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1.
Last interglacial sediments in unglaciated Alaska and Yukon (eastern Beringia) are commonly identified by palaeoecological indicators and stratigraphic position ~2–5 m above the regionally prominent Old Crow tephra (124 ± 10 ka). We demonstrate that this approach can yield erroneous age assignments using data from a new exposure at the Palisades, a site in interior Alaska with numerous exposures of last interglacial sediments. Tephrochronology, stratigraphy, plant macrofossils, pollen and fossil insects from a prominent wood‐rich organic silt unit are all consistent with a last interglacial age assignment. However, six 14C dates on plant and insect macrofossils from the organic silt range from non‐finite to 4.0 14C ka BP, indicating that the organic silt instead represents a Holocene deposit with a mixed‐age assemblage of organic material. In contrast, wood samples from presumed last interglacial organic‐rich sediments elsewhere at the Palisades, in a similar stratigraphic position with respect to Old Crow tephra, yield non‐finite 14C ages. Given that local permafrost thaw since the last interglaciation may facilitate reworking of older sediments into new stratigraphic positions, minimum constraining ages based on 14C dating or other methods should supplement age assignments for last interglacial sediments in eastern Beringia that are based on palaeoecology and stratigraphic association with Old Crow tephra. Copyright © 2011 John Wiley & Sons, Ltd.  相似文献   

2.
Steppe‐tundra is considered to have been a dominant ecosystem across northern Eurasia during the Last Glacial Maximum. As the fossil record is insufficient for understanding the ecology of this vanished ecosystem, modern analogues have been sought, especially in Beringia. However, Beringian ecosystems are probably not the best analogues for more southern variants of the full‐glacial steppe‐tundra because they lack many plant and animal species of temperate steppes found in the full‐glacial fossil record from various areas of Europe and Siberia. We present new data on flora, land snails and mammals and characterize the ecology of a close modern analogue of the full‐glacial steppe‐tundra ecosystem in the southeastern Russian Altai Mountains, southern Siberia. The Altaian steppe‐tundra is a landscape mosaic of different habitat types including steppe, mesic and wet grasslands, shrubby tundra, riparian scrub, and patches of open woodland at moister sites. Habitat distribution, species diversity, primary productivity and nutrient content in plant biomass reflect precipitation patterns across a broader area and the topography‐dependent distribution of soil moisture across smaller landscape sections. Plant and snail species considered as glacial relicts occur in most habitats of the Altaian steppe‐tundra, but snails avoid the driest types of steppe. A diverse community of mammals, including many species typical of the full‐glacial ecosystems, also occurs there. Insights from the Altaian steppe‐tundra suggest that the full‐glacial steppe‐tundra was a heterogeneous mosaic of different habitats depending on landscape‐scale moisture gradients. Primary productivity of this habitat mosaic combined with shallow snow cover that facilitated winter grazing was sufficient to sustain rich communities of large herbivores.  相似文献   

3.
Botanical analyses of fossil and modern arctic ground squirrel (Urocitellus parryii) caches and nests have been used to reconstruct the past vegetation from some parts of Beringia, but such archives are understudied in Alaska. Five modern and four fossil samples from arctic ground squirrel caches and nests provide information on late Pleistocene vegetation in Eastern Beringia. Modern arctic ground squirrel caches from Alaska's arctic tundra were dominated by willow and grass leaves and grass seeds and bearberries, which were widespread in the local vegetation as confirmed by vegetation surveys. Late Pleistocene caches from Interior Alaska were primarily composed of steppe and dry tundra graminoid and herb seeds. Graminoid cuticle analysis of fossil leaves identified Calamagrostis canadensis, Koeleria sp. and Carex albonigra as being common in the fossil samples. Stable carbon isotopes analysis of these graminoid specimens indicated that plants using the C3 photosynthetic pathways were present and functioning with medium to high water-use efficiency. Fossil plant taxa and environments from ground squirrel caches in Alaska are similar to other macrofossil assemblages from the Yukon Territory, which supports the existence of a widespread mammoth steppe ecosystem type in Eastern Beringia that persisted throughout much of the late Pleistocene.  相似文献   

4.
The lithostratigraphy and biostratigraphy of two sites (Allt Odhar and Dalcharn) in north-central Scotland are described, where pollen spectra of temperate affinity have been obtained from organic deposits that underlie till. The pollen record from Allt Odhar, in association with evidence from plant macrofossils and Coleoptera, shows the expansion of birch woodland and its eventual replacement by open grassland under a climatic regime slightly cooler than that prevailing in the northern highlands of Scotland at the present day. The organic sediments accumulated during an Early Devensian interstadial episode, which has been dated by the uranium series disequilibrium method to ca. 106 ka BP. Evidence for one and possibly two Devensian glaciations may be preserved at the site. The pollen record from Dalcharn, by contrast, reflects the middle and later stages of an interglacial cycle with the transition from pine forest to grassland. The overlying till sequence contains evidence of at least two separate glacial episodes. The age of the warm stage cannot be established precisely on present evidence, but there are indications that it may predate the last (Ipswichian) interglacial. These are the first sites from the mainland of Scotland to provide evidence of wooded conditions during interstadial and interglacial episodes of the Middle/Late Pleistocene.  相似文献   

5.
A 14-m-thick section of marine and nonmarine sediments of the Gubik Formation of northern Alaska, exposed in bluffs near Ocean Point on the Colville River, has been studied by means of pollen analysis. Pollen from the marine sediments, of probable late Pliocene age, records a boreal forest of spruce and birch with minor amounts of alder in the adjacent terrestrial vegetation. Pine and perhaps true fir were probably at or near their northern limit here, but hemlocks and hardwoods were absent. The suggested environment for the Arctic Slope during the time represented by the marine sediments is similar to that of present-day Anchorage. Pollen floras from the overlying fluvial strata, of early or middle Pleistocene age, record predominantly herbaceous taxa indicating tundra conditions probably more severe than those of the present day. These deposits were most likely contemporaneous with glacial conditions in the Brooks Range to the south. Pollen of woody taxa (spruce, alder, birch, heaths) is rare through most of the section, although birch and alder percentages similar to those found in modern river sediments indicate an interstadial or interglacial warming in midsection. Inland climates during glacial episodes may have been similar to those of the present Arctic coast.  相似文献   

6.
During the Quaternary glacial episodes, when sea level was considerably lower, Asia and North America were linked by large extensions of circumarctic land (Beringia), which remained unglaciated. This land mass served not only as a biogeographical bridge for plants, animals, and humans, but also supported a biome very different from present tundra or boreal coniferous forests, which was dominated by steppes and a rich mammalian megafauna. Carbon stable isotope ratios of Beringian late Pleistocene mummified remains of bison, equids, mammoth, caribou, musk-ox, moose, woolly rhino, and other undetermined species, found preserved in permafrost, indicate that these megaherbivores fed exclusively on C3 plants, and that C4 grasses were not differentially ingested by bison, as previously suggested. Paleoclimatic constraints probably prevented the formation of a warm-season (C4) guild during the later part of the growing season in the steppes of Beringia during the last glaciation.  相似文献   

7.
《Quaternary Science Reviews》2003,22(18-19):1947-1986
Loess is one of the most widespread subaerial deposits in Alaska and adjacent Yukon Territory and may have a history that goes back 3 Ma. Based on mineralogy and major and trace element chemistry, central Alaskan loess has a composition that is distinctive from other loess bodies of the world, although it is quartz-dominated. Central Alaskan loess was probably derived from a variety of rock types, including granites, metabasalts and schists. Detailed stratigraphic data and pedologic criteria indicate that, contrary to early studies, many palaeosols are present in central Alaskan loess sections. The buried soils indicate that loess sedimentation was episodic, or at least rates of deposition decreased to the point where pedogenesis could keep ahead of aeolian input. As in China, loess deposition and pedogenesis are likely competing processes and neither stops completely during either phase of the loess/soil formation cycle. Loess deposition in central Alaska took place before, and probably during the last interglacial period, during stadials of the mid-Wisconsin period, during the last glacial period and during the Holocene. An unexpected result of our geochronological studies is that only moderate loess deposition took place during the last glacial period. Our studies lead us to conclude that vegetation plays a key role in loess accumulation in Alaska. Factors favouring loess production are enhanced during glacial periods but factors that favour loess accumulation are diminished during glacial periods. The most important of these is vegetation; boreal forest serves as an effective loess trap, but sparsely distributed herb tundra does not. Thus, thick accumulations of loess should not be expected where tundra vegetation was dominant and this is borne out by modern studies near the treeline in central Alaska. Much of the stratigraphic diversity of North American loess, including that found in the Central Lowlands, the Great Plains, and Alaska is explained by a new model that emphasizes the relative importance of loess production factors versus loess accumulation factors.  相似文献   

8.
Fossil arctic ground squirrel (Spermophilus parryii) middens were recovered from ice-rich loess sediments in association with Sheep Creek-Klondike and Dominion Creek tephras (ca 80 ka) exposed in west-central Yukon. These middens provide plant and insect macrofossil evidence for a steppe-tundra ecosystem during the Early Wisconsinan (MIS 4) glacial interval. Midden plant and insect macrofossil data are compared with those previously published for Late Wisconsinan middens dating to ~25–2914C ka BP (MIS 3/2) from the region. Although multivariate statistical comparisons suggest differences between the relative abundances of plant macrofossils, the co-occurrence of steppe-tundra plants and insects (e.g., Elymus trachycaulus, Kobresia myosuroides, Artemisia frigida, Phlox hoodii, Connatichela artemisiae) provides evidence for successive reestablishment of the zonal steppe-tundra habitats during cold stages of the Late Pleistocene. Arctic ground squirrels were well adapted to the cold, arid climates, steppe-tundra vegetation and well-drained loessal soils that characterize cold stages of Late Pleistocene Beringia. These glacial conditions enabled arctic ground squirrel populations to expand their range to the interior regions of Alaska and Yukon, including the Klondike, where they are absent today. Arctic ground squirrels have endured numerous Quaternary climate oscillations by retracting populations to disjunct “interglacial refugia” during warm interglacial periods (e.g., south-facing steppe slopes, well-drained arctic and alpine tundra areas) and expanding their distribution across the mammoth-steppe biome during cold, arid glacial intervals.  相似文献   

9.
Although there is evidence from molecular studies for the existence of central European last glacial refugia for temperate species, there is still a great lack of direct fossil records to confirm this theory. Here we bring such evidence in the form of fossil shells from twenty strictly forest land snail species, which were recorded in radiocarbon-dated late glacial or older mollusc assemblages of nine non-interrupted mollusc successions situated in the Western Carpathians, and one in the Bohemian Massif. We proposed that molluscs survived the last glacial period in central Europe in isolated small patches of broadleaf forest, which we unequivocally demonstrate for two sites of last glacial maximum age.  相似文献   

10.
Rodent middens from ice-rich loess deposits are important new paleoenvironmental archives for Eastern Beringia. Plant macrofossils recovered from three middens associated with Dawson tephra (ca. 24,000 14C yr B.P.) at two sites in Yukon Territory include diverse graminoids, forbs, and mosses. These data suggest substantial local scale floristic and habitat diversity in valley settings, including steppe-tundra on well-drained soils, moist streamside meadows, and hydric habitats. Fossil arctic ground squirrel burrows and nesting sites indicate that permafrost active layers were thicker during Pleistocene glacial periods than at present on north-facing slopes.  相似文献   

11.
Studies of invertebrates from steppe patches in the tundra and taiga zones of Beringia provide additional evidence that these areas could be relict steppes. A number of insect species common to both modern relict steppes and fossil Beringian insect faunal assemblages have been found. These provide important information on the moisture and temperature preferences of some of the surviving members of Pleistocene steppe-tundra insect communities. The most significant species of West Beringian insects are weevils in the genus Stephanocleonus (Coleoptera, Curculionidae), indicators of thermophytic steppe, and the pill beetle Morychus viridis (Coleoptera, Byrrhidae), the indicator of hemicryophytic steppe. The East Beringian invertebrate population of relict steppe is substantially different. Fossil evidence suggests that biotic exchange between the two parts of Beringia was limited during the Pleistocene; populations of steppe insects did not move across the Bering Land Bridge (BLB), while tundra species had more flexibility. The tundra environment reconstructed for the Pleistocene BLB should have facilitated amphi-beringian distributions for most tundra invertebrate species, but apparently only a few species achieved this.  相似文献   

12.
Pollen analysis of a sediment core from Zagoskin Lake on St. Michael Island, northeast Bering Sea, provides a history of vegetation and climate for the central Bering land bridge and adjacent western Alaska for the past ≥30,000 14C yr B.P. During the late middle Wisconsin interstadial (≥30,000-26,000 14C yr B.P.) vegetation was dominated by graminoid-herb tundra with willows (Salix) and minor dwarf birch (Betula nana) and Ericales. During the late Wisconsin glacial interval (26,000-15,000 14C yr B.P.) vegetation was graminoid-herb tundra with willows, but with fewer dwarf birch and Ericales, and more herb types associated with dry habitats and disturbed soils. Grasses (Poaceae) dominated during the peak of this glacial interval. Graminoid-herb tundra suggests that central Beringia had a cold, arid climate from ≥30,000 to 15,000 14C yr B.P. Between 15,000 and 13,000 14C yr B.P., birch shrub-Ericales-sedge-moss tundra began to spread rapidly across the land bridge and Alaska. This major vegetation change suggests moister, warmer summer climates and deeper winter snows. A brief invasion of Populus (poplar, aspen) occurred ca.11,000-9500 14C yr B.P., overlapping with the Younger Dryas interval of dry, cooler(?) climate. During the latest Wisconsin to middle Holocene the Bering land bridge was flooded by rising seas. Alder shrubs (Alnus crispa) colonized the St. Michael Island area ca. 8000 14C yr B.P. Boreal forests dominated by spruce (Picea) spread from interior Alaska into the eastern Norton Sound area in middle Holocene time, but have not spread as far west as St. Michael Island.  相似文献   

13.
In 1980 a large proboscidean femur, probably Mammuthus sp., was found in situ in a bluff exposure at the mouth of the Tyone River in the northwestern part of the Copper River Basin, Alaska. The regional setting, stratigraphy, radiocarbon chronology, flora, and implications of the fossil locality, which represents the first documented occurrence of Pleistocene terrestrial mammalian fauna in southern Alaska, are described. Radiocarbon dates and stratigraphic relations at the site indicate that the sediments containing the fossil accumulated during the transition from interstadial to glacial conditions during terminal middle Wisconsin time. During this interval the immediate vicinity was unforested and large areas of south-central Alaska may have been available for faunal and possibly human habitation. This documented find, dated at 29,450 ± 610 14C yr B.P., extends the known range for Pleistocene mammals and possibly steppe-tundra conditions south-ward at least 150 km, and suggests that mountain passes through the Alaska Range to the north were ice free during the last part of the middle Wisconsin interstadial.  相似文献   

14.
The development of a glacial lake impounded along the retreating, northeastern ice margin of the Fennoscandian Ice Sheet during the last deglaciation and environmental conditions directly following the early Holocene deglaciation have been studied in NE Finland. This so‐called Sokli Ice Lake has been reconstructed previously using topographic and geomorphologic evidence. In this paper a multiproxy approach is employed to study a 3‐m‐thick sediment succession consisting of laminated silts grading into gyttja cored in Lake Loitsana, a remnant of the Sokli Ice Lake. Variations in the sediment and siliceous microfossil records indicate distinct changes in water depth and lake size in the Loitsana basin as the Sokli Ice Lake was drained through various spillways opening up along the retreating ice front. Geochemical data (XRF core‐scanning) show changes in the influence of regional catchment geochemistry (Precambrian crystalline rocks) in the glacial lake drainage area versus local catchment geochemistry (Sokli Carbonatite Massif) within the Lake Loitsana drainage area during the lake evolution. Principal component analysis on the geochemical data further suggests that grain‐size is an additional factor responsible for the variability of the sediment geochemistry record. The trophic state of the lake changed drastically as a result of morphometric eutrophication once the glacial lake developed into Lake Loitsana. The AMS radiocarbon dating on tree birch seeds found in the glaciolacustrine sediment indicates that Lake Loitsana was deglaciated sometime prior to 10 700 cal. a BP showing that tree Betula was present on the deglaciated land surrounding the glacial lake. Although glacial lakes covered large areas of northern Finland during the last deglaciation, only few glaciolacustrine sediment successions have been studied in any detail. Our study shows the potential of these sediments for multiproxy analysis and contributes to the reconstruction of environmental conditions in NE Finland directly following deglaciation in the early Holocene.  相似文献   

15.
Studies of interglacial successions are critical to our understanding of the environmental history of an area. Analyses of macrofossil remains of plants and invertebrates from Eemian sediments exposed in a coastal cliff section at Borðoyarvík near Klaksvík, Bordoy, northeastern Faroe Islands, indicate that the sediments accumulated in a coastal lagoon. The fossil flora comprises tree birch Betula sect. Albae and we suggest that birch forests were found locally at sheltered sites in the area. Tree birch also occurred on the islands during the mid‐Holocene. The only other woody plant recovered from the Eemian deposit is the dwarf‐shrub Empetrum nigrum, which is common on the islands today. Remains of herbaceous plants are rare but include Viola, Ajuga, Myosotis, Urtica dioica and Ranunculus. The bryophyte flora is species‐rich and most of the fragments belong either to stream species or to species of humid or wet habitats. The fossil flora and fauna also comprise a number of freshwater species that probably lived in an oligotrophic lake and in streams in the catchment of the lagoon. The climate during deposition of the lagoonal sediments was similar to the Holocene oceanic climate of the Faroe Islands. The study adds to our understanding of Eemian environments in the North Atlantic region and helps to fill a knowledge gap about the history of the flora and fauna of the Faroe Islands, which is of biogeographical importance.  相似文献   

16.
Western camel (C. hesternus) fossils are rare from Eastern Beringia, thus there is little available information on their chronology, paleoecology, and biogeography in this region. In August of 2010, a partial proximal phalanx of a western camel was recovered from a sedimentary exposure along the White River, in the formerly glaciated terrain of southwest Yukon, northwest Canada. The fossil specimen was recovered in situ from sediments that are correlated by stratigraphic, tephra and radiocarbon data to the Marine Isotope Stage (MIS) 5 interglacial period (Sangamonian). Associated paleoenvironmental data indicates that this western camel inhabited a shrub tundra ecosystem that did not include spruce trees or boreal forest during a relatively cold interval between MIS 5e and 5a. This is the oldest reliably dated western camel fossil from Eastern Beringia and supports the model of range expansion for this species to the high latitudes of northwest North America during the last interglacial (sensu lato).  相似文献   

17.
Fossils of forest habitat beetles and leaves of Nothofagus menziesii provide evidence of a forest refugium at times between ca. 34 000 and ca. 18 500 cal. a BP at an upland site in Howard Valley, located adjacent to glaciated valleys in South Island, New Zealand. The stratigraphy of the glacial‐aged terrace sequence of organic‐rich silts and fluvial sand/gravels indicates that soil development occurred episodically for around 15 000 a. Fifty‐four beetle taxa represent seven habitat types: forest, forest or scrub, riparian and aquatic, litter, grass/tussock, marshland and moss habitats. Leaf and beetle fossils indicate that forest dominated by N. menziesii persisted at the site for most of the time period represented, and tree line taxa such as Taenarthrus sp. 1 (Carabidae) and Podocarpus sp. (Podocarpaceae) indicate that the site may represent the upper tree limit for full‐glacial time. The finding of forest at this elevated site adds to the growing fossil evidence for multiple forest refugia in New Zealand during the last glaciation and is consistent with the pollen records, which have consistently indicated the presence of forest species during the last glaciations. Copyright © 2009 John Wiley & Sons, Ltd.  相似文献   

18.
We present a multi‐proxy reconstruction from a well‐preserved vegetation surface (ca. 32 000 14C a BP) from the Fox Permafrost tunnel near Fairbanks, Alaska. A thick litter layer of plant material on the vegetation surface is consistent with the vegetation lacking evidence of disturbance. Plant macrofossils and graminoid cuticle analysis show the presence of a graminoid assemblage consistent with phytolith data. The pollen data indicate that trees were not local to the site and that Artemisia sp. was present in the region. The insect and bryophyte reconstructions are consistent with the vascular plant reconstruction, indicating the site was at least periodically wet. δ13C values from the graminoids present show a large range encompassing both the wet and dry range displayed by modern graminoids in Alaska. Sequential δ13C analyses conducted along the length of leaves attached to the vegetation surface indicate a seasonal shift towards relatively higher water use efficiency. The lower water use efficiency earlier in the growing season may have stemmed from the use of winter season meltwater by plants at the site – a scenario consistent with the site's cryostratigraphy. Our multi‐proxy reconstruction contributes to the limited palaeoecological data available for graminoid‐dominated vegetation present in Eastern Beringia and particularly the interior of Alaska during the mid‐Wisconsinan interstadial. Copyright © 2011 John Wiley & Sons, Ltd.  相似文献   

19.
Improved chronological control on the penultimate advance of the Cordilleran Ice Sheet in northwest Canada (the Reid glaciation) is required for a better understanding of late Quaternary palaeoclimatic and palaeoenvironmental change in eastern Beringia. However, reliable dating of glaciation events beyond the last glacial maximum is commonly hindered by a lack of directly dateable material. In this study we (i) provide the first combined minimum and maximum age constraint on the Reid glaciation at Ash Bend, its reference locale in the Stewart River valley, northwestern Canadian Cordillera, using single-grain optically stimulated luminescence dating of quartz; and (ii) compare the timing of the Reid glaciation with other penultimate ice sheet advances in the region with the aim of establishing improved glacial reconstructions in eastern Beringia. We obtain ages of 158 ± 18 ka and 132 ± 18 ka for glaciofluvial sands overlying and underlying the Reid till, respectively. These ages indicate that the Reid advance, at its reference locale, occurred during MIS 6. This precludes an earlier MIS 8 age, and suggests that the Reid advance may have been synchronous with the Delta glaciation of central Alaska, and is likely correlative with the Mirror Creek glaciation in southern Yukon.  相似文献   

20.
We present 10Be exposure ages from moraines in the Delta River Valley, a reference locality for Pleistocene glaciation in the northern Alaska Range. The ages are from material deposited during the Delta and Donnelly glaciations, which have been correlated with MIS 6 and 2, respectively. 10Be chronology indicates that at least part of the Delta moraine stabilized during MIS 4/3, and that the Donnelly moraine stabilized ∼ 17 ka. These ages correlate with other dates from the Alaska Range and other regions in Alaska, suggesting synchronicity across Beringia during pulses of late Pleistocene glaciation. Several sample types were collected: boulders, single clasts, and gravel samples (amalgamated small clasts) from around boulders as well as from surfaces devoid of boulders. Comparing 10Be ages of these sample types reveals the influence of pre/post-depositional processes, including boulder erosion, boulder exhumation, and moraine surface lowering. These processes occur continuously but seem to accelerate during and immediately after successive glacial episodes. The result is a multi-peak age distribution indicating that once a moraine persists through subsequent glaciations the chronological significance of cosmogenic ages derived from samples collected on that moraine diminishes significantly. The absence of Holocene ages implies relatively minor exhumation and/or weathering since 12 ka.  相似文献   

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