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This paper provides comprehensive evidence that sediment routing around pools is a key mechanism for pool‐riffle maintenance in sinuous upland gravel‐bed streams. The findings suggest that pools do not require a reversal in energy for them to scour out any accumulated sediments, if little or no sediments are fed into them. A combination of clast tracing using passive integrated transponder (PIT) tagging and bedload traps (positioned along the thalweg on the upstream riffle, pool entrance, pool exit and downstream riffle) are used to provide information on clast pathways and sediment sorting through a single pool‐riffle unit. Computational fluid dynamics (CFD) is also used to explore hydraulic variability and flow pathways. Clast tracing results provide a strong indication that clasts are not fed through pools, rather they are transported across point bar surfaces, or around bar edges (depending upon previous clast position, clast size, and event magnitude). Spatial variations in bedload transport were found throughout the pool‐riffle unit. The pool entrance bedload trap was often found to be empty, when the others had filled, further supporting the notion that little or no sediment was fed into the pool. The pool exit slope trap would occasionally fill with sediment, thought to be sourced from the eroding outer bank. CFD results demonstrate higher pool shear stresses (τ ≈ 140 N m–2) in a localized zone adjacent to an eroding outer bank, compared to the upstream and downstream riffles (τ ≈ 60 N m–2) at flows of 6 · 2 m3 s–1 (≈ 60% of the bankfull discharge) and above. There was marginal evidence for near‐bed velocity reversal. Near‐bed streamlines, produced from velocity vectors indicate that flow paths are diverted over the bar top rather than being fed through the thalweg. Some streamlines appear to brush the outer edge of the pool for the 4 · 9 m3 s–1 to 7 · 8 m3 s–1 (between 50 and 80% of the bankfull discharge) simulations, however complete avoidance was found for discharges greater than this. Copyright © 2013 John Wiley & Sons, Ltd.  相似文献   

4.
Average pool spacing between five and seven bankfull widths has been documented in environments throughout the world, but has limited theoretical justification in coarse‐bedded and bedrock environments. Pool formation in coarse‐bedded and bedrock channels has been attributed to bedrock and boulder constrictions. Because the spacing of these constrictions may be irregular in nature, it is difficult to reconcile pool‐formation processes with the supposedly rhythmic spacing of pools and riffles. To address these issues, a simulation model for pool and riffle formation is used to demonstrate that semi‐rhythmic spacing of pools with an approximate spacing of five to seven bankfull widths can be recreated from a random distribution of obstructions and minimum pool‐ and riffle‐length criteria. It is assumed that a pool–riffle couplet will achieve a minimum length based on dominant‐discharge conditions. Values for the minimum‐length assumption are based on field data collected in New England and California, while the theoretical basis relies on the demonstrated hydraulic response of individual pools to elongation. Results from the simulations show that the location of pools can be primarily random in character, but still assume an average spacing between four and eight bankfull widths for a variety of conditions. Field verification data generally support the model but highlight a highly skewed distribution of pool‐forming elements and pool spacing. The relation between pool spacing and bankfull widths is attributed to the common geometric response of these features to dominant‐discharge conditions. Copyright © 2001 John Wiley & Sons, Ltd.  相似文献   

5.
Riffle‐pool sequences are a common feature of gravel‐bed rivers. However, mechanisms of their generation and maintenance are still not fully understood. In this study a monitoring approach is employed that focuses on analysing cross‐sectional and longitudinal channel geometry of a large floodplain river (Vereinigte Mulde, Sachsen‐Anhalt, Germany) with a high temporal and spatial resolution, in order to conclude from stage‐dependant morphometric changes to riffle and pool maintaining processes. In accordance with previous authors, pool cross‐sections of the Mulde River are narrow and riffle cross‐sections are wide suggesting that they should rather be addressed as two general types of channel cross‐sections than solely as bedforms. At high flows, riffles and pools in the study reaches changed in length and height but not in position. Pools were scoured and riffles aggraded, a development which was reversed during receding flows below the threshold of 0·4Qbf (40% bankfull discharge). An index for the longitudinal amplitude of riffle‐pool sequences, the bed undulation intensity or bedform amplitude, is introduced and proved to be highly significant as a form parameter, its first derivative as a process parameter. The process of pool scour and riffle fill is addressed as bedform maintenance or bedform accentuation. It is indicated by increasing longitudinal bed amplitudes. According to the observed dynamics of bed amplitudes, maintenance of riffle‐pool sequences lags behind discharge peaks. Maximum bed amplitudes may be reached with a delay of several days after peak discharges. Increasing bed undulation intensity is interpreted to indicate bed mobility. Post‐flood decrease of the bed undulation intensity indicates a retrograde phase when transport from pools to riffles has ceased and bed mobility is restricted to riffle tails and heads of pools. This type of transport behaviour is referred to as disconnected mobility. The comparison of two river reaches, one with undisturbed sediment supply, the other with sediment deficit, suggests that high bed undulation intensity values at low flows indicate sediment deficit and potentially channel degrading conditions. It is more generally hypothesized that channel bed undulations constitute a major component of form roughness and that increased bed amplitudes are an important feature of channel bed adjustment to sediment deficit be it temporally during late floods or permanently due to a supply limitation of bedload. Copyright © 2011 John Wiley & Sons, Ltd.  相似文献   

6.
Bedload transport data from planebed and step‐pool reach types are used to determine grain size transport thresholds for selected upland streams in southeast Australia. Morphological differences between the reach types allow the effects of frictional losses from bedforms, microtopography and bed packing to be incorporated into the dimensionless critical shear stress value. Local sediment transport data are also included in a regime model and applied to mountain streams, to investigate whether empirical data improve the delineation of reach types on the basis of dimensionless discharge per unit width (q*) and dimensionless bedload transport (qb*). Instrumented planebed and step‐pool sites are not competent to transport surface median grains (D50s) at bankfull discharge (Qbf). Application of a locally parametrized entrainment equation to the full range of reach types in the study area indicates that the majority of cascades, cascade‐pools, step‐pools and planebeds are also not competent at Qbf and require a 10 year recurrence interval flood to mobilize their D50s. Consequently, the hydraulic parameters of the regime diagram, which assume equilibrium conditions at bankfull, are ill suited to these streams and provide a poor basis of channel delineation. Modifying the diagram to better reflect the dominant transported bedload size (equivalent to the D16 of surface sediment) made only slight improvements to reach delineation and had greatest effect on the morphologies with smaller surface grain sizes such as forced pool‐riffles and planebeds. Likewise, the Corey shape factor was incorporated into the regime diagram as an objective method for adjusting a base dimensionless critical shear stress (τ*c50b) to account for lithologically controlled grain shape on bed packing and entrainment. However, it too provided only minor adjustments to reach type delineation. Copyright © 2007 John Wiley & Sons, Ltd.  相似文献   

7.
Despite the occurrence of riffle–pool sequences in many rivers there are few data concerning riffle–pool unit morphology. Of many criteria proposed to identify riffle–pool units, only three methods can be regarded as objective and robust. These are the ‘zero‐crossing’, the ‘spectral analysis’ and the ‘control‐point’ methods. In this paper statistics are developed using the first two of these methods to describe the streamwise morphology of 275 riffles and 285 pools which form a continuous 32·1 km reach of the bed of the River Severn in Shropshire, England. Yalin's theoretical relationship between the average riffle:pool unit length ( λ p) and channel width ( W ), λ p = 3 W , applies to the River Severn. Reach‐average riffle height ( H ) is a constant proportion of bankfull depth ( h ); typically H ≅ 0·16 h . Riffle height is a positive function of riffle length. Pool depth is a positive function of pool length. However, both riffle length and pool length increase more rapidly than the bed‐level amplitude, such that long riffles or pools are relatively ‘flat’. As channel gradient reduces, bedforms flatten and become more asymmetric as riffle stoss sides and the proximal slope of pools lengthen at the expense of riffle lee sides and pool distal slopes. The statistical relationships between riffle steepness (H/L) and water depth are similar to those for equilibrium subaqueous dunes. The Severn data are consistent with Yalin's theoretical analysis relating riffle bedform length (L r) to water depth, i.e. L r = α2π h, wherein α ≅ 1 for steep near‐equilibrium bedforms but α ≅ 2 to 3 as the relative depth decreases and riffles become long, low features. Theoretical consideration and turbulence data indicate that the frequency of coherent turbulent‐flow structures associated with the riffle–pool mixing length in the Severn should be of the order of 50 to 100 s. The morphological similarity of the steepest River Severn riffles with dunes raises intriguing questions with respect to self‐similar, convergent organization of periodic alluvial bedforms and to bedform dynamic classification particularly. Copyright © 2000 John Wiley & Sons, Ltd.  相似文献   

8.
The ‘velocity-reversal hypothesis’ is the linchpin for a number of recent conceptual models pertaining to sediment sorting and the maintenance of pool-riffle sequences in gravel-bedded streams. The literature in support of the hypothesis together with published adverse criticism is reviewed. It is concluded that convincing evidence for the ubiquitous occurrence of such a reversal in a range of channel geometries is currently unavailable. Continuity considerations indicate that riffles need to be considerably wider than pools for a reversal in the mean velocity to occur under conditions of subcritical flow, high stage, and stable morphology. These observations are substantiated by a detailed study of the hydraulic geometry of stable pool-riffle sequences in the River Severn, England. Neither the sectionally-averaged velocity nor the near-bed shear velocity is sensibly greater in the pools than over the riffles during bankfull or near bankfull flow. Instead a tendency towards equalization of the values of average hydraulic variables is noted as discharge increases. A detailed investigation of the three-dimensional character of the flow is required to demonstrate whether the entrainment forces within pools can locally exceed those over neighbouring riffles. Unusual behaviour of the energy gradient over riffles during moderate discharges is related to backwater effects as mediated by the spacing of the riffles. The hydraulic data are used to comment on the stability of the test reaches in the context of the development of the River Severn in the vicinity of Shrewsbury.  相似文献   

9.
A field‐based project was initiated in order to characterize velocities and sediment entrainment in a forced‐pool and riffle sequence. Three‐dimensional velocities and turbulence intensities were measured with an acoustic Doppler velocimeter at 222 different points at three similar flows that averaged approximately 4·35 m3 s−1 within a large pool–riffle unit on North Saint Vrain Creek, Colorado. Sediment‐sorting patterns were observed with the introduction of 500 tracer particles painted according to initial seeding location. Tracer particles moved sporadically during a 113 day period in response to the annual snowmelt peak flow, which reached a maximum level of 14·8 m3 s−1. Velocity data indicate high instantaneous velocities and turbulence levels in the centre of pools. Patterns of sediment deposition support the notion that stream competence is higher in the pool than the downstream riffle. Flow convergence around a large channel constriction appears to play a major role in multiple processes that include helical flow development and sediment routing, and backwater development with low velocities and turbulence levels above the constriction that may locally limit sediment supply. Jet flow, flow separation, vortex scour and turbulence generation enhance scour in the centre of pools. Ultimately, multiple processes appear to play some role in maintenance of this forced pool and the associated riffle. Copyright © 2008 John Wiley & Sons, Ltd.  相似文献   

10.
Experimental removal of woody debris from a small, gravel-bed stream in a forested area resulted in a four-fold increase in bedload transport at bankfull discharge. This was caused by increased transportability of sediment previously stored upslope of debris buttresses or in low-energy hydraulic environments related to debris. Bank erosion delivered additional sediment to the channel, and transport energy was increased by an inferred increase in the component of total boundary shear stress affecting grains on the bed. Increased transport following debris removal in May 1987 continued throughout the entire autumn storm season through late November 1987, indicating persistent adjustment of the stream bed and banks despite marked response to earlier flows as large as bankfull. Stream bed adjustments included development of a semi-regular sequence of alternate bars and pools, many of which were spaced independently of former pool locations.  相似文献   

11.
Computer flow simulations using the HEC-2 step-backwater routine are used to demonstrate the effect of systematically varying river channel width, riffle spacing and channel roughness on the shear velocity, section-mean velocity and energy slope in fixed-bed pool-riffle sequences. Initial scaling is obtained by utilizing published information on hydraulic parameters within reaches of the River Severn. Subsequently this restriction is relaxed and the effect of varying parameter combinations within realistic limits is explored. The purpose of this exercise is to isolate those scenarios which may preclude or promote the occurrence of a competence ‘reversal’, such that pools scour at high flow whilst deposition occurs on riffles. It is concluded that rivers in which pools are hydraulically rougher than riffles are likely to demonstrate a competence reversal. For prescribed conditions, the critical discharge at which a reversal occurs is a negative function of riffle spacing and riffle width relative to pool width. Downstream variation in hydraulic roughness also has implications for the phase relationship of shear velocity maxima and minima in relation to the extremes in pool-riffle topography.  相似文献   

12.
Studies on pool morphologies include reports of over 80% or 90% of pools being associated with structural controls and large obstructions that include boulders, bedrock outcrops and large woody debris (LWD). A Monte Carlo simulation approach and developmental computer model was created to predict pool formation, spacing and the percentage length covered by pools, riffles, scour holes and runs based on input data that include channel slope, width, the number of small and large boulders, and the number of 10–30 cm, 30–60 cm and >60 cm pieces of wood. The statistical‐empirical model is founded on the idea that boulders, bedrock outcrops and large woody debris provide a physical framework that then controls local water‐surface slopes, velocity patterns and the locations of pools and riffles. The spacing values of individual types and sizes of obstructions are modeled as log‐normal distributions with separate distributions for each obstruction type. Pools are assigned different probabilities of development depending on the obstruction type. Pool and riffle lengths used to create the subsequent morphology follow their own slope‐dependent, log‐normal trends. A minimum distance develops between successive pools because of the backwater and turbulent conditions needed for pool formation. The total number and spacing of pools, riffles and scour holes thus reflects the number and locations of obstructions and characteristics of the pool–riffle couplet. The simulation model accurately captures the number of pools in the modeled data range at 65% of all the verification field sites, and 86% of the verification field sites with a more limited range of width and slope characteristics. Lower levels of prediction capabilities are associated with modeled numbers of scour holes and log jams. The model accurately mimics some statistical attributes of pool spacing, and future versions of the model could be developed to improve overall predictive capabilities. Copyright © 2011 John Wiley & Sons, Ltd.  相似文献   

13.
Sediment grains in a bedrock‐alluvial river will be deposited within or adjacent to a sediment patch, or as isolated grains on the bedrock surface. Previous analysis of grain geometry has demonstrated that these arrangements produce significant differences in grain entrainment shear stress. However, this analysis neglected potential interactions between the sediment patches, local hydraulics and grain entrainment. We present a series of flume experiments that measure the influence of sediment patches on grain entrainment. The flume had a planar bed with roughness that was much smaller than the diameters of the mobile grains. In each experiment sediment was added either as individual grains or as a single sediment pulse. Flow was then increased until the sediment was entrained. Analysis of the experiments demonstrates that: (1) for individual grains, coarse grains are entrained at a higher discharge than fine grains; (2) once sediment patches are present, the different in entrainment discharge between coarse and fine grains is greatly reduced; (3) the sheltering effect of patches also increases the entrainment discharge of isolated grains; (4) entire sediment patches break‐up and are eroded quickly, rather than through progressive grain‐by‐grain erosion; (5) as discharge increases there is some tendency for patches to become more elongate and flow‐aligned, and more randomly distributed across the bed. One implication of this research is that the critical shear stress in bedrock‐alluvial channels will be a function of the extent of the sediment cover. Another is that the influence of sediment patches equalizes critical shear stresses between different grain sizes and grain locations, meaning that these factors may not need to be accounted for. Further research is needed to quantify interactions between sediment patches, grain entrainment and local hydraulics on rougher bedrock surfaces, and under different types of sediment supply. Copyright © 2016 John Wiley & Sons, Ltd.  相似文献   

14.
Past research investigated the surpassing of mean velocity at riffle cross sections by that at pool cross sections for flows up to bankfull, termed ‘velocity reversals’, to understand one mechanism by which riffle–pool relief is maintained. This study reenvisioned the classic velocity reversal by documenting stage‐dependent changes to the locations of peak velocity without cross sections. Instead, the dynamics of peak velocity patches were considered for flows spanning 0.2 to 22 times bankfull discharge through the use of a high‐resolution DEM and two‐dimensional hydrodynamic modelling. A remarkable diversity in peak velocity patch behaviour was found across discharges, including gradual expansion and shifting as well as abrupt disappearance and emergence relative to the low‐flow patch locations. These behaviours blended together to varying degrees to produce many reversals in peak velocity across morphological units, but it took substantially higher than bankfull discharge for peak velocities to move from riffles and chutes to fast glides and pools. The discharges at which reversals occurred among morphological units were significantly higher for the valley‐confined reach than for the anastomosing reach. Copyright © 2016 John Wiley & Sons, Ltd.  相似文献   

15.
Tian Zhou  Ted Endreny 《水文研究》2012,26(22):3378-3392
River restoration projects have installed j‐hook deflectors along the outer bank of meander bends to reduce hydraulic erosion, and in this study we use a computational fluid dynamics (CFD) model to document how these deflectors initiate changes in meander hydrodynamics. We validated the CFD with streamwise and cross‐channel bankfull velocities from a 193° meander bend flume (inlet at 0°) with a fixed point bar and pool equilibrium bed but no j‐hooks, and then used the CFD to simulate changes to flow initiated by bank‐attached boulder j‐hooks (1st attached at 70°, then a 2nd at 160°). At bankfull and half bankfull flow the j‐hooks flattened transverse water surface slopes, formed backwater pools upstream of the boulders, and steepened longitudinal water slopes across the boulders and in the conveyance region off the mid‐channel boulder tip. Streamwise velocity and mass transport jets upstream of the j‐hooks were stilled, mid‐channel jets were initiated in the conveyance region, eddies with a cross‐channel axis formed below boulders, and eddies with a vertical axis were shed into wake zones downstream of the point bar and outer bank boulders. At half bankfull depth conveyance region flow cut toward the outer bank downstream of the j‐hook boulders and the secondary circulation cells were reshaped. At bankfull depth the j‐hook at 160° was needed to redirect bank‐impinging flow sent by the upstream j‐hook. The hooked boulder tip of both j‐hooks funneled surface flow into mid‐channel plunging jets, which reversed the secondary circulation cells and initiated 1 to 3 counter rotating cells through the entire meander. The main outer bank collision zone centered at 50° without the j‐hook was moved by the j‐hook to within and just beyond the 70° j‐hook boulder region, which displaced other mass transport zones downstream. J‐hooks re‐organized water surface slopes, streamwise and cross‐channel velocities, and mass transport patterns, to move shear stress from the outer bank and into the conveyance and mid‐channel zones at bankfull flow. At half bankfull flows a patch of high shear re‐attached to the outer bank below the downstream j‐hook. J‐hook geometry and placement within natural meanders can be analyzed with CFD models to help restoration teams reach design goals and understand hydraulic impacts. Copyright © 2011 John Wiley & Sons, Ltd.  相似文献   

16.
This paper focuses on surface–subsurface water exchange in a steep coarse‐bedded stream with a step‐pool morphology. We use both flume experiments and numerical modelling to investigate the influence of stream discharge, channel slope and sediment hydraulic conductivity on hyporheic exchange. The model step‐pool reach, whose topography is scaled from a natural river, consists of three step‐pool units with 0.1‐m step heights, discharges ranging between base and over‐bankfull flows (scaled values of 0.3–4.5 l/s) and slopes of 4% and 8%. Results indicate that the deepest hyporheic flow occurs with the steeper slope and at moderate discharges and that downwelling fluxes at the base of steps are highest at the largest stream discharges. In contrast to findings in a pool‐riffle morphology, those in this study show that steep slopes cause deeper surface–subsurface exchanges than gentle slopes. Numerical simulation results show that the portion of the hyporheic zone influenced by surface water temperature increases with sediment hydraulic conductivity. These experiments and numerical simulations emphasize the importance of topography, sediment permeability and roughness elements along the channel surface in governing the locations and magnitude of downwelling fluxes and hyporheic exchange. Our results show that hyporheic zones in these steep streams are thicker than previously expected by extending the results from streams with pool‐riffle bed forms. Copyright © 2014 John Wiley & Sons, Ltd.  相似文献   

17.
Seventy-eight riffle to riffle and 80 bend spacings along eleven coarse-bedload, low sinuosity stream channels in upland Britain have been surveyed. Frequency distributions of these spacings are notably right-skewed. The most common repeating distances between riffles and bend inflections are between 4 and 6 channel widths although spacings up to 20 widths are also present. Riffle and pool locations around bends at different stages of planform development indicate that change is largely through increased sinuosity between two consecutive riffles of an original straight reach. Observed straight segments exhibit alternating riffles and pools evenly spaced at 4-6 widths, and most bends have similarly spaced riffles at their inflections in plan, with the intermediate pool at their apex. However angular deflections between axial lines joining inflections indicate existing sequences of bends did not develop from a single straight reach. Bends which are significantly longer than 4-6 widths are of low sinuosity and represent variability in naturally irregular planforms rather than arcs in advanced stages of meander growth. Adjusted bed topography around such long bends takes two forms. Either a single riffle-pool cycle is present with one or both bed forms being longer than average, or a 4-6 widths spacing is maintained by more than one riffle-pool sequence. Locally, the cross-section characteristics of riffles and pools are also influenced by planform location.  相似文献   

18.
A simple analytic model is presented relating local sediment transport capacity to variance in the transverse shear stress distribution in a stream channel. The model is used to develop a physically based conceptual model for the initiation of meandering in straight, bedload‐dominated streams as a result of a feedback mechanism. The feedback maximizes the cross‐sectional shear stress variance and – in order to achieve stability – ultimately minimizes the energy slope at repeated locations along the channel, subject to steady‐state mass flux and the stability of the channel boundary. These locations develop into pools in a fully developed meandering channel; they represent attractor states wherein sediment continuity is satisfied using the least possible energy expenditure per unit length of channel. However, since the cross‐sectional geometry of a pool (and the adjacent bar) is asymmetric, these attractor states are only conditionally stable, requiring strong, curvature‐induced secondary circulation to maintain their asymmetry. Between two successive pools, a stream occupies a metastable, higher energy state (corresponding to a riffle) that requires greater energy expenditure per unit length of channel to transport the same volume of sediment. The model we present links processes at the scale of a channel width to adjustments of the channel sinuosity and slope at the scale of a channel reach. We argue that the reach‐scale extremal hypotheses employed by rational regime models are mathematical formalisms that permit a one‐dimensional theory to describe the three‐dimensional dynamics producing stream morphology. Our model is consistent with the results from stream table experiments, with respect to both the rate of development of meandering and the characteristics of the equilibrium channel morphology. Copyright © 2006 John Wiley & Sons, Ltd.  相似文献   

19.
David Milan 《水文研究》2017,31(12):2179-2195
Fine sediment infiltration into gravel interstices is known to be detrimental to incubating salmonid embryos. Infiltration into spawning riffles can show large spatial variations at the scale of a morphological unit and over time, with significant implications for embryo survival. Furthermore, some process‐based infiltration studies, and incubation‐to‐emergence models assume that fines are delivered to redds via suspension rather than bedload. This process‐based 12‐month study examined spatial patterns of predominantly sand infiltration into gravels in an upland trout stream, using infiltration baskets. An assessment of Rouse numbers for infiltrated sand indicated that it was transported predominantly as bedload at flow peaks. Clear temporal and spatial patterns existed, with highest rates of infiltration strongly associated with higher discharges (r2 = 0.7, p < .05). Seasonal variations in the delivery of different grain sizes were also a feature, with enhanced contributions of 0.5–2 mm sediment during elevated winter flows and 0.125–0.5 mm sediment during spring and summer; the latter is potentially harmful to the later stages of embryo incubation. Clear spatial patterns were also evident across riffles, with highest rates of infiltration tending to occur in areas of lower relative roughness—the areas competent to transport sand for longer periods. Incubation‐to‐emergence models should take into consideration spatial patterns of fine sediment dynamics at the pool–riffle scale, to improve prediction.  相似文献   

20.
Drought and summer drying can be important disturbance events in many small streams leading to intermittent or isolated habitats. We examined what habitats act as refuges for fishes during summer drying, hypothesizing that pools would act as refuge habitats. We predicted that during drying fish would show directional movement into pools from riffle habitats, survival rates would be greater in pools than in riffles, and fish abundance would increase in pool habitats. We examined movement, survival and abundance of three minnow species, bigeye shiner (Notropis boops), highland stoneroller (Campostoma spadiceum) and creek chub (Semotilus atromaculatus), during seasonal stream drying in an Ozark stream using a closed robust multi-strata mark-recapture sampling. Population parameters were estimated using plausible models within program MARK, where a priori models are ranked using Akaike’s Information Criterion. Creek chub showed directional movement into pools and increased survival and abundance in pools during drying. Highland stonerollers showed strong directional movement into pools and abundance increased in pools during drying, but survival rates were not significantly greater in pools than riffles. Bigeye shiners showed high movement rates during drying, but the movement was non-directional, and survival rates were greater in riffles than pools. Therefore, creek chub supported our hypothesis and pools appear to act as refuge habitats for this species, whereas highland stonerollers partly supported the hypothesis and bigeye shiners did not support the pool refuge hypothesis. Refuge habitats during drying are species dependent. An urgent need exists to further understand refuge habitats in streams given projected changes in climate and continued alteration of hydrological regimes.  相似文献   

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