summary data for all previously described fossils. All are placed in extant genera, and have visible diagnostic generic-level characters including some putative synapomorphies as judged by recent phylogenetic work. Including O. electrospinosus sp. nov., there are now four known Cretaceous species, all of which belong to either Octavius, Nordenskioldia, or Stenus. To explain the long-term morphological stasis in this group of rove beetles, we suggest that the continuous presence of mesic habitats may have buffered these lineages from strong selection for morphological change. Considering the fossils along with phylogenetic hypotheses we suggest the Euaesthetine subgroup originated in the Late Jurassic– Early Cretaceous and the Staphylinine group in the Early Jurassic. We emphasize the derived status of Cretaceous fossils in assessing possible divergence times and the significance of the pre-Cretaceous taphonomic bias for restricting more robust estimates. Further detailed morphological study of available fossils in a phylogenetic framework is badly needed to clarify the phylogenetic positions of these taxa.  相似文献   

4.
A field study was conducted on two mountain streams in the Cascade Mountains of Washington State on the morphological characterization of cluster microforms. Morphological characterization of clusters is presented in terms of: (i) cluster shape; (ii) cluster geometric properties; and (iii) the spatial arrangement of clusters in the horizontal plane. Clusters were differentiated from other microtopography features such as reticulate structures and transverse ribs, and identified clusters were categorized by shape as being of pebble, line, comet, heap or ring type. The complex spatial arrangement of clusters at the sites was characterized by using a two‐dimensional correlation function, which allowed for measurement of the average cluster‐spacing properties. For the rivers examined, pebble‐shaped clusters were the most frequently observed cluster shape. Cluster geometric properties were found to be controlled by particles of the largest size fraction in the bed and the projected frontal width of the cluster – with cluster length being linearly related to cluster width for cluster width‐to‐height ratios <3·5. Results of the cluster‐spacing analysis suggest that cluster spacing increases with cluster size and decreases with local slope. Application of this principle to the available spacing data shows that cluster spacing λ scales with the ratio of S/d0 such that λS/d0 = constant, where S is the local slope and d0 the diameter of the largest particle in the cluster.  相似文献   

5.
Cupressinocladus Seward is a fossil genus of conifers and conifer fossils with reproductive organs are very rare. In general, it is difficult to understand the natural affinities with other conifers. In this paper, a new species, Cupressinocladus guyangensis P.H. Jin et B.N. Sun sp. nov., is reported based on branches with immature female cones from the Lower Cretaceous Guyang Formation of the Guyang Basin in Inner Mongolia, northern China. The foliage shoots are decussate. Leaves are decussate, imbricate, scale-like, weakly dimorphic, and bear longitudinal glands on the abaxial view. Stomata complexes are haplocheilic, monocyclic, irregularly arranged, and spread along the leaf margin. Immature female cones are subglobose with 6–8 cone scales, and three subglobose ovules arranged in a row at the base of the cone scales. Moreover, we performed cluster analysis using a statistics and machine learning toolbox for 23 fossils and extant species based on 16 morphological characters. The result implies that the new species bears a close resemblance to the extant Cupressus funebris Endl. and might have nearest systematic affinities to it.  相似文献   

6.
The appearance of coelenterates marks the real beginning of metazoan evolution. It therefore has a prominent position in the origin and evolutionary history of organisms, and is also a pivotal question of evolutionary biology. Punctatus is an extinct, ancient marine animal from the early stage of the Cambrian explosion, occurring at the lowermost Cambrian of both Kuanchuanpu (Ningqiang, Shaanxi) and Maidiping (Emei, Sichuan) areas. Punctatus has been studied for many years since the discovery of its fragments. Systematic and phylogenetic analysis has long been limited because of the rarity of complete specimens. In order to improve research into Punctatus, more than ten thousand globular fossils were recovered by means of “Chemistry Retting”. On the basis of the study of these globular fossils, a series of Punctatus fossils with cone parts and finely preserved soft-tissue mouthparts and fossilized metazoan embryo were recovered. Through research on characteristics such as shape, modality and structure of these fossils symbiotic with Punctatus emeiensis, the author found many possible embryo fossils including the evidence of gastrula-stage animal fossils. The sequence of fetation might have appeared on the corolliform oral region of Punctatus emeiensis. A study of the soft tissues, functional morphology and the sequence of embryo fossils shows evidence that Punctatus resembles coelenterate polyps in systematic classification. Perhaps it also represents an ancestor of the coelenterate with an original tentacle. A comparison with the real “tentacle animal” found in the Chengjiang Fauna, shows that the original tentacle is very tiny and its function range is limited. This reveals the primitive nature of the animal. Although the original tentacle is so small, it does exist, representing the first big step towards the real flexible tentacle with a strong function from the early evolving tentacle. Translated from Journal of Acta Palaeontologica Sinica, 2006, 45(2): 182–194 [译自: 古生物学报]  相似文献   

7.
Sediments contained in the river bed do not necessarily contribute to morphological change. The finest part of the sediment mixture often fills the pores between the larger grains and can be removed without causing a drop in bed level. The discrimination between pore‐filling load and bed‐structure load, therefore, is of practical importance for morphological predictions. In this study, a new method is proposed to estimate the cut‐off grain size that forms the boundary between pore‐filling load and bed‐structure load. The method evaluates the pore structure of the river bed geometrically. Only detailed grain‐size distributions of the river bed are required as input to the method. A preliminary validation shows that the calculated porosity and cut‐off size values agree well with experimental data. Application of the new cut‐off size method to the river Rhine demonstrates that the estimated cut‐off size decreases in a downstream direction from about 2 to 0·05 mm, covariant with the downstream fining of bed sediments. Grain size fractions that are pore‐filling load in the upstream part of the river thus gradually become bed‐structure load in the downstream part. The estimated (mass) percentage of pore‐filling load in the river bed ranges from 0% in areas with a unimodal river bed, to about 22% in reaches with a bimodal sand‐gravel bed. The estimated bed porosity varies between 0·15 and 0·35, which is considerably less than the often‐used standard value of 0·40. The predicted cut‐off size between pore‐filling load and bed‐structure load (Dc,p) is fundamentally different from the cut‐off size between wash‐load and bed‐material load (Dc,w), irrespective of the method used to determine Dc,p or Dc,w. Dc,w values are in the order of 10?1 mm and mainly dependent on the flow characteristics, whereas Dc,p values are generally much larger (about 100 mm in gravel‐bed rivers) and dependent on the bed composition. Knowledge of Dc,w is important for the prediction of the total sediment transport in a river (including suspended fines that do not interact with the bed), whereas knowledge of Dc,p helps to improve morphological predictions, especially if spatial variations in Dc,p are taken into account. An alternative to using a spatially variable value of Dc,p in morphological models is to use a spatially variable bed porosity, which can also be predicted with the new method. In addition to the morphological benefits, the new method also has sedimentological applications. The possibility to determine quickly whether a sediment mixture is clast‐supported or matrix‐supported may help to better understand downstream fining trends, sediment entrainment thresholds and variations in hydraulic conductivity.  相似文献   

8.
《第四纪科学杂志》2017,32(5):653-660
Our knowledge of past animal populations, including the geographical ranges of extinct species, has largely been derived from morphological analyses of skeletal fossil remains. However, a major barrier to the identification of the remains of extinct megafaunal species in archaeological and palaeontological sites is the highly fragmented nature of the material, which often precludes confident taxonomic identifications based on morphology. Biomolecular techniques are able to go beyond these limitations and are increasingly being used to make such identifications. Protein analysis offers a promising alternative to DNA techniques because they can be much cheaper, more amenable to high‐throughput processing and work on much older specimens. Here we demonstrate the potential of collagen fingerprinting in an Australian context by extracting collagen from 50‐ka kangaroo fossils from two caves in Tasmania, and identify several species including the extinct short‐faced kangaroo Simosthenurus occidentalis . Importantly, of the five fossil bones sampled that had hitherto been ascribed morphology‐based identifications below the family level, three had been incorrectly identified during an initial assessment of photographs taken in the field. Our results highlight the utility of using protein‐based methods for making genus‐level identification of marsupial bone, especially those that may form a basis for broader arguments such as that of late‐surviving megafaunal species.
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9.
3D finite strain analyses and kinematic vorticity measurements were carried out on the Loghon Anticline within the HP‐LT Sanandaj–Sirjan metamorphic belt (Neyriz area, SW Iran). Rƒ/φ and Fry methods were used on the strain markers (e.g. deformed fossils) to interpret geometric relationships between the fold axis, strain ellipsoid axes and shear zone boundaries. The results indicate the predominance of prolate strain in the anticline. Quantitative kinematic analyses show that the Wk parameter is 0. 67 ± 0. 06 (i.e. pure‐shear dominated non‐coaxial flow). This study quantitatively supports the establishment of a dextral transpressive system, which is responsible for the development of the large‐scale right‐lateral shear zones that strike sub‐parallel to the major folds. Flexural shear combined with regional dextral‐shear is suggested to be the most common mechanism of folding in this area. Copyright © 2011 John Wiley & Sons, Ltd.  相似文献   

10.
A biostratigraphic study on calcareous nannofossils from the CM3D06 Co-rich ferromanganese crust from the Magellan seamounts in the northwestern Pacific enabled estimation of depositional age.The bio-imprinting of calcareous nannofossils and other fossil species suggests six age ranges for the nannofossils: late Cretaceous,late Paleocene,(early,middle,late) Eocene,middle Miocene,late Pliocene,and Pleistocene.Gas chromatography(GC) and gas chromatography-mass spectrometry(GC-MS) were used to test the Co-rich crusts,and a variety of molecular fossils were detected,such as chloroform bituminous "A",n-alkane,isoprenoid and sterol.Peak carbon and molecular indices(such as ΣC23 /ΣC24 +,CPI,Pr/Ph,Pr/nC17,Ph/nC18 and δ13C) indicate that the parent organic matter is dominated by marine phytoplankton and thallogen whereas there is little input of terrestrial organic matter.Researches on calcareous nannofossils,molecular fossils and molecular organic geochemistry data reveal that the Paleocene/Eocene(P/E) global event is recorded in the cobaltrich crusts from the northwestern Pacific Ocean.A succession of biomes can be observed near the 85 mm boundary(about 55 Ma),i.e.,the disappearance of the late Cretaceous Watznaueria barnesae and Zigodicus spiralis,and Broisonia parka microbiotas above the P/E boundary,and the bloom of Coccolithus formosus,Discoaster multiradiatus,Discoaster mohleri and Discoaster sp.below the boundary.Typical parameters of molecular fossils,such as saturated hydrocarbon components and carbon-number maxima,Pr/Ph,Pr/C17,Ph/C18,distribution types of sterols,Ts/Tm ratios and bacterial hopane,also exhibit dramatic changes near the P/E boundary.These integrated results illustrate that the biome succession of calcareous nannofossils,relative content of molecular fossils and molecular indices in the cobalt-rich crusts near the 85 mm boundary faithfully record the P/E global event.  相似文献   

11.
Carbonate drifts have so far not been as intensely investigated as their siliciclastic equivalents, especially from an ichnological perspective. The aim of this work is therefore to provide an overview of the different bioturbation styles in carbonate drifts for ichnologists and sedimentologists working in such deposits. Different types of carbonate drifts from the Maldives were studied to address this objective. The cores recovered during International Ocean Discovery Program Expedition 359 were examined to provide the sedimentological and ichnological data for a detailed analysis of the ichnology of carbonate drifts. The ichnological characteristics of the Maldives drifts are compared to other carbonate drifts in order to discuss similarities and differences, and thus provide an overview of the general characteristics of carbonate drift ichnology. These drifts are located in the Santaren Channel which lines Great Bahama Bank, along the Marion Plateau in Australia, in the Limassol and Larnaca basins in Cyprus and in the Danish Basin in Denmark. The common characteristics of bioturbation in carbonate drifts are: (i) the complete bioturbation of the sediment with bioturbation indexes between four and six; (ii) the occurrence of distinctive trace fossils limited to facies contacts or condensed intervals; (iii) a typical ichnoassemblage consisting of Thalassinoides, Scolicia, Planolites, Zoophycos, Chondrites, Phycosiphon and Palaeophycus; (iv) the contiguous occurrence of ichnogenera from different tiers, with only Zoophycos and Chondrites as deep tiers; and (v) distinct infills of the traces including particulate organic matter, pyrite, silica and celestine. In addition, the main ichnofacies of carbonate drifts is the Zoophycos ichnofacies. Ichnofabrics grade from coarse‐grained and completely bioturbated to ichnofabrics with present to rare trace fossils and preserved sedimentary structures. The type and intensity of the bioturbation is controlled by the amount of organic matter and the oxygenation at the sea floor that is determined by the action of bottom currents and the sea‐level fluctuations affecting the carbonate factory in carbonate platforms bordering the basins where the carbonate drifts form. The study of the bioturbation in core and outcrop provides palaeoenvironmental information about carbonate‐drift deposits that complement the classical sedimentological data.  相似文献   

12.
Halkieria is an Early Cambrian fossil genus with proposed phylogenetic affinities ranging from stem group lophotrochozoan to stem-group aculiferan. Skeletal similarities are often cited as evidence in support of a mollusk interpretation for Halkieria and related Cambrian fossils. The skeletal microstructures of the polyplacophoran mollusk (chiton) Ischnochiton hakodadensis, which bears perinotum elements resembling sclerites of halkieriids and other coeloscleritophoran small shelly fossils (SSFs), were studied to evaluate the interpretations. There are both similarities and differences between chiton skeletal structures and halkieriid sclerites: 1) the head valve of L hakodadensis and the dorsal shell of Halkieria evangelista share a similar pattern of fine concentric growth lines, but halkieriid dorsal shells lack other polyplacophoran valve characteristics, including V-shaped indentation, radiate ribs; 2) the arrangement of I. hakodadensis perinotum elements in three zones is somewhat similar to the arrangement pattern of halkieriid sclerites; 3) the inner perinotum scales of L hakodadensis (-200 μm in diameter) have a base and a blade, remarkably similar in morphology to halkieriid palmate sclerites (-400 μm in diameter). However, I. hakodadensis perinotum scales are nearly solid whereas halkieriid sclerites are hollow. These observations add a microstructural dimension to the skeletal comparison between mollusks and halkeriids, and may prove to be useful in resolving the phylogenetic affinity of Halkieria and the Halwaxiidae.  相似文献   

13.
We describe a large collection of fossil ‘waterfowl’ bones that are referable to the extinct clade Presbyornithidae (Anseriformes). All of these fossils were collected between 1971 and 1994 from Upper Paleocene and Lower Eocene sediments at the Tsagaan Khushuu site in the Gobi Desert of southern Mongolia. The collection includes specimens referred to a new small species within the genus Presbyornis Wetmore, 1926 as well as large numbers of bones that we place in the genus Presbyornis. On this basis of the Tsagaan Khushuu collection we suggest that several species of Presbyornis likely coexisted in this region; indeed, the presence of large numbers of middle‐sized, morphologically consistent but probably ecologically disparate species at the Tsagaan Khushuu site is consistent with the range of variation seen, for example, in taxa of extant dabbling ducks (Anatini). Although the anatomy and phylogenetic position of Presbyornithidae (in particular Presbyornis) are well known, this material from Mongolia further demonstrates the prevalence of these birds in aquatic and semi‐aquatic habitats by the earliest Paleogene. Because presbyornithids are also well documented from the late Cretaceous, their palaeoecology and morphological diversity provides a clue to selective avian survivorship across the Cretaceous‐Paleogene (K‐Pg) boundary. Copyright © 2009 John Wiley & Sons, Ltd.  相似文献   

14.
Macroscopic plant remains, pollen, insect and mollusc fossils recovered from a cut bank on the Red River in North Dakota, USA, provide evidence that an extensive wetland occupied the southern basin of Lake Agassiz from 10 230 to 9900 14C yr BP. Marsh‐dwelling plants and invertebrates had colonised the surface of a prograding delta during the low‐water Moorhead Phase of Lake Agassiz. A species of Salix (willow) was abundant along distributary channels, and stands of Populus tremuloides (aspen), Ulmus sp. (elm), Betula sp. (birch), and Picea sp. (spruce) grew on the better‐drained sand bars and beach ridges. Most of the species of plants, insects, and molluscs represented as fossils are within their existing geographic ranges. Based on a few species with more northerly distributions, mean summer temperature may have been about 1–2°C lower than the present day. No change in species composition occurred in the transition from the Younger Dryas to Preboreal. At the time that the wetland existed, Lake Agassiz was draining either eastward to the North Atlantic Ocean or northwestward to the Arctic Ocean. The wetland was drowned during the Emerson Phase transgression that resulted in meltwater draining southward to the Gulf of Mexico after 9900 14C yr BP. Copyright © 2005 John Wiley & Sons, Ltd.  相似文献   

15.
成吉思汗驿站龙江组剖面位于扎兰屯成吉思汗镇东南部,组内大化石稀少,但孢粉化石丰富,下部岩性为沉凝灰岩、凝灰岩、凝灰质砂岩、砂岩、砾岩,上部岩性为安山岩、英安岩。通过对成吉思汗驿站龙江组孢粉化石的分析,在龙江组中识别出圆形粒面孢-二连粉孢粉组合带(Cyclogranisporites-Erlianpollis)。通过对孢粉化石属种的分析,根据具有时代指示意义的孢粉化石,如Erlianpollis,Cicatricosisporites,Pilosisporites等,认为成吉思汗地区龙江组孢粉化石形成时代为早白垩世,结合地质、古植物及孢粉化石的定量分析,认为龙江组植被类型反映亚热带气候,指示温暖潮湿的环境。  相似文献   

16.
The rich Quaternary fossil record from the Cantabrian region (NW Iberia) is virtually restricted to the Late Pleistocene, with Middle Pleistocene findings very rare. This work presents the study of two Middle Pleistocene palaeontological sites in the Cantabrian region: Llantrales quarry and Mestas de Con mine, both discovered and first described in the 1950s but never studied in detail thereafter. The material was here re‐analysed and re‐evaluated. Taxonomic attributions have been changed in most cases and, consequently, chronological assignments of the fossils have also been changed. This first detailed study of the assemblage from Llantrales quarry revealed the occurrence of the large‐sized deer Praemegaceros solilhacus, which has never been reported previously in North Iberia. Cervus cf. elaphus and Stephanorhinus cf. hundsheimensis were also identified. The age of these fossils was originally regarded as Pontian (late Miocene), but new palaeontological results indicate a much younger age, most likely from around 0.8 to 0.5 Ma. A taxonomic re‐evaluation of the fossils from Mestas de Con provided different results with respect to the 1950s works. The faunal association is composed of a large‐sized deer (probably corresponding to Praemegaceros), Cervus elaphus, Capreolus cf. capreolus, Bison sp. (small sized), Equus sp. (large sized), Stephanorhinus hemitoechus, Ursus sp. and Homotherim latidens. This faunal association was evaluated within a western European context and proves to be highly analogous to other contemporary assemblages there. The chronology was initially regarded as late Villafranchian (in a broader sense). New results, documenting the co‐occurrence of Stephanorhinus hemitoechus and Homotherium latidens, indicate a more precise and younger age, most likely between 0.6 and 0.4 Ma. These two fossil assemblages provide new and relevant information on the Middle Pleistocene faunas from a geographical region where this information, compared with other western European areas, is rare and poorly known.  相似文献   

17.
The Aiketik Group, distributed at the western end of the South Tianshan Mountains, China, is an important lithostratigraphic unit involved in the South Tianshan orogen. It is separated from the adjacent rocks by faults. Generally, the geologists ascribed it to the Upper Carboniferous according to Pseudostaffella sp., Profusulinella sp. and Fusulinella sp. found from the limestone and sandy limestone of Aiketik. Our radiolarian fossils were obtained from the chert samples collected from the Haladaok section located at the upper Tuoshihan River. The fossils mainly include Albaillella undulata Deflandre, Albaillella paradoxa Deflandre, Albaillella sp. aff. A. paradoxa Deflandre, Albaillella sp. cf. A. deflandrei Gourmelon, Albaillella sp., Albaillella excelsa Ishiga, Kito and Imoto (?), Belowea variabilis (Ormiston et Lane), Callella cf. C. parvispinosa Won, Entactinia cf. E. tortispina Ormiston et Lane, Entactinia aff. E. tortispina Ormiston et Lane, Entactinia variospina Won, Entactinia sp., Eostylodicty  相似文献   

18.
Changes in the ostracod assemblages from two sediment cores collected from Lago Cardiel in southeastern Patagonia (49°S) reflect the main regional abrupt climatic changes over the last 15.6 cal. ka BP. Shifts in species abundance and switches in dominances suggest that these were mainly driven by variable salinity. During the Late Pleistocene, Limnocythere rionegroensis was abundant and dominant, indicating waters with high salinity and prevalence of evaporative processes. Between 12.6 and 10.8 cal. ka BP, Lago Cardiel expanded markedly and reached an Early Holocene highstand of +55 m above present lake level. A major change in ostracod assemblage in which Limnocythere patagonica appears as the dominant species in parallel with the disappearance of L. rionegroensis and Eucypris aff. cecryphalium mirrored this transitional period between the cold and dry Late Pleistocene and the humid and warm Early Holocene. Over the last 4 cal. ka BP, L. rionegroensis returned to the species assemblage and Riocypris whatleyi increased its abundance pointing towards increasing salinities. The variations in size, shape and ornamentation of L. rionegroensis and R. whatleyi fossil valves were examined using geometric morphometric techniques and further compared to those of modern Patagonian sites. Limnocythere rionegroensis specimens displayed high morphological variability during the evolution of Lago Cardiel. More specifically, the switch in reproductive mode – from sexual to parthenogenetic– and the increase in valve ornamentation around 12.7 cal. ka BP suggest that these changes were promoted by the hydrological alteration that occurred in the Late Pleistocene. This exercise provides a robust range of morphological variation for these proxies, which will be useful in further taxonomic and palaeoenvironmental studies adding more information about different factors influencing the observed morphological trends.  相似文献   

19.
An electron probe and chemical study of bulk phosphorite samples and separated constituents from various Negev deposits was carried out together with XRD, FTIR spectroscopy and textural analysis. The results allow a better understanding of the distribution of fluorine in these Upper Cretaceous phosphorite sequences and shed light on variations in the composition of the carbonate fluorapatite (CFA) phase during phosphogenesis. Two facies are recognized: (1) a pristine, microbially generated phosphorite facies; (2) a recycled, peloidal and biodetrital facies. Fluorine distribution in the Negev phosphorites is facies controlled: F/P2O5 is much lower in the pristine facies (0·090–0·107) than in the recycled facies (0·107–0·120). In addition, F/P2O5 varies considerably between the various constituents of the phosphate fraction; F‐poor francolites (F/P2O5 as low as 0·080) co‐exist with F‐rich francolites (F/P2O5 as high as 0·135) in the same phosphorite bulk sample. A lower F/P2O5 in francolite is associated with higher Cd and Zn concentrations in the phosphorite, an increase in Fe‐rich smectites in the clay fraction and the presence of structural OH in the francolite. The lower F/P2O5 ratios in the pristine facies are attributed to high organic deposition rates during the formation of these matted sediments, leading to rapid burial of the in situ‐forming CFA. This is possibly coupled with diffusion of F from sea water into bottom sediments being hampered by microbial mat coatings. These conditions resulted in O2‐depleted porefluids, inducing the precipitation of Cd‐rich Zn sulphides and the formation of Fe‐rich smectites. F‐enrichment probably takes place when the earlier formed F‐poor ‘primary’ CFA is relocated close to the sea floor and bathed with interstitial sea water solutions of higher F concentrations. Oxidation and removal of the sulphide‐bound Cd and Zn apparently occurred together with enrichment in F of the francolite. Combining chemical data with XRD and FTIR results suggests a multistage growth for the Negev phosphate constituents in shifting formational sites and porefluids of varying F concentrations. This multiphase growth is reflected in the patchy distribution of F in the Negev constituents and might explain the inverse correlation between mean CO2/F and F/P2O5 ratios of the analysed phosphorites in the two facies. It also suggests that CFA (or an amorphous precursor) initially formed with some OH groups in the apatite structure, which were subsequently substituted by F ions in recycled francolite through re‐equilibration with porefluids of higher F concentrations.  相似文献   

20.
The Yixian Formation (Lower Cretaceous) of China is famous for its fossils of early angiosperms, and these fossils have shed otherwise unavailable light on the evolution of angiosperms. The seeds in Archaefructus and Nothodichocarpum are inserted along the dorsal of the fruits while those in Sinocarpus are along the ventral of the fruits, suggesting different Bau‐plans and pathways deriving the carpels in these plants of the Early Cretaceous. Adding further fossil evidence, here we report a novel fossil angiosperm, Neofructus lingyuanensis gen. et sp. nov, with its seeds inserted on both ventral and dorsal sides of the fruits documented through light microscopic and SEM observations, suggesting a novel way forming gynoecium in the Yixian Formation. These fossil angiosperms indicate that angiosperm gynoecia in the Yixian Formation (the Barremian to Aptian) are derived in obviously different ways, implying an evolutionary scenario for angiosperms quite different from formerly assumed.  相似文献   

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