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1.
Helius ewa sp. nov., one of the oldest representative of the genus Helius Lepeletier and Serville 1828 (Diptera: Limoniidae) from the Lebanese amber (Lower Cretaceous) is characterized, illustrated and described. The evidences of rapid evolution of the genus Helius are provided. The hypothesis on the origin of the evolution of this genus in Gondwana and the possibility of rapid radiation and expansion in Laurasia are discussed. A complete list of Cretaceous limoniids belonging to Helius is given. 相似文献
2.
Lepisosteid fishes are well known from the Upper Cretaceous of Europe, but only by fragmentary remains from some Cenomanian and Campanian–Maastrichtian deposits. Here we report various cranial and postcranial remains of gars, discovered in the Upper Cretaceous (Santonian) Csehbánya Formation of Iharkút (Bakony Mountains, Hungary). These remains represent one of the most diverse assemblages of lepisosteid fish material from Upper Cretaceous continental deposits of Europe. Based on tooth morphology, scale-microstructure and the features of the supracleithrum we refer these remains to the genus Atractosteus. Besides some uncertain remains from the Cenomanian of France and Spain, the Santonian aged fossils from Iharkút represent the oldest undisputable occurrence of the family Lepisosteidae in the European continental Cretaceous. Using tooth crown morphology, the surface microstructure of the ganoid scales and the anatomy of the supracleithrum a review of the Late Cretaceous lepisosteid record suggests the occurrence of both Atractosteus and Lepisosteus in the European archipelago. 相似文献
3.
Patagonia has yielded the most comprehensive fossil record of Cretaceous theropods from Gondwana, consisting of 31 nominal species belonging to singleton taxa and six families: Abelisauridae, Noasauridae, Carcharodontosauridae, Megaraptoridae nov. fam., Alvarezsauridae, and Unenlagiidae. They provide anatomical information that allows improved interpretation of theropods discovered in other regions of Gondwana. Abelisauroids are the best represented theropods in Patagonia. They underwent an evolutionary radiation documented from the Early Cretaceous through to the latest Cretaceous, and are represented by the clades Abelisauridae and Noasauridae. Patagonian carcharodontosaurids are known from three taxa (Tyrannotitan, Giganotosaurus and Mapusaurus), as well as from isolated teeth, collected from Aptian to Cenomanian beds. These allosauroids constituted the top predators during the mid-Cretaceous, during which gigantic titanosaur sauropods were the largest herbivores. Megaraptorans have become better documented in recent years with the discovery of more complete remains. Megaraptor, Aerosteon and Orkoraptor have been described from Cretaceous beds from Argentina, and these taxa exhibit close relationships with the Aptian genera Australovenator, from Australia, and Fukuiraptor, from Japan. The Gondwanan megaraptorans are gathered into the new family Megaraptoridae, and the Asiatic Fukuiraptor is recovered as the immediate sister taxon of this clade. Although megaraptorans have been recently interpreted as members of Allosauroidea, we present evidence that they are deeply nested within Coelurosauria. Moreover, anatomical information supports Megaraptora as more closely related to the Asiamerican Tyrannosauridae than thought. Megaraptorans improve our knowledge about the scarcely documented basal radiation of Gondwanan coelurosaurs and tyrannosauroids as a whole. Information at hand indicates that South America was a cradle for the evolutionary radiation for different coelurosaurian lineages, including some basal forms (e.g., Bicentenaria, Aniksosaurus), megaraptorans, alvarezsaurids less derived than those of Laurasia, and unenlagiids, revealing that Gondwanan coelurosaurs played sharply differing ecological roles, and that they were taxonomically as diverse as in the northern continents. The unenlagiids represent an endemic South American clade that has been recently found to be more closely related to birds than to dromaeosaurid theropods. Analysis of the theropod fossil record from Gondwana shows the highest peak of origination index occurred during the Aptian–Albian and a less intense one in the Campanian time spans. Additionally, peaks of extinction index are recognized for the Cenomanian and Turonian–Coniacian time spans. In comparison, the Laurasian pattern differs from that of Gondwana in the presence of an older extinction event during the Aptian–Albian time-span and a high origination rate during the Cenomanian time-bin. Both Laurasian and Gondwanan theropod records show a peak of origination rates during the Campanian. 相似文献
4.
Lycian Nappes (in SW Turkey) lie between the Menderes Massif and Bey Dağları carbonates and comprise thrust sheets (nappes piles) of Paleozoic-Cenozoic rocks, ophiolitic and tectonic mélanges and serpentinized peridodites. This study focuses on identification of rudists and their palaeoenvironmental features observed within the Cretaceous low grade metamorphic successions (dominated by recrystallized limestones) from the Tavas and Bodrum nappes. The study is based on fifteen stratigraphic sections measured from Tavas, Fethiye, Köyceğiz, Bodrum, Ören and Bozburun areas. The Lower Cretaceous successions with rudists are very sparse in the Lycian Nappes and a unique locality including a Berriasian epidiceratid-requieniid assemblage is reported so far. A new requieniid-radiolitid assemblage was found within the pre-Turonian (?Albian-?Cenomanian) limestones. Four different Late Cretaceous rudist assemblages were firstly identified as well: 1) Caprinid-Ichthyosarcolitid assemblage (middle-late Cenomanian); 2) Distefanellid assemblage (late Turonian); 3) Hippuritid-Radiolitid assemblage (late Coniacian-Santonian-Campanian); 4) Radiolitid-Hippuritid assemblage (‘middle’-late Maastrichtian). Microfacies data and field observations indicate that the rudists lived in the inner and outer shelves of the Cretaceous carbonate platform(s) in this critical part of the Neotethys Ocean. Rudists formed isolated patchy aggregations in very shallow palaeoenvironments and deposited as shell fragments particularly on the outer shelf environment, which is characterized by higher energy and platform slope characteristics. 相似文献
5.
T. N. Palechek D. P. Savel’ev O. L. Savel’eva 《Stratigraphy and Geological Correlation》2010,18(1):63-82
The Albian—Cenomanian sediments belong to the oldest deposits bearing fossils in the Eastern Kamchatka. Described in this
work is one carbonate-cherty section of the Smaginsk Complex in the Kamchatskii Mys Peninsula. Two interlayers enriched in
sapropelic organic matter represent a peculiar feature of the studied section. The interlayers originated most likely under
the influence of Cretaceous anoxic events. All radiolarian taxa identified formerly and now in rocks of the Smaginsk Complex
are considered. New data clarify the taxonomic composition of radiolarian assemblages, characterizing the Smaginsk Complex.
Some of the currently defined radiolarian species were known before only from the DSDP Sites in the Pacific Ocean. According
to data of radiolarian analysis, the section part sandwiched between two interlayers enriched in organic matter is attributed
to the Cenomanian. The early Cenomanian age is suggested for one of the studied samples. 相似文献
6.
L. G. Bragina 《Stratigraphy and Geological Correlation》2013,21(1):79-95
Recently, representatives of the genus Cuboctostylus Bragina (order Entactinaria) were included in the Upper Cretaceous radiolarian regional stratigraphic scale of Sakhalin. The Late Cretaceous species Hexacromyum pergamenti Bragina (order Spumellaria) has morphological similarity to representatives of the genus Cuboctostylus. Peculiar features of H. pergamenti internal structure are considered. Collections of Upper Cretaceous radiolarians from southern Cyprus, Serbia, northern Turkey, Crimean Mountains, East European Platform, northwestern Kamchatka, eastern slope of the Sredinnyi Range in Kamchatka, and Shikotan Island (Lesser Kurile Range) were used for the analysis of the taxonomic composition of Late Cretaceous representatives of the genera Cuboctostylus Bragina and Hexacromyum Haeckel as well as their stratigraphic and paleobiogeographic distribution. It is established that Cuboctostylus is distributed from tropical to south boreal realms. This genus is shown to exist through almost the entire Late Cretaceous: from the middle Cenomanian to initial Maastrichtian. Hexacromyum Haeckel populated both the south boreal realm and marginal areas of the Tethys Ocean in the Late Cretaceous. The new data presented may be used for distant interregional correlations. Cuboctostylus stellatus sp. nov. and several other Cuboctostylus taxa identified in open nomenclature are described; some morphological features of Hexacromyum pergamenti are specified. 相似文献
7.
《Journal of African Earth Sciences》2000,30(2):401-416
Land-derived pollen and spores and marine dinoflagellate cysts were extracted from the Jurassic and Cretaceous sediments of the West Tiba-1 borehole, northern Western Desert, Egypt, On the basis of the recovered palynomorphs, of known stratigraphical significance, the following stages were assessed: Bathonian-Oxfordian (Middle-Late Jurassic) and Hauterivian, Aptian-Early Albian, Late Albian-Early Cenomanian, Early Cenomanian and Late Cenomanian (Early-Middle Cretaceous). No palynomorphs diagnostic for the Berriasian, Valanginian and Barremian stages (Early Cretaceous) were depicted. Based on the nature and composition of the identified palynomorph content, five informal palynomorph assemblage zones were recognised. These are: the Gonyaulacysta jurassica-Korystocysta kettonensis Assemblage Zone (PI, Bathonian-Oxfordian), Ephedripites-Aequitriradites verrucosus Assemblage Zone (PII, Hauterivian), Afropollis jardinus-Duplexisporites generalis-Tricolpites Assemblage Zone (PIIl, Aptian-Early Albian), Nyssapollenites-Elaterosporites Assemblage Zone (PIV, Late Albian-Early Cenomanian) and Assemblage Zone PV (Early-Late Cenomanian). The latter zone was differentiated into two subzones, namely the Classopollis brasiliensis-Elaterosporites klaszii Assemblage Subzone (PVa, Early Cenomanian) and Afropollis kahramanensis-Triporates Assemblage Subzone (PVb, Late Cenomanian). The time stratigraphy of the studied interval was revised. The occurrences and types of the dinoflagellate cysts, extracted from the studied succession, reflect a general shallow (shelf) marine pal˦oenvironment. 相似文献
8.
Salah El Beialy Maher El-Soughier Sayed Abdel Mohsen Haytham El Atfy 《Journal of African Earth Sciences》2011,59(2-3):215-226
Palynological investigation of the Cretaceous Abu Roash, Bahariya, Kharita, Alamein, Alam El Bueib and Betty formations, encountered in the Gebel Rissu-1 well, north Western Desert, Egypt yielded 27 species of pteridophytic spores, 24 of gymnosperm pollen, 25 of angiosperm pollen and 11 of dinoflagellate cysts in addition to some acritarchs, foraminiferal test linings and freshwater algae. This enabled us to recognize five miospore biozones arranged from youngest to oldest as: Classopollis brasiliensis–Afropollis cf. kahramanensis–Dichastopollenites ghazalataensis Assemblage Zone (Late Cenomanian); Elaterosporites klaszii–Sofrepites legouxae–Afropollis jardinus Assemblage Zone (Middle/Late Albian–Early Cenomanian); Pennipollis peroreticulatus–Duplexisporites generalis-Tricolpates Assemblage Zone (Early Aptian–Early Albian); Tucanopollis crisopolensis–Afropollis sp. Assemblage Zone (Barremian) and Appendicisporites cf. tricornitatus–Ephedripites spp. Assemblage Zone (Late Neocomian).The Early Cretaceous Kharita, Alam El Bueib and the Betty formations encountered in the Gebel Rissu-1 well are interpreted to indicate oxic proximal and distal shelf deposits, characterized by type III/IV, V kerogen, which is gas prone but having little potential to produce hydrocarbons. The Upper Cretaceous Abu Roash and Bahariya formations are characterized by a distal suboxic–anoxic and marginal dysoxic–anoxic environment, and their kerogen type III/II indicates gas/oil prone nature. The Bahariya and Kharita Albian–Cenomanian sediments in the present study witnessed the onset of a semi-arid to arid climate, with local or seasonal humid conditions, based on the continuous high abundance of the elaterates pollen and Afropollis-producing plants that inhabited the paleotropical humid coastal plains. 相似文献
9.
The oldest laniatorean harvestman, Petrobunoides sharmai gen. et sp. nov. (Opiliones: Laniatores) is described, from the Upper Cretaceous (lowermost Cenomanian) amber of Myanmar. This is the first fossil Laniatores recovered from Southeast Asia, which is placed in the extant family Epedanidae. It is also the first fossil known within the superfamily Epedanoidea; a relatively derived clade of Laniatores restricted to Southeast Asia today. At ca. 99 Ma, this new amber inclusion is substantially older than the previous oldest record of a member of Laniatores from Baltic amber (ca. 44–49 Ma); however, given the Palaeozoic age of Laniatores implied by molecular data, the new record from Burmese amber is probably still too young for constraining or calibrating the date of cladogenesis for total group Laniatores. Nevertheless, it provides a much better constraint for Epedanoidea, adding the first useful laniatorean fossil to provide an internal calibration point for a clade of Grassatores. 相似文献
10.
A new Upper Cretaceous genus and species of soldier beetles, Archaeomalthodes rosetta gen. et sp. nov., is described and illustrated from an individual preserved in Upper Cretaceous (Cenomanian, ca. 99 Ma) amber from northern Myanmar. It is undoubtedly placed in extant subfamily Malthininae based on its small-sized body, somewhat abbreviated elytra and fusiform terminal maxillary palpomere, representing the oldest documented occurrence of Malthininae. It suggests that this subfamily is an ancient group, which originated at least in the earliest Late Cretaceous. Our discovery sheds light on the palaeodiversity of Cantharidae in the Late Mesozoic. Together with other previously reported fossil cantharids, it is likely that Malthininae has been fairly diverse during the early evolution of Cantharidae. On the other hand, a morphological similarity between Archaeomalthodes and Recent malthinines and the occurrence of flowering plants in the Burmese amber implies a potential flower-visiting behaviour of this fossil species. 相似文献
11.
Ornithuromorph birds (the clade which includes modern avian radiation) first appeared in the Early Cretaceous in Asia and achieved a great diversity during the latest ages of the Late Cretaceous (Campanian and Maastrichtian). The evolutionary history of orithuromorphs during the first 17 MYAs of the Late Cretaceous (Cenomanian to Santonian ages) remains very poorly known, as the fossil record for this time interval is largely restricted to several isolated finds of the classic avian genus Ichthyornis in North America. Here we describe an isolated distal tibiotarsus of an evolutionary advanced bird, morphologically similar to Ichthyornis, from the middle Cenomanian of Saratov Province, European Russia. This is the first documentation of an Ichthyornis-like bird in the Old World. The find further constitutes only the second pre-Campanian record of the Late Cretaceous Ornithuromorpha in Eurasia, the second record of Cenomanian birds in Russia. This discovery shows that Ichthyornis-like birds enjoyed a wide geographical distribution as early as the beginning of the Late Cretaceous. Given that the earliest and the most primitive ornithuromorph birds are known from Asia, the new find supports a Eurasian origin for Ichthyornithidae. 相似文献
12.
The phylostratigraphy, taphonomy and palaeoecology of the Late Cretaceous neoselachian Ptychodus of northern Germany appears to be facies related. Ptychodus is not present in lower Cenomanian shark-tooth-rich rocks. First P. oweni records seem to relate to middle Cenomanian strata. P. decurrens appears in the middle to upper Cenomanian mainly in non-coastal environments of the shallow marine carbonate ramp and swell facies which isolated teeth were found partly in giant ammonite scour troughs on the Northwestphalian-Lippe High submarine swell in the southern Pre-North Sea Basin. They are recorded rare in deeper basin black shales facies (upwelling influenced, OAE Event II). P. polygyrus seems to be restricted to upwelling influenced basin and deeper ramp facies mainly of the uppermost Cenomanian and basal lower Turonian (OAE II Event). P. mammillaris is mostly represented during the lower to middle Turonian in the inoceramid-rich ramp and the near shore greensand facies along the Münsterland Cretaceous Basin coast north of the Rhenish Massif mainland. Finally, P. latissimus is recorded by two new tooth sets and appears in the upper Turonian basin swell facies and the coastal greensands. Autochthonous post-Turonian Ptychodus remains are unrecorded in the Santonian–Campanian of Germany yet. Reworked material from Cenomanian/Turonian strata was found in early Santonian and middle Eocene shark-tooth-rich condensation beds. With the regression starting in the Coniacian, Ptychodus disappeared in at least the Münster Cretaceous Basin (NW-Germany), but remained present at least in North America in the Western Interior Seaway. The Cenomanian/Turonian Ptychodus species indicate a rapid neoselachian evolution within the marine transgression and global high stand. A correlation between inoceramid shell sizes, thicknesses and their increasing size during the Cenomanian and Turonian might explain the more robust and coarser ridged enamel surfaces in Ptychodus teeth, if Ptychodus is believed to have preyed on epifaunistic inoceramid bivalves. 相似文献
13.
Eurypoda, the major radiation of armoured dinosaurs, comprises the ankylosaurs and their sister group, the stegosaurs. As the earliest-branching major clade of ornithischian dinosaurs, the evolutionary history of Eurypoda is significant for understanding both the palaeobiology of bird-hipped dinosaurs and the composition of middle Mesozoic ecosystems. Eurypodans were diverse and abundant throughout the Late Jurassic and Cretaceous in Laurasia; in contrast, their remains are extremely rare in Gondwana. Herein, we describe a new genus and species of stegosaur from the Middle Jurassic of Morocco, Adratiklit boulahfa. Adratiklit is the first eurypodan from north Africa and the oldest definitive stegosaur from anywhere in the world. The genus is more closely related to the European stegosaurs Dacentrurus and Miragaia than it is to the southern African taxa Kentrosaurus and Paranthodon. Statistically significant correlations between the number of dinosaur-bearing formations, dinosaur-bearing collections, and eurypodan occurrences in Gondwana indicates that their fossil record is biased by both geological and anthropogenic factors. Tantalizing but fragmentary remains and trackways suggest that eurypodan diversity in Gondwana may have been as rich as that of Laurasia, and the prospects for future discoveries of new genera across Gondwana are therefore very good. 相似文献
14.
《Proceedings of the Geologists' Association. Geologists' Association》2023,134(1):107-114
New material of pterasterid asteroids from the UK chalk is described on the basis of ossicles recovered from washed residues. A new species, Pteraster lyddenensis sp. nov., is erected for oral and adambulacral ossicles and a primary radial ossicle from the Cenomanian Grey Chalk Subgroup of Dover (Kent), and the first UK record of Pteraster kutscheri Gale, 2022 is described from the upper Campanian Chalk of Norwich (Norfolk); both taxa belong to extant groups of Pteraster. Pteraster lyddenensis sp. nov. is the oldest known representative of the genus. The benthic invertebrate fauna of the Cretaceous chalk facies includes a number of extant genera which at the present day dwell in the deep sea. However, their presence was probably due to the low-productivity oceanic palaeoenvironment of the Chalk Sea, simulating deep-ocean conditions, rather than its depth. 相似文献
15.
In this paper we describe previously unpublished trionychid turtle material, consisting of numerous shell fragments, from the Late Cretaceous (late Turonian) Bissekty Formation of the Dzharakuduk locality in Uzbekistan. This material is assigned to two shell-based taxa: Aspideretoides cf. riabinini and “Trionyx” cf. kansaiensis. The material which cannot be confidently attributed to these two taxa is identified as Trionychidae indet. In addition to these shell-based trionychid taxa, the Dzharakuduk turtle assemblage includes two skull-based taxa of trionychids (Khunnuchelys kizylkumensis and Trionychini indet.). The trionychids from the Bissekty Formation are most similar to trionychids from the younger (Santonian – early Campanian) Bostobe Formation of Kazakhstan, represented by three shell-based taxa (Aspideretoides riabinini, Paleotrionyx riabinini and “Trionyx” kansaiensis), and one skull-based taxon (Khunnuchelys sp.). We provide an improved understanding of the subtle similarities and differences between four closely related Cretaceous turtle assemblages of Middle Asia and Kazakhstan. 相似文献
16.
The stratigraphy of the western Portugal on-shore Cretaceous record (western Iberian margin, Lusitanian Basin) is described, including formal units and a selection of informal units prevailing in the geological literature. This paper is a synthesis based on a review of previous works, but with an innovative emphasis on the interpretation of eustatic and tectonic controls. The sedimentary record is dominated by siliciclastics and comprises fluvial and deltaic coastal marine siliciclastic systems, as well as extensive deposits of shallow marine carbonate platforms, both open and rimmed. Several regional unconformities and transgressive/regressive cycles are identified and the allogenic controls interpreted, namely the geodynamic events along the boundaries of the Iberian plate. Above the Berriasian deposits belonging to the Upper Jurassic cycle, the five main unconformity-bounded units are: (1) upper Berriasian–lower Barremian, (2) upper Barremian–lower Aptian, (3) upper Aptian–uppermost Cenomanian, (4) mid lower Turonian–lower Campanian and (5) middle Campanian–Maastrichtian. These units show transgressive peaks in the lower Hauterivian, lower Aptian, base of the upper Cenomanian and mid lower Turonian. The general trend of the Lower Cretaceous reflects the transition from late rifting to passive margin, with the last break-up unconformity dated as late Aptian. The Lusitanian Basin achieved full infill by the Cenomanian, when a large carbonate platform extended far inland. The later deposits were preserved only in the northern sector and the accompanying unconformities reflect transpressive intraplate stresses generated in boundaries of the plate with Africa and Eurasia. With very low accommodation being created throughout the Late Cretaceous, fluvial deposits were dominant, including a few marine levels related with eustatic rises in the early Turonian, the Coniacian, the early Campanian and the Maastrichtian. 相似文献
17.
The extant fern genus Asplenium Linnaeus, 1753 is widely distributed in tropical-temperate regions and shows a high diversity. The oldest fossils assigned to it comprise fronds and spores preserved in situ from the Lower Cretaceous of northeast Asia. However, molecular dating suggests that Asplenium diverged during the Paleocene (∼57.7 Ma). Here we present some explanations for the disparity in fossil and molecular age estimates, and new insights into the origin and evolution of the genus. We suggest that Cretaceous Asplenium most likely represents a stem-group member, and that lineage extinction during the early evolutionary stages of the genus resulted in missing nodes. 相似文献
18.
A remarkable new genus and species, Mesallotrochus longiantennatus n. gen. n. sp., is described and figured based on a well-preserved individual in the lowermost Upper Cretaceous (Cenomanian) amber from Myanmar. Mesallotrochus is placed in the extant tribe Thoracophorini based on its general habitus, including the protibia with inner edge straight, without ctenidium, exposed protrochantins, open procoxal cavities, and more or less flatted body. Mesallotrochus is separated from other allied genera by the long antennae, very long maxillary palpomere 4, contiguous procoxae, and well-developed anterolateral pronotal angles. The new discovery of the oldest Osoriinae from about 99 million years not only suggests the antiquity of the subfamily, but also bears significant biogeographic implications. 相似文献
19.
Maher I. El-Soughier Amr S. Deaf Magdy S. Mahmoud 《Arabian Journal of Geosciences》2014,7(8):3051-3068
Palynological and palynofacies analyses were carried out on some Cretaceous samples from the Qattara Rim-1X borehole, north Western Desert, Egypt. The recorded palynoflora enabled the recognition of two informal miospore biozones arranged from oldest to youngest as Elaterosporites klaszii-Afropollis jardinus Assemblage Zone (mid Albian) and Elaterocolpites castelainii–Afropollis kahramanensis Assemblage Zone (late Albian–mid Cenomanian). A poorly fossiliferous but however, datable interval (late Cenomanian–Turonian to ?Campanian–Maastrichtian) representing the uppermost part of the studied section was also recorded. The palynofacies and visual thermal maturation analyses indicate a mature terrestrially derived organic matter (kerogen III) dominates the sediments of the Kharita and Bahariya formations and thus these two formations comprise potential mature gas source rocks. The sediments of the Abu Roash Formation are mostly dominated by mature amorphous organic matter (kerogen II) and the formation is regarded as a potential mature oil source rock in the well. The palynomorphs and palynofacies analyses suggest deposition of the clastics of the Kharita and Bahariya formations (middle Albian and upper Albian–middle Cenomanian) in a marginal marine setting under dysoxic–anoxic conditions. By contrast, the mixed clastic-carbonate sediments of the Abu Roash Formation (upper Cenomanian–Turonian) and the carbonates of the Khoman Formation (?Campanian–Maastrichtian) were mainly deposited in an inner shallow marine setting under prevailing suboxic–anoxic conditions as a result of the late Cenomanian and the Campanian marine transgressions. This environmental change from marginal to open (inner shelf) basins reflects the vertical change in the type of the organic matter and its corresponding hydrocarbon-prone types. A regional warm and semi-arid climate but with a local humid condition developed near/at the site of the well is thought to have prevailed. 相似文献
20.
A new genus and species of cimicomorphan bug, Aphrastomedes anthocoroides Yamada and Yamamoto, gen. and sp. nov., is described and illustrated based on two specimens preserved in Upper Cretaceous (Cenomanian) amber from the Hukawng Valley of northern Myanmar (Burma). This remarkable cimicomorphan species is tentatively assigned to the family Velocipedidae primarily on the basis of the structure of the labium, forewings, and genitalia. Aphrastomedes is considered best placed in the Cimiciformes as a member of the Velocipedidae, and it also has a close relationship to the Medocostidae. 相似文献