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1.
A new genus and species of ripiphorid beetle is described based on a female specimen preserved in Upper Cretaceous (Cenomanian, ca. 99 Ma) amber from northern Myanmar, under the name of Spinotoma ruicheni gen. et sp. nov. It belongs to the extant Pelecotominae and represents the second documented occurrence of this subfamily in the Late Cretaceous. This new taxon is morphologically distinguishable in Pelecotominae by the combination of following characters: eyes oval, without incision; antennomeres V–X serrate, with XI fusiform; maxillary palpi unmodified; protarsi shorter than protibiae; tibiae sparsely provided with spines; tibial spurs formula 2-0-2. Together with other previously reported fossil ripiphorids, it highlights the species diversification of wedge-shaped beetles during their early evolution. On the other hand, a morphological similarity between Spinotoma and modern pelecotomines and the occurrence of xylophagous beetles in the Burmese amber suggest a similar host preference in wood-boring beetles for this species, shedding lights on an ancient host-parasitoid relationship in the late Mesozoic. This paper also provides a brief discussion on the taxonomy and evolution of Mesozoic wedge-shaped beetles. The wedged-shaped body form and fully-developed elytra in Recent Pelecotominae and Ptilophorinae are considered to be derived from their early ancestors rather than evolved with the onset of flowering plants during the Cretaceous as commonly thought.  相似文献   

2.
A new polypore fungus beetle is described and illustrated, under the name of Thescelostrophus cretaceus gen. et sp. nov., representing the first documented occurrence of the tribe Eustrophini. The well-preserved specimen was collected from the Upper Cretaceous (Cenomanian, ca. 99 Ma) amber near the Hukawng Valley of northern Myanmar. This fossil species can be assigned to the extant subfamily Eustrophinae based on its elongate oval and strongly narrowed posteriorly body, simple and narrow tarsi, and somewhat clubbed antennomeres. The comparison among the extinct and extant eustrophines supports the previous hypothesis that antennal morphology of early eustrophines was very diverse. Additionally, an overall similarity between Thescelostrophus and extant eustrophines suggests a potential fungivory of this fossil species. Morphological characters preserved in the Burmese amber highlight the diversity of tetratomids during the Late Mesozoic and provide data for future phylogenetic studies of Tetratomidae.  相似文献   

3.
The enigmatic staphylinid subfamily Dasycerinae consists of only 17 species within a single extant genus Dasycerus, but it is easily distinguished from other rove beetles by overall, latridiid beetle-like body and extremely slender, verticillate antennae. Direct fossil evidence is lacking for this group. Here, I describe the first fossil of this peculiar subfamily, Protodasycerus aenigmaticus gen. and sp. n., from the Upper Cretaceous Burmese amber. Despite considerable external similarities to extant dasycerine species, this new genus possesses several important morphological differences: a much smaller body size (ca. 1.1 mm), narrowly elongate antennomeres 1 and 2, not strongly transverse metaventrite, elytra striate but not tricostate, and truncate posterior margin of the elytra exposing four abdominal tergites dorsally. The new finding of a Cretaceous representative of the Dasycerinae helps elucidate the origin and early evolution of the omaliine group subfamilies, implying rather close similarities between the Dasycerinae and the monobasic subfamily Neophoninae. Discovery of P. aenigmaticus suggests a rather ancient origin of the subfamily by the Upper Cretaceous.  相似文献   

4.
The exclusively Cretaceous braconid wasp subfamily Protorhyssalinae is reported from the Upper Cretaceous deposits of northern Myanmar. Archaeorhyssalus subsolanus Engel, gen. et sp. nov., is represented by a well-preserved female in Burmese amber and is similar to other protorhyssalines, but differs in numerous details of forewing venation and flagellomere number. This species is the first definitive Oriental occurrence for protorhyssalines, expanding their palaeogeographic distribution from the Lower Cretaceous of Spain, and the Upper Cretaceous of France and New Jersey, USA. A key to the known genera of protorhyssalines is provided.  相似文献   

5.
A new genus and species of female phantom midge Neuseptychoptera carolinensis gen. et sp. nov. in the fossil subfamily Eoptychopterinae is described and illustrated; this ptychopterid is from Upper Cretaceous (Campanian) amber of North Carolina. This is the first member of Eoptychopterinae from North America and the first found in Campanian age amber. We propose to treat the extinct subfamily Eoptychopterininae Lukashevich, 1995, as a junior synonym of the subfamily Eoptychopterinae Handlirsch, 1906, syn. nov. The elongated mouthparts of the new species have broad labella, and these are functionally interpreted as feeders for honeydew, fermenting plant juices, and nectar rather than for biting.  相似文献   

6.
We describe two new species of extinct biting midges in the fossil genus Protoculicoides: Protoculicoides hispanicus Szadziewski and Arillo, sp. nov. and Protoculicodes sanjusti Szadziewski and Arillo, sp. nov., from Albian (101–113 Ma) amber of San Just, Spain. Atriculicoides Remm, 1976 is recognized as a new junior synonym of Protoculicoides Boesel, 1937 (NEW SYNONYM). Devalquia Choufani et al., 2013 is regarded here as a new junior synonym of the extant genus Culicoides Latreille, 1809 (NEW SYNONYM). Metahelea roggeroi Choufani et al., 2013 from Upper Cretaceous French amber, is excluded from the tribe Heteromyiini and placed in the extant genus Stilobezzia Kieffer, 1911, tribe Ceratopogonini: Stilobezzia roggeroi (Choufani et al., 2013), comb. nov. The extinct tribe Atriculicoidini Szadziewski, 1996 is regarded as a new subfamily, Atriculicoidinae, stat. nov., that includes the fossil genus Protoculicoides, with 13 species reported from Albian, Turonian, Cenomanian, Coniacian, Santonian and Campanian ambers (78–113 Ma). Protoculicoides skalskii Szadziewski & Arillo, 1998 from Lower Cretaceous amber of Álava, Spain, Protoculicoides succineus Szadziewski, 1966 from Lower Cretaceous Lebanese amber and Protoculicoides burmiticus Szadziewski & Poinar, 2005 from Upper Cretaceous Burmese amber are transferred to the extinct genus Archiculicoides Szadziewski, 1996, comb. nov. Protoculicoides krzeminskii Choufani et al., 2014 from Lower Cretaceous Lebanese amber is transferred to the extinct genus Archiaustroconops Szadziewski, 1996, comb. nov. The Cretaceous subfamily Atriculicoidinae forms an unresolved trichotomy with the extant subfamilies Forcipomyiinae and Dasyheleinae, both of which date to the Eocene. The fossil record indicates that wings with macrotrichia in biting midges evolved during the mid-Cretaceous greenhouse climate. We suggest that this was an evolutionary adaptation to new atmospheric conditions with higher levels of CO2 in order for the antennal Johnston's organs of males to receive the vibrational sex signals produced by females during flight.  相似文献   

7.
Mesozoic whip-scorpions are very rare, with only two Cretaceous species known to date. Here we describe a new genus and species of Thelyphonidae, Mesothelyphonus parvus gen. & sp. nov., based on a very well-preserved male in Upper Cretaceous amber from Myanmar. Mesothelyphonus is firmly placed in the extant subfamily Thelyphoninae as supported by the abdominal tergites with a median longitudinal suture. Mesothelyphonus differs from other fossil and recent genera primarily by its very small body size, the absence of ommatoids on abdominal segment XII, and the elongate, slender and toothed patellar apophysis of the male pedipalp. The new discovery represents the oldest definitive fossil record for Thelyphoninae, highlighting the antiquity of the whip-scorpion group.  相似文献   

8.
A new genus and species of extinct rove beetle, Procileoporus burmiticus gen. and sp. n., is described from Upper Cretaceous Burmese amber (Cenomanian; ca. 99 Ma). Due to the condition of the fossil, parts of the body could not be examined, but sufficient characteristics were visible to allow the specimen to be placed within the subfamily Tachyporinae (tribe Tachyporini). Procileoporus is most similar to the extant genus Cileoporus Campbell in having a slender abdomen, but it differs in having much shorter abdominal setae, prominent striated microreticulation on the elytra, non-lobed tergite VIII, and in other details of the male genital segments. This is the first tachyporine rove beetle from Burmese amber and the oldest record of the subfamily in amber. The evolution of sexual dimorphism in the genital segments is briefly discussed.  相似文献   

9.
Fossil smicripid beetles are very rare. Here we describe and figure the first Mesozoic representative of Smicripidae. Smicrips cretacea sp. nov., a new species belonging to the extant Smicrips LeConte is preserved in the Upper Cretaceous amber from northern Myanmar. Our discovery suggests that Smicrips is an ancient group, originating at least in the earliest Late Cretaceous. Together with other two fossil Smicrips species from the Eocene of Europe, it reinforces the hypothesis that the Recent distribution of Smicripidae is relict, and the family was more widespread in the Mesozoic and early Cenozoic than it is in the Recent.  相似文献   

10.
A new genus and species of tetrablemmid spider, Electroblemma bifida n. gen. et sp., is described, from two adult males found in Upper Cretaceous (Cenomanian) amber of Myanmar. The genus is distinguished by its enormous dorsal carapace projection and highly modified chelicerae. The new genus is referred to the tribe Tetrablemmini within the subfamily Tetrablemminae. The presence of a relatively derived tetrablemmid on the south-east Eurasian continent during the Late Cretaceous suggests that the family was already well diversified in tropical rainforests at this time.  相似文献   

11.
A new subfamily of Ichneumonidae, Novichneumoninae subfam. nov., is established based on two new genera with two new species: Novichneumon longus gen. et sp. nov. and Caloichneumon perrarus gen. et sp. nov. These two new species are the first ichneumonids described from the Upper Cretaceous Myanmar (Burmese) amber. A list of all described Mesozoic ichneumonid fossil species with their respective localities and ages is summarized. The distribution of Ichneumonoidea during the Cretaceous indicates that Cretaceous ichneumonids were documented from localities at high latitudes while braconids were distributed worldwide, a pattern consistent with the distribution of extant ichneumonids and braconids.  相似文献   

12.
A new species of Dicranoptycha Osten Sacken, 1860, Dicranoptycha plicativa sp. nov., and a unassigned species are illustrated and described from Myanmar (Burmese) amber (lowermost Cenomanian, Upper Cretaceous). A complete list of fossil species of the genus Dicranoptycha is summarized. In addition, an updated key to the known fossil species of the Dicranoptycha is provided.  相似文献   

13.
14.
The new genus and species Angustaeshna magnifica of Burmaeshnidae is described on the basis of a new fossil from Burmese amber. The genus Cretaeshna from the same amber is transferred from the Telephlebiidae into the Burmaeshnidae. We redefine this last family, no longer considered as the sister group of the Late Cretaceous Enigmaeshnidae, but as putative sister group of the Telephlebiidae in the Aeshnoidea. No known fossil belongs to the Telephlebiidae.  相似文献   

15.
Mesozoic leiodids are poorly known, and only one definitive leiodid is formally described from Burmese amber. Here we describe and illustrate the second definitive Mesozoic leiodid, Cretagyrtodes glabratus gen. et sp. nov., based on a single specimen from the Upper Cretaceous Burmese amber. The fossil is placed in Agyrtodini (subfamily Camiarinae) after maxillary palpomere 4 as wide as palpomere 3, and procoxal cavities closed behind. Cretagyrtodes is tentatively attributed to the extant “Eupelates group”. The discovery of Cretagyrtodes in Burmese amber suggests that the south hemisphere endemic tribe Agyrtodini is probably an ancient group, which has showed its first appearance before the breakup of Pangaea.  相似文献   

16.
<正>Whalfera wiszniewskii sp.nov.is described from the Late Eocene Baltic amber.The genus Whalfera is considered as the only fossil genus confidently assigned to the Rhachlberothinae.Others previously placed in this subfamily belong to Paraberothinae(except perhaps for Oisea).The Late Eocene/present Rhachiberothinae and the Cretaceous Paraberothinae are considered to be the subfamilies of Berothidae.  相似文献   

17.
Cretaceous bryophyte fossils are known only from a few localities and only a few are known in ultrastructural fidelity. We describe a fossil moss gametophyte from Upper Cretaceous Taimyr amber and place it in the fossil genus Taimyrobryum martynoviorum gen. et sp. nov. (Bryopsida). Its unbranched, somewhat zig-zagged stems with distantly spaced, spreading, narrowly lanceolate leaves and the unbranched costa match the gross morphology of Dicranidae; however, the uniformly prosenchymatous leaf cells contradict an affiliation to this subclass. The Yantardakh in the Taimyr Peninsula is the fourth and northernmost Cretaceous amber locality yielding inclusions of bryophytes.  相似文献   

18.
The new tribe Taimyralticini trib. nov., new genus Taimyraltica gen. nov. and new species Taimyraltica calcarata sp. nov. (Coleoptera: Chrysomelidae: Galerucinae) are described from Upper Cretaceous (Santonian) Taimyr amber of northern Siberia (Yantardakh locality). The new genus shares characters with genera of the tribe Alticini (small size, transverse antebasal groove on pronotum, regular punctate-striate elytra) and the Galerucini (narrow, non-swollen metafemora). A possible apomorphy of the new genus and tribe is a large and acute metatibial spur, erect and oriented transversely to the longitudinal axis of tibia. This character state is unknown in recent and fossil genera of Galerucinae. Early evolution and divergence of leaf beetles are discussed, and a Jurassic and Early Cretaceous subfamily radiation is proposed. Fossil galerucines are reviewed. The oldest Phyllotreta is recorded from the lower Miocene of Izarra (Spain). Our findings show an extreme rarity of Phytophaga in Taimyr amber from Yantardakh. Absence of Phytophaga was recorded earlier at Obeschayuschiy (Santonian-Campanian of the Magadan Region) and compared with their much larger share in Khetana (north of Khabarovsk Krai); we find nearly the same differences between Taimyr amber from Yantardakh and Canadian amber. In Obeschayuschiy this appears due to the Mesophytic character of the flora. It is assumed that the Santonian amber gymnosperm forest of the Khatanga Basin had the same character.  相似文献   

19.
Oxyporinae is a distinct staphylinid subfamily characterized by a large body, remarkably long and projecting mandibles, and large, crescent-shaped terminal labial palpomeres. Previously, only three compression fossils from the Lower Cretaceous Yixian Formation in China were known for the subfamily from the Mesozoic Era. This study describes a remarkable new species, Oxyporus cretaceous sp. nov., based on a single specimen in Upper Cretaceous Burmese amber. This finding is the earliest record of Oxyporinae preserved as amber inclusion. The well preserved mouthparts suggest that mycophagous feeding habits had already been acquired by the mid-Cretaceous.  相似文献   

20.
A new genus and species of the ant-like stone beetle supertribe Mastigitae, Cretoleptochromus archaicus gen. et sp. nov., is described and figured based on an exceptionally well-preserved individual in the Upper Cretaceous (lowermost Cenomanian, ca. 99 Ma) amber from Myanmar. Cretoleptochromus is undoubtedly placed in Mastigitae based on its distinctly geniculate antennae with strongly elongate antennal scape, and pointed labial palpomere III. Cretoleptochromus is most likely the sister taxon to the extinct Palaeoleptochromus O'Keefe from the Late Cretaceous Canadian amber. Cretoleptochromus shares with Clidicini the distinct posterior collar of pronotum; with Leptomastacini the presence of mesocoxal lateroventral bristles. The new discovery may help to elucidate the relationships among the Recent three tribes, particularly for Leptomastacini and Clidicini.  相似文献   

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