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1.
Baurusuchids are among the most common and diverse crocodyliform fossils from the Late Cretaceous Bauru Basin of Brazil. This inland continental basin was the habitat of a rich crocodyliform fauna containing five mesoeucrocodylian families, of which the Baurusuchidae represents highly specialized predatory crocodyliforms of terrestrial habits as indicated by their dental, cranial, and postcranial features. The large size they achieved, together with likely predatory adaptations, would suggest they competed and occupied theropod ecological niches in the Bauru Basin. Here we describe Gondwanasuchus scabrosus gen. et sp. nov., a medium-sized baurusuchid with a strongly laterally compressed skull, bearing unique dentition with deep apicobasal sulci and probably well-developed binocular vision. The cranial and dental features in Gondwanasuchus suggest that this active predator would have fed on small vertebrates and took the role of small theropods in terrestrial guild. Gondwanasuchus is the most distinctive baurusuchid known to date and enriches the knowledge on these important Gondwanan terrestrial predatory crocodyliforms.  相似文献   

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3.
The skin of Crocodyliformes is characterized by osteoderms of various sizes and shapes. It plays roles in defence, thermoregulation, sexual attraction, calcium reserves, and locomotion. This study presents the morphometric characteristics of osteoderms preserved in the nuchal, dorsal, ventral and appendicular shield of Montealtosuchus arrudacamposi, a Peirosauridae crocodyliform from the Turonian-Santonian (Upper Cretaceous) of Adamantina Formation, Bauru Basin, Brazil. The results of the analysis show that its dermal shield had protective and thermoregulatory functions and a well-defined mechanical function allowing terrestrial locomotion and enabling good agility and capacity for movement.  相似文献   

4.
Upper Cretaceous vertebrate accumulations from the Adamantina Formation are known due to their high taxonomic diversity. On the other hand, taphonomic analyses still are rare, limiting the understanding of processes related to the biostratinomic and fossildiagenetic histories of this lithostratigraphic unit. In 2005, fossils were collected from an outcrop located at Jales municipality, state of São Paulo, southeastern Brazil. From this outcrop, a well-preserved Baurusuchus was recovered, which displays a peculiar set of taphonomic signatures. This paper identifies and interprets taphonomic features of a specimen of Baurusuchus (Crocodyliformes, Baurusuchidae; UFRJ DG 418-R) from the Adamantina Formation (Upper Cretaceous of the Bauru Basin), in Jales municipality, state of São Paulo. Brief taphonomic comparisons with other specimens previously studied (crocodiles and dinosaurs) and a lithofaciological analysis of the outcrop were undertaken in order to enhance the knowledge of the stratigraphy and paleoenvironment and improve the time resolution for the Adamantina Formation in the region of Jales. Furthermore, paleoecological data were interpreted based on the taphonomic analysis.The fossil is composed of an articulated segment of vertebral column, ribs, part of the pelvic girdle and gastralia. There is no hydraulic equivalence between both the specimen of Baurusuchus and the size of quartz grain predominant in the fossiliferous layer, suggesting death in situ or short transport as a “water carcass”. Teeth marks identified on the pubes were assigned to a small/juvenile baurusuchid crocodyliform or a theropod dinosaur. The repositioning of some elements (ribs and dorsal osteoderms) is suggestive of mummification. Desiccation marks were observed and attributed to the stage 1 of weathering. These features suggest subaerial exposure of the carcass prior to burial, however, probably after the mummification. On the other hand, the subaerial exposure was short, because the individual was not fully disarticulated. Furthermore, the degrees of articulation and preservation of the specimen nullify the hypothesis of reworking.Lithofaciological and taphonomic analyses suggest a well-drained floodplain as the burial environment and an arid or semi-arid climate in the region of Jales outcrop. In addition, the taphonomic signatures seem to indicate a time resolution about 100–104 years for the layer where the crocodyliforms were collected, but it seems to have, within this time limit, temporal-mixing among terrestrial crocodiles and dinosaurs collected from the same layer, suggesting time-averaging in this outcrop.  相似文献   

5.
A small to medium-sized pterodactyloid pterosaur (wingspan approximately 1.10 m) from the Upper Jurassic (middle-late Tithonian) marine deposits of the Vaca Muerta Formation of Patagonia (Los Catutos area, central Neuquén Province, Argentina) is reported. The specimen lacks the skull but constitutes a nearly complete postcranial skeleton, which includes cervical and dorsal vertebrae; a few thoracic ribs; both pectoral girdles; the left pelvic girdle; a proximal right wing (humerus, ulna, and radius) and metacarpal IV; a left wing that lacks only wing phalanx four; and both hindlimbs, the right one without the foot. Ontogenetic features suggest that the new fossil corresponds to a relatively mature individual, probably a subadult. Observed characters support its assignment to the Archaeopteroactyloidea, a basal clade within the Pterodactyloidea. This specimen is the second pterosaur from Los Catutos and the most complete Jurassic pterosaur so far known from South America.  相似文献   

6.
Mongolian Paleontological Center (MPC) 100/1305 is one of the most complete ankylosaurid skeletons ever collected, and includes much of the postcranial skeleton and numerous in situ osteoderms. This specimen has been referred to Saichania chulsanensis, based on the similarity of the skull compared to the holotype of Saichania, MPC 100/151. However, MPC 100/1305 does not include a skull, and so referral of MPC 100/1305 to Saichania must be based on postcranial characters. Comparison of the postcrania of MPC 100/1305 and MPC 100/151 reveals several differences in the scapula, humerus, and metacarpals, indicating that MPC 100/1305 should not be referred to Saichania. Additionally, although it was previously reported that MPC 100/1305 was collected from the Baruungoyot Formation at Khulsan in Mongolia, collection records indicate that this specimen was instead collected from the Djadokhta Formation at Zamyn Khond. Two ankylosaurid species are known from the Djadokhta Formation of Mongolia and China, Pinacosaurus grangeri and P. mephistocephalus. There are no diagnostic characters in MPC 100/1305 that can be used to refer this specimen to P. grangeri or P. mephistocephalus, and there are a few differences between MPC 100/1305 and Pinacosaurus, such as the number of caudal vertebrae, and morphology of the coracoid, which have uncertain taxonomic implications. At present, MPC 100/1305 is best referred to Ankylosauridae indet., or cf. Pinacosaurus, based on its generally congruent morphology with Pinacosaurus and its provenance from the Djadokhta Formation, in which Pinacosaurus is the only recognized ankylosaurid taxon.  相似文献   

7.
Among titanosaurs, osteoderms are morphologically diverse and have been reported from deposits in South America, Africa, Madagascar, Eurasia and Oceania. In Brazil, titanosaur osteoderms are rare and have only been recorded from Bauru Basin sedimentary rocks. Here, we describe a keeled titanosaur osteoderm, which is the first occurrence of an osteoderm specimen in the São Luís Basin, Northeastern Brazil. This osteoderm is characterized by an external bulb, an internal root, and a very rugose cingulum limiting the external and internal surfaces. These characteristics are typical of Titanosauria dermal bones, and this specimen strengthens the idea that the armored sauropods were present during the early Late Cretaceous of Northeastern Brazil, extending their distribution in the northernmost portion of South America.  相似文献   

8.
The postcranial skeleton of a new specimen of the long-tailed lizard Dalinghosaurus longidigitus was excavated from the Yixian Formation in Lingyuan, western Liaoning. The new specimen provides more anatomical information about this species, especially about the anterior dorsal vertebrae, shoulder girdle and forelimbs. This lizard can be included within the clade Scleroglossa by its 27 or more presacrals, moderately long pubis, and gently notched distal end of tibia. But the detailed systematic position for this taxon remains undetermined. The features of the much longer hind limbs and pes compared with forelimbs and manus, metatarsal Ⅳ longer than Ⅲ, pedal phalanges robust, and penultimate phalanx not longer than other phalanges etc. suggest that this lizard was a running and ground swelling animal.  相似文献   

9.
Combined methods of fission track (FTM) and U–Pb in situ zircon dating were applied to sedimentary samples from the Vale do Rio do Peixe Formation, Bauru Basin, Brazil. Detrital zircons of nine samples were determined by the FTM, and the obtained ages varied from 239 Ma–825 Ma, which can be grouped into four main populations as the 230–300 Ma, 460–490 Ma, 500–650 Ma and 696–825 Ma groups. The U–Pb data show two clear source areas: the Early Paleozoic to Neoproterozoic zircons, ranging from 445 ± 14 to 708 ± 18 Ma, and the Paleoproterozoic zircons, ranging from 1879 ± 23 to 2085 ± 27 Ma. Subordinate occurrences of Early Neoproterozoic to Mesoproterozoic zircons (836 ± 15 and 1780 ± 38 Ma) were identified. The combined information allows us to characterize Early Brazilian, Brazilian and Rhyacian material as the main source for the zircons, which are areas situated to west of the Bauru Basin (e.g., Goiás Massif) that have been incorporated into the sedimentary cycles in the Phanerozoic (mainly in the Paraná Basin). FT zircon ages reflect the main denudation processes of the South American Plate from Neoproterozoic to Early Triassic as those related to orogenic cycles of Early Brazilian, Brazilian, Famatinian/Cuyanian and Gondwanide.  相似文献   

10.
Cervical vertebrae of elasmosaurids (Sauropterygia, Plesiosauroidea) have proven to be informative even at family level. The present paper uses a previously developed methodology based on bivariate graphic analysis that evaluates the proportion indices of cervical vertebrae in elasmosaurids, which is applied to a well-preserved postcranial specimen of this group recovered from upper Maastrichtian beds of central Chile. The plotting of such indices groups the Chilean material with other specimens from the Late Cretaceous of Argentina and Antarctica, the latter referred to the genus Aristonectes. This result allows us to identify the first relatively complete individual of the genus based on non-cranial characters. The identification of the Chilean specimen as Aristonectes sp. provides important new data on the postcranial skeleton of the genus, that has remained poorly known until now.  相似文献   

11.
A new species of Macrocnemus is described on the basis of two incomplete specimens from the Lower Ladinian Prosanto Formation of southeastern Switzerland. The new form can be distinguished by its gracile limb elements and having a noticeably longer tibia than either Macrocnemus bassanii or Macrocnemus fuyuanensis. One of the new specimens exhibits soft part preservation in the region of the pelvic girdle. The hind part of this individual is preserved fully articulated and allows a complete count of the caudal vertebrae to be made for the first time with 52 caudals. It is postulated that this specimen was predated upon and that the anterior part of the animal was consumed by a large predator such as a nothosaurid reptile or the actinopterygian fish Saurichthys.  相似文献   

12.
The Sphagesauridae is a family of Crocodyliformes exclusively known for the Brazilian Late Cretaceous Bauru Basin. This lineage reveals how diverse was the morphology and ecology of terrestrial Crocodyliformes during the Late Cretaceous of Gondwana. Here is described Armadillosuchus arrudai gen. et sp. nov., a sphagesaurid that presents some mammal-like morphological features, such as propalinal and alternate unilateral jaw occlusion pattern and heavy body armor, composed of a rigid shield and mobile-banded section as in extant armadillos (Xenarthra, Dasypodidae). These unusual morphological features contrast to the double row of osteoderms observed on the closest relatives of A. arrudai. As its mammal analogs, A. arrudai presents some evidence of fossoriality and an exclusive terrestrial life style in contrast to the extant alligatorids and crocodylids.  相似文献   

13.
A new genus and species of an Aptian–Albian sauropod from the Itapecuru Formation, northern Brazil, Amazonsaurus maranhensis, is described. It is known from an incomplete, but diagnostic postcranial skeleton. The new taxon possesses several autapomorphies, such as the anterior caudal vertebrae with lateral laminae formed by the coalescence of the spinoprezygapophyseal and postzygodiapophyseal laminae and, to a lesser extent, of the postzygodiapophyseal laminae. It exhibits many synapomorphies supporting its inclusion in the Diplodocoidea. These include high caudal neural arches and anterior caudal neural arches with spinoprezygapophyseal laminae on the lateral aspect of the neural spine. This record is consistent with previous hypotheses on the existence of a community of Afro-South American dinosaurs.  相似文献   

14.
Yabeinosaurus was the first lizard genus described from the Early Cretaceous Jehol Biota of China. The holotype of the type species, Y. tenuis, is an immature skeleton but it has been lost for decades. A second species, Y. youngi, was erected based on another immature skeleton and is distinguished by its longer, more gracile limbs. In 2001, a juvenile skeleton from the Jingangshan locality of Liaoning was designated as the neotype for Y. tenuis. Subsequently, several further specimens from other localities, juvenile and adult, have been attributed to this species. Here we describe a second lizard specimen from the neotype locality. In anterior tooth shape, finger-like mandibular angular process, cranial sculpture, maxilla shape, and short, robust humerus, the new specimen resembles adult and subadult material currently attributed to Y. tenuis, but it differs in having bicuspid posterior teeth and a straight rather than a hooked angular process of the mandible. We propose a new species (Y. bicuspidens sp. nov.) for this specimen which represents the first occurrence of bicuspid teeth in an Early Cretaceous lizard from China. However, the recognition of a second species at the neotype locality raises a taxonomic problem. Due to the immaturity of the designated holotype and neotype of Y. tenuis, they cannot be coded for the jaw and dental characters that distinguish the two robust-limbed species. This renders Yabeinosaurus tenuis a nomen dubium. Here we propose that Yabeinosaurus tenuis should be treated as a historic taxon, permitting retention of the generic name under ICZN rules, and erect a second new species (Y. robustus sp. nov.) for the principal robust-limbed morphotype in the Jehol Biota with monocuspid posterior teeth and a hooked angular process.  相似文献   

15.
A new ornithischian dinosaur, Yueosaurus tiantaiensis gen. et sp. nov., is erected based on a partial postcranial skeleton from the Liangtoutang Formation (Aptian–Cenomanian) of Zhejiang Province, China. It differs from other ornithischians in possessing the following unique combination of characters: prominent and slightly ventrally directed cervical parapophyses; anteroposteriorly narrow neural spines on caudal vertebrae; scapula with a ventroscapular groove, supraglenoidal buttress, supraglenoid fossa, and a strong anteroventral expansion of the scapular blade. Yueosaurus represents the first basal ornithopod taxon from southeastern China. It expands our understanding of the Cretaceous dinosaurian fauna of Zhejiang Province.  相似文献   

16.
We report new theropod dinosaur material from the Presidente Prudente Formation (Campanian–Maastrichtian), Bauru Group, in southwestern São Paulo state. The material comprises a fragment of right maxilla of Carcharodontosauridae, an anterior portion of a left ilium of Abelisauroidea and a proximal portion of a right fibula of a coelurosaurian. Previous theropod records from the Bauru Basin comprise Abelisauridae and Tetanurae and in São Paulo state these have been represented by only a right premaxilla of an abelisaurid and isolated abelisaurid and carcharodontosaurid teeth. The new material reported here represents the first theropod remains from the Presidente Prudente Formation, and includes the first abelisauroid and coelurosaurian postcranial remains from the Bauru Basin in São Paulo state and the first latest Cretaceous carcharodontosaurid known from non-dental remains anywhere.  相似文献   

17.
The holotype of Libonectes atlasense is an almost complete skeleton from Upper Cretaceous (mid-Turonian) rocks of the Goulmima area in eastern Morocco. Initial assessment of this specimen in 2005 proposed generic referral based on stratigraphical contemporaneity with Libonectes morgani from the Cenomanian–Turonian of Texas, U.S.A. Nevertheless, relative differences in the profile of the premaxillary-maxillary tooth row, position of the external bony nasal opening, number of teeth and rostrad inclination of the mandibular symphysis, proportions of the axial neural arch, and number of cervical and pectoral vertebrae were used to distinguish between these species. As part of an on-going comparative appraisal of elasmosaurid plesiosaurian osteo-anatomy, we re-examined the type and formally referred material of both L. atlasense and L. morgani in order to establish species validity, as well as compile a comparative atlas for use in future works. Our inspections revealed that these reportedly distinct species-level fossils are in fact virtually indistinguishable in gross morphology. Indeed, the only substantial difference occurs in relative prominence of the midline keel along the mandibular symphysis, which might be explained by intraspecific variation. Furthermore, our observations permit an amendment to the published generic diagnosis of Libonectes with the confirmation of important states such as the likely presence of a pectoral bar, distocaudad expansion of the humerus, and an epipodial foramen. In addition, novel features include a prominent ‘prong-like’ ventral midline process on the coracoids, and the development of a median pelvic bar that encloses a central fenestration. The composite remains of L. morgani thus constitute one of the most complete elasmosaurid skeletal hypodigms documented worldwide, and evidence a trans-Atlantic distribution for this apparently dispersive species during the early–Late Cretaceous.  相似文献   

18.
Among extant vertebrates, only birds have a respiratory system associated with pneumatic diverticula. However, several extinct clades also had pneumatic diverticula, including the sauropod dinosaurs. Among sauropods, Saltasaurini are characterized by extreme postcranial pneumaticity, which extends as far down the spinal column as the posterior caudal vertebrae. In this paper the pneumatic foramina in both the neural arches and the centra are described in detail, and the relative proportion of air spaces in the caudal vertebrae is established, revealing that the skeleton of Rocasaurus muniozi was more pneumatized than that of Neuquensaurus australis, with Saltasaurus loricatus intermediate. The level of pneumatization varies between the three saltasaurine taxa: in Neuquensaurus, only the neural arch is pneumatized, whereas in the other two saltasaurines both neural arches and centra are pneumatized. This allows us to hypothesize that the timing of pneumatization varied between the three species, with Rocasaurus muniozi pneumatized earliest in ontogeny. This ontogenetic pattern is correlated with evolutionary derivation: the most derived taxa show pneumatization in both the neural arch and the centrum.  相似文献   

19.
Saltasaurine titanosaurs are characterized by their relatively small size compared to other sauropods, extreme postcranial pneumaticity, and dermal armour covering the body. This group has been reported in the Upper Cretaceous of the Lecho, Allen, and Anacleto formations of Argentina. We describe here a new saltasaurine specimen (MACN-Pv RN 233) from the Campanian of the Angostura Colorada Formation (Río Negro Province) that is represented by eight caudal vertebrae and six osteoderms. This specimen is described in detail and its phylogenetic relationships with the other three known saltasaurines, as well as its implications to the knowledge of caudal vertebra and osteoderm anatomy, are discussed. Our results place MACN-Pv RN 233 more closely related to Saltasaurus loricatus and Rocasaurus muniozi than to Neuquensaurus australis. MACN-Pv RN 233 possesses a combination of features that differ from other saltasaurines, but because of the fragmentary nature of the specimen we decided for the sake of taxonomic stability to not erect a new taxon. This specimen shows the first unambiguous evidence of chevron pneumatisation for a sauropodomorph, implying a broader osteological invasion of the diverticula from the abdominal air sac than previously thought for this group of dinosaurs. MACN-Pv RN 233 preserves two osteoderm morphotypes, one similar to those reported for Neuquensaurus australis and Saltasaurus loricatus. This new specimen expands the distribution of the group to a new geological unit and increases the dinosaur diversity known for the Angostura Colorada Formation.  相似文献   

20.
A new theropod dinosaur, Shidaisaurus jinae gen. et sp. nov., has been described on the basis of an incomplete skeleton. The specimen was found near the base of the Upper Lufeng Formation (early Middle Jurassic) in Yunnan, China. It is the first theropod dinosaur from the Middle Jurassic of Yunnan. Shidaisaurus jinae is distinguishable from other Jurassic theropods by certain features from the braincase, axis, and pelvic girdle. The absence of any pleurocoels in the axis or in any anterior dorsal vertebrae suggests that the new Lufeng theropod is relatively primitive and more plesiomorphic than most of the Middle to Late Jurassic theropods from China. Most Chinese taxa of Jurassic theropod dinosaurs have not been well described; a further detailed study will be necessary for us to determine their phylogenetic relationships with Shidaisaurus jinae.  相似文献   

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