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1.
Exposures of upper Paleozoic slates of the metamorphic basement near Concepción, central Chile, are covered by transgressive deposits belonging to the Upper Cretaceous Quiriquina Formation. Presence of clusters of the bivalve boring Gastrochaenolites isp. at the irregular and erosive contact between these two units indicates the development of an ancient rocky shoreline, illustrating a rare example of bioerosion in a metamorphic substrate. Coarse-grained deposits mantling the bioeroded surface represent a transgressive lag produced due to ravinement. In sequence-stratigraphic parlance, the bioeroded surface is classified as a FS/SB or co-planar surface formed due to amalgamation of erosion during lowstand and the subsequent transgression. Unburrowed trough cross-bedded upper-shoreface to intensely bioturbated middle-shoreface deposits record continuous transgression. Logs bioeroded by Teredolites clavatus are also present. Middle-shoreface deposits are dominated by deep Ophiomorpha isp., commonly showing laminated infill. Thalassinoides suevicus occurs locally. Intense bioturbation also suggests that the shoreface developed in embayed shorelines, protected from oceanic waves. While rocky shorelines in limestone are characterized by a high abundance and diversity of bioerosion structures, those formed in other types of substrates contain less diverse, commonly monospecific, suites of borings. In terms of Seilacherian ichnofacies, the bioerosion structures analyzed are attributed to a low-diversity expression of the Trypanites ichnofacies. It is proposed that the Trypanites ichnofacies thus may display two expressions: an archetypal one characterized by high diversity in carbonate substrates, along with a depauperate expression in other types of substrates (e.g., metamorphic and igneous rocks). The extreme hardness of the substrate is regarded as the stress factor responsible for the reduction in ichnodiversity.  相似文献   

2.
Trace fossils provide detailed palaeoenvironmental and palaeoecological information of both ancient and modern sedimentary systems. During middle Miocene times the Aures Massif located in the northeastern part of Algeria, was affected by, at least, one marine transgression. The latter led to the installation of a carbonate platform, which is placed, for the first time, in a Mediterranean context. In the Rhassira basin, the Middle Miocene marine succession is characterised by carbonate platform deposits dominated by rhodolith beds, typical of those known throughout the Mediterranean area. This succession can be divided into many units separated by discontinuities interpreted here as omission surfaces. The Djebel Arhane section shows two omission surfaces characterised by a pre-omission suite (firmground) represented by Balanoglossites burrows for the first surface and Gastrochaenolites ornatus burrows/borings for the second one, and an omission suite (hardground) as evidenced by the bioerosive structures Trypanites and Caulostrepsis, in both surfaces, respectively. Gastrochaenolites ornatus traces were formed and preserved in firm, compact, semi-lithified and fine-grained substrates (firm- to hardground), indicating the Glossifungites ichnofacies. They show bioglyphs which have been formed during contraction of the posterior adductor muscles. These suggest that their tracemakers were represented by suspension-feeding bivalves, most probably Pholadidae or Mytilidae, which rotated during penetration. The fill of these traces is composed of marine deposits related to a transgressive lag. The omission suite is divided into two ichnocoenoses: (i) pre-lithification burrows/borings, and (ii) post-lithification borings. This is the first report of the ichnotaxon G. ornatus from Algeria.  相似文献   

3.
The Callovian-Oxfordian Kuldhar Member of Middle-Upper Jurassic Jaisalmer Formation, Rajasthan is characterized by a rich cephalopod (ammonites and belemnites) fauna along with other invertebrates. They are embedded in oolitic limestone, at places in nodular limestone, grey shale and occasional red mud. The belemnite rostra bear bioerosions in the form of borings of circular or elliptical type. The present study identified those as Trypanites isp. and Rogerella isp. and characterized the hitherto unnoticed bioerosions of these belemnite rostra to be post-mortal. Different dispositions of major diameter of belemnite rostra has been interpreted to represent churning effect of ichnofossil producing organisms which put evidences of several generations of omission surfaces. Vertical and horizontal orientations of stephanoceratid ammonites side by side have been critically analysed in the light of lithology and prevailing environmental condition and inferred to be an artifact of gravity and thixotropic property of clay. Ammonites like Stephanoceratids and Macrocephalitids inhabit within 110m depth which is also permissible depth of belemnites as well as probable depth of formation of oolites, carbonate mud and aluminosilicate mud. Considering the lithology of the rocks like oolites, carbonate mud, aluminosilicate, the habitat of stephanoceratid and habitat of acrothoracid barnacles (organisms responsible for Rogerella isp.), the environment has been delineated to be sublittoral which was previously designated as shoreface zone to offshore transition zone.  相似文献   

4.
The litho- and biostratigraphy of the Craie de Villedieu Formation (Coniacian-Santonian)of western France are described in detail. The formation is subdivided into three members each containing a number of lithologically distinct named hardgrounds and marker beds. These constitute an onlapping sequence that thins from > 15 m in the NE around Cangey and Villedieu-le-Château, to < 2 m in the SW around St Michel-sur-Loire, a distance of 70 km. Thickness variation is related to the interaction of differential subsidence with eustatic transgression. Comparison with the Chalk Rock Formation of southern England indicates that transgressive and regressive hardground suites may be differentiated on bed geometry and hardground surface characteristics. The Craie de Villedieu rests everywhere on a regional hardground that coincides with the Turonian/Coniacian boundary in expanded successions, but probably marks a significant hiatus. South-west of Tours, onlap results in Santonian strata resting disconformably on strata of Turonian age. The basal Craie de Villedieu contains a succession of three Coniacian ammonite faunas characterized by Peroniceras and Forresteria (Harleites) (oldest), Gauthiericeras margae (Schlüter), and Protexanites (youngest). Volviceramus ex gr. involutus (J. de C. Sowerby) occurs with the two uppermost ammonite assemblages. A Santonian ammonite fauna dominated by Placenticeras polyopsis (Dujardin) occurs with Texanites gallicus Collignon and common Cladoceramus in the middle of the formation. Cordiceramus ex gr. cordiformis (J. de C. Sowerby) is recorded with Santonian ammonites in the upper part of the formation. A correlation with the Micraster zones of chalk facies is suggested, based on the inoceramid stratigraphy. The record of T. gallicus in association with Cladoceramus affords the first direct evidence for the position of the base of the Santonian in the Anglo-Paris Basin.  相似文献   

5.
New finds reveal that cases of larval caddisflies (indusifauna) are widespread aquatic domichnia in Eurasian non-marine deposits. 16 new ichnospecies are described in 6 ichnogenera from the Wealden Supergroup (Lower Cretaceous) of southeast England, mainly from the Ashdown Formation (Valanginian), but some also from the Weald Clay Group (Hauterivian–Barremian). New ispp. are: Terrindusia valdensis isp. nov., T. anomala isp. nov.; Secrindusia sarahae isp. nov.; Conchindusia dianae isp. nov., C. elderi isp. nov., C. goodmani isp. nov.; Pelindusia duprati isp. nov.; ?Ostracindusia mixta isp. nov.; Folindusia stouti isp. nov., F. bipennis isp. nov., F. ruffordi isp. nov., F. chiasma isp. nov., F. woodhamsi isp. nov., F. boothi isp. nov., F. avancnae isp. nov., and F. whitei isp. nov. Two new forms, a variety and aberration of Conchindusia (an igen. with no modern analogue) are also recognised. The first Chinese isp. (Conchindusia sinensis isp. nov.) is described from the Yixian Formation (Aptian) and 31 morphotypes are listed from eight Lower Cretaceous Chinese formations.The aquatic palaeoenvironment and palaeoecology are discussed. The UK indusifauna is dominated by ConchindusiaFolindusia in contrast to a Terrindusia dominance in China; furthermore, the UK EPT (Ephemeroptera–Plecoptera–Trichoptera) richness is skewed towards Trichoptera. This reflects differences in the fluvio-lagoonal and fluvio-lacustrine settings, respectively.  相似文献   

6.
The robust calcite rostra (or guards) of belemnites were attractive substrates for some boring organisms in the Mesozoic. The rostra of belemnitellid coleoids are common at certain levels within the Gulpen and Maastricht formations of the extended type area of the Maastrichtian Stage (Upper Cretaceous) in the south-east Netherlands and north-east Belgium. A range of episkeletozoans have been reported from these substrates hitherto. A new specimen preserved as a natural mould in flint from the province of Liège, north-east Belgium, is distinctive, preserving a dense three-dimensional infestation of long, slender, unbranched and unsculptured borings of constant diameter, Trypanites? isp. These may be the spoor of polychaetes. Variations in diameter between borings suggests that the traces in this specimen may represent four separate infestations.  相似文献   

7.
The Marwar Supergroup of the Bikaner-Nagaur Basin is composed of sediments deposited from the late Neoproterozoic (Ediacaran) to Upper Cambrian. The Nagaur Sandstone Formation of the Nagaur Group (uppermost division of the Marwar Supergroup) preserves trace fossils significant for establishing Early Cambrian biostratigraphic zones and depositional facies. Fifteen ichnospecies (and eight ichnogenera) identified in the Nagaur Sandstone Formation include “Treptichnus” pedum, Cruziana cf. tenella, Cruziana isp., Diplichnites ispp. A, B, and C, Gyrophyllites isp., Lockeia isp., Merostomichnites isp., Monomorphichnus gregarius isp. nov., Monomorphichnus isp., Planolites isp., Psammichnites isp., Rusophycus bikanerus isp. nov., Rusophycus cf. carbonarius, Rusophycus isp. and radial trace fossils.These trace fossils belong to ethological categories pascichnia, repichnia, cubichnia, and fodinichnia and represent arthropod and worm-like burrowing biota. The assemblage and a regional comparison with contemporaneous trace fossils in the eastern Gondwanan realm suggest that the sequence in the study area belongs to the Cruziana tenella Ichnozone and to Stage 2 (upper part of Terreneuvian), however the Middle Cambrian is not excluded. The trace fossil assemblage belongs to the archetypal Cruziana ichnofacies. Cross bedded sandstone, mud cracks and rainprints in the ichniferous strata of the Nagaur Sandstone Formation indicate deposition in an intertidal sand flat with channels that was exposed episodically.  相似文献   

8.
The Upper Emsian to Frasnian Ia-Ib strata of the Marhouma area (or “km 30” outcrop), exposed in the Ougarta Range (SW Algeria) belong to the Chefar El Ahmar Formation. On the basis of distinct lithological and palaeontological features, this formation is subdivided into three members (Lower Marly Limestones Member, Middle Marly Limestones Member, and Upper Marly Limestones Member). The studied beds show low to moderate diversity of trace fossil assemblage which contains thirteen ichnotaxa: Chondrites intricatus, Chondrites isp., Chondrites cf. targionii, Circulichnis cf. montanus, Cochlichnus isp., Neonereites biserialis, Neonereites multiserialis, Nereites isp., Palaeophycus isp., Planolites isp., Thalassinoides isp., Zoophycos aff. cauda-galli, and Zoophycos isp. A. The two latter ichnotaxa are the most common trace fossils in the assemblage and occur at three different levels showing different bioturbation intensities. The first Zoophycos-bearing level (Zl 1) is characterised by an overall high bioturbation intensity reflecting a very high oxygenation rate and nutrient supply, allowing the development of large and dense Zoophycos specimens. The second Zoophycos-bearing level (Zl 2) has a considerable reduction of bioturbation intensity as compared to the previous level, with an abundance of Chondrites, which is probably due to radical palaeoecological changes that suggests dysoxic and stressful conditions. The third Zoophycos-bearing level (Zl 3) is characterised by an overall moderate bioturbation intensity. The distribution of trace fossils was influenced by lithology, sedimentation rate, energy level (storm events), bottom oxygenation, and nutrient supply. The lithofacies and trace fossils of the Chefar El Ahmar Formation both indicate a depositional environment fluctuating from the lower shoreface to lower offshore zone.  相似文献   

9.
The poor preservation and apparent monospecifity of Permian tetrapod footprints from eolian paleoenvironments have thus far hampered their reliable interpretation. This study clarifies how this is due to distinct and repeated ichnotaphonomic effects on trackway pattern and footprint morphology on originally inclined planes. Once these effects are excluded, the anatomy-consistent ichnotaxobases useful for ichnotaxonomy can be recognized. Several nomina dubia are identified, among these the ichnogenus Chelichnus, here considered a taphotaxon. The eolian ichnoassociations from the Lopingian of Scotland and Germany include six different ichnotaxa: cf. Capitosauroides isp. (?eutheriodont therapsid), Dicynodontipus geinitzi (cynodont therapsid), Dolomitipes isp. (dicynodont therapsid), Pachypes loxodactylus n. comb. (pareiasaurian parareptile), Procolophonichnium isp. (small parareptile) and Rhynchosauroides isp. (non-archosauriform neodiapsid). This is completely different from the interpretations of the last 20 years, which postulated that these paleoenvironments comprised monospecific associations of synapsid tracks. These ichnoassociations are instead moderately diverse, similar to low-latitude marginal marine to floodplain ichnoassociations and belong to the Lopingian Paradoxichnium footprint biochron. The Cornberg Formation of Germany, being constrained between the Illawarra reversal and the mid-Wuchiapingian Kupferschiefer at the Rotliegend/Zechstein transition, constitutes the earliest evidence of Lopingian tetrapod faunas at low paleolatitudes and the first evidence of low-paleolatitude faunal turnover related to the end-Guadalupian mass extinction from both the skeleton and the track record. This suggests a global extension of the dinocephalian extinction event, which occurred at high-mid (South Africa and Russia) and low (Western Europe) paleolatitudes of Pangea about 259–260 Ma and was probably triggered by the eruption of the Emeishan Large Igneous Province of SW China, which considerably changed global environmental conditions in both marine and continental settings.  相似文献   

10.
In the first ever systematic study of trace fossils from the Badhaura Formation, the authors described a nesting burrow, which they ascribed to a stomatopod. The purpose of this paper is two-fold: primarily, to document ichnofauna from (post-glacial marine late Palaeozoic rocks of peninsular India) the Badhaura Formation (Sterlitmakian) representing marine rocks deposited following the Late Palaeozoic glaciation and secondly to contribute to the data on post-glacial ichnofauna from constituent continents of the Gondwanaland. Trace fossils described here are from the Harbans Bed, the topmost lithounit of the Badhaura Formation. The ichnofauna includes Arenicolites tenuis, Beaconites isp., Curvolithus isp., Cylindrichnus concentricus, Didymaulichnus lyelli, Ophiomorpha isp., Palaeophycus tubularis, Planolites beverleyensis, P. montanus, Rosselia chonoides, R. socialis, Skolithos linearis, Taenidium cameronensis, Thalassinoides paradoxicus, Thalassinoides isp. and a flask-shaped brood chamber assigned to a stomatopod crustacean. This mixed assemblage is assigned to distal Skolithos ichnofacies and is suggestive of a period of relatively quiet, shallow water conditions of deposition. The ichnofauna, when viewed in context of peri-gondwanic ichnofaunas, mainly consisting of simple tracks and trails, from late Palaeozoic post-glacial deposits of other Gondwanan continents, is interesting due to dominance of domichnia. Profusion of brood chambers along with Thalassinoides in the Badhaura Formation validates the concept of pre-Mesozoic Thalassinoides being non-decapod in origin and suggestive of adaptive convergence.  相似文献   

11.
The Almería-Níjar Basin is a Neogene, intermontane depression marginal to the Mediterranean in southern Spain in the vicinity of El Argamasón, Andalucia. The Pliocene Cuevas Formation rests unconformably on the Upper Messinian rock succession in the Carboneras Fault Zone. The Cuevas Formation is a coarse-grained, bioclastic-rich, calcarenite to calcirudite shoreface deposit. Oysters, namely Saccostrea cucullata (Born), are locally common and preserve a moderate diversity of borings: Caulostrepsis taeniola Clarke; Entobia isp.; Gastrochaenolites isp. aff. G. lapidicus Kelly and Bromley; Maeandropolydora isp. cf. M. sulcans Voigt; Oichnus paraboloides Bromley; and Talpina isp. aff. T. hirsuta Voigt. All represent domiciles except for the predatory O. paraboloides trace. This suite of ichnotaxa is assigned to the Entobia ichnofacies sensu Bromley and Asgaard; they are comparable, particularly, with the Boulder Assemblage of the Pliocene of Rhodes, Greece. Physical disturbance is an important parameter in favouring this pattern of infestation, whether the bored clasts are boulders or oyster valves.  相似文献   

12.
The Early Cretaceous (latest Hauterivian) Faraoni Oceanic Anoxic Event (F-OAE) in the Río Argos section (Caravaca region, southern Spain), the candidate for the Global Boundary Stratotype Section and Point (GSSP) of the Hauterivian–Barremian transition, has been studied. Its ichnological bed-by-bed analysis allows for interpretation of oxygenation changes through the Faraoni Level interval and improvement upon previous characterization of oxygen conditions prior, during and after the F-OAE. The trace fossil assemblage belongs to the Zoophycos ichnofacies and it includes Chondrites intricatus, Chondrites targionii, Halimedides isp., Palaeophycus isp., ?Patagonichnus isp., Planolites isp., Rhizocorallium isp., Taenidium isp., Thalassinoides isp., Trichichnus linearis, Zavitokichnus fusiformis, and Zoophycos isp. Their diversity in particular beds fluctuates. Beds with four to six ichnotaxa reflect a multi-tiered macrobenthic trace maker assemblage living in good oxic and trophic conditions. In one bed below and one bed above the Faraoni Level (both marls without primary lamination), there are only two or three, mostly opportunistic ichnotaxa (Trichichnus, Chondrites, Planolites). They record dysoxic conditions. At the base of the Faraoni Level, one thicker and two thinner beds of marly mudstones (21.2 and 3.5 cm thick, respectively) are characterized by primary lamination. At the top and in the basal part of the thicker bed and in the thinner beds some trace fossils are present. These beds were deposited under anoxic conditions and later colonized by trace makers either from overlying beds deposited under oxic conditions or from the level of a greenish lamina in the lower part of the thicker bed, recording short episodes of dysoxic conditions. The thinner anoxic beds are separated by marls deposited under dysoxic conditions.  相似文献   

13.
The fossil record of mid to late Permian terrestrial vertebrates in the South African Karoo Basin is regarded as the most abundant and diverse in the world. Despite the extensive research on body fossils, to-date the vertebrate footprint sites have not been subjected to an anatomy-consistent ichnotaxonomic investigation. Here we present a comprehensive ichnotaxonomic revision of Permian-Early Triassic tracksites in the main Karoo Basin of South Africa. Furthermore, a track-trackmaker correlation for all Permian synapsid groups is provided for the first time, based on the analysis of the functional morphology of potential producers. The following ichnotaxa and their proposed trackmakers are recognized: Brontopus giganteus (dinocephalians), cf. Capitosauroides isp. (therocephalians), cf. Dicynodontipus isp. (cynodonts), Dolomitipes accordii (small dicynodonts), Dolomitipes icelsi n. comb. (large dicynodonts), Karoopes gansfonteinensis n. igen. n. isp. (gorgonopsids), Procolophonichnium nopcsai (procolophonids) and Rhynchosauroides isp. (non-archosauriform diapsids). Three different footprint assemblages (FA I–III) are proposed for footprint biostratigraphy: FA I (lower Tapinocephalus AZ), a Guadalupian assemblage dominated by dinocephalian tracks; FA II (topmost Tapinocephalus-Cistecephalus AZ), a latest Guadalupian-Wuchiapingian assemblage dominated by gorgonopsid and dicynodont tracks in association with subordinate therocephalian tracks and FA III (lower Lystrosaurus AZ), an Induan assemblage with dicynodont, cynodont, procolophonid and diapsid tracks. The lower FA II includes the earliest ichnofauna with Lopingian affinity all over the world (topmost Tapinocephalus-Pristerognathus AZ, ~260–259 Ma) and could indicate an early recovery phase after the end-Guadalupian mass extinction, because of the high abundance of large gorgonopsid tracks and absence of dinocephalian tracks. This footprint record may also predate the body fossil record, suggesting an earlier gorgonopsid radiation. FA III represents the earliest and most complete post end-Permian extinction ichnofauna, which includes an early phase of abundant small dicynodont tracks, potentially indicating a stressed post-event community. Nevertheless, this ichnofauna looks very similar to pre-extinction ichnofaunas from Europe, in agreement with the skeletal record at the Daptocephalus-Lystrosaurus AZ transition.  相似文献   

14.
The Late Oligocene oyster Hyotissa antiguensis (Brown) is locally common in the Antigua Formation of Hughes Point, eastern Antigua, Lesser Antilles; it was not commonly bored at that time. Its valves and shells are robust, and reworked into the shallow water near-shore environment in Antigua; it could potentially be incorporated into younger rocks. Its neoichnology includes clues that would facilitate identification of these oysters as reworked fossils. The suite of modern borings found in these specimens includes common Caulostrepsis taeniola Clarke, Gastrochaenolites isp. cf. G. turbinatus Kelly and Bromley and Entobia isp., and rare Oichnus simplex Bromley and Rogerella? isp. The latter three taxa are limited to oyster shell substrates. Of the common ichnotaxa, Caulostrepsis and Gastrochaenolites are particularly prominent in limestone clasts and limestone cemented to oyster shells, which would be an indicator of reworking if found in a post-Oligocene lithified deposit. Caulostrepsis and Gastrochaenolites are relatively less common in oyster shells and valves, and in many specimens are seen to terminate against the shell. Entobia is the only common boring limited to the shell substrate. The fidelity of preservation of modern borings is also superior in limestone clasts. This suite of borings is comparable with those found in the Neogene of the Antillean region.  相似文献   

15.
The limestone succession of the Middle Ordovician Volkhov Formation in the St. Petersburg region (Russia) exposes numerous horizons with firm- and hardgrounds below omission surfaces, which contain trace fossils attributable to the ichnogenus Balanoglossites Mägdefrau, 1932. Although known from the literature since 75 years, such trace fossils were previously only informally described as “Korrosionsgruben”, “Karandashi”, or ascribed to different ichnotaxa such as Trypanites, Arenicolites and Pseudopolydorites. Owing to their complexity and often high bioturbation density, the morphology of these trace fossils is difficult to capture. Ichnofabrics containing Balanoglossites triadicus were studied in detail from sawn rock faces, broken rock blocks and sectioned slabs, including those from historical buildings in St. Petersburg. Accordingly, different trace-fossil elements can be revealed in dependence on the original substrate consistency, reflecting various stages of lithification: Mineral-stained and Trypanites-perforated hardground surfaces are bioeroded with long elongated grooves which are assigned to the ichnogenus Sulcichnus. Subtle openings lead into the partly lithified limestone where they branch into complex galleries of B. triadicus. They are characterized by J-, U- and Y-shaped shafts and multiply branched tunnels, which gradually continue into the underlying firmground. Other portions of the ichnofabric only exhibit biodeformational structures or the strongly compacted and branched burrow networks Labyrintichnus, which is due to the original soft sediment consistency. Balanoglossites ichnofabrics demarcate certain omission surfaces within the Dikari Limestone and can be traced for more than 300 km, supporting the regional lithostratigraphical correlation. The trace maker of B. triadicus and related trace fossils is interpreted to be a eunicid polychate with the ability to bioerode and burrow the sediment. The studied material from the Ordovician is similar to the Balanoglossites from the type area, the Triassic of Germany, in many respects. B. triadicus is a very common trace fossil in Ordovician and other Palaeozoic rocks of Baltoscandia and North America but so far has been seldom identified as such but instead is commonly confused with Thalassinoides, from which, however, it differs in several aspects.  相似文献   

16.
The mixed carbonate-siliciclastic Weißenegg (Allo-) Formation records three depositional sequences corresponding approximately to the TB 2.3, TB 2.4 and TB 2.5 global cycles. Sea-level fluctuations were of the order of at least 30 m. Siliciclastic lowstand systems tracts comprise lignite deposits, reworked basement and tidal siltstones (above a tectonically enhanced sequence boundary) as well as coastal sand bars. Coastal sands of the transgressive systems tract contain distinct layers of well cemented nodules. They are interpreted as the first stage in hardground formation and record superimposed minor sea-level fluctuations. Coral patch reefs and rhodolith platforms developed during transgressive phases and were subsequently drowned and/or suffocated by siliciclastics during early highstand. Shallowing upwards siliciclastic parasequences, each terminated by a bank of rhodolith limestone, form the (late) highstand systems tract. The limestone beds record superimposed fourth-order transgressive pulses. Occasionally a carbonate highstand wedge developed. Lowstand carbonate shedding occurred where the top of a platform which suffered incipient drowning during highstand was near sealevel again during the following lowstand. Late highstand delta progradation is common.  相似文献   

17.
Plug-shaped trace fossils B. perata, B. hemispherica, Bergaueria isp., Calycraterion samsonowiczi, Conostichus ornatus and Conostichus isp. are found in the middle-upper Oxfordian fine-grained sandstone of the Argiles de Saïda Formation (northwestern Algeria). Except for the trace Calycraterion which is interpreted as an annelid trace, these traces correspond to distinct behaviours of sea anemones, and their occurrence within tempestite deposits is influenced by a stress factors in a storm-dominated environment. The abundance of these plug-shaped trace fossils suggests that the upper Jurassic siliciclastic shelves with fine-grained soft clastic substrate were colonised by actinarian sessile benthos.  相似文献   

18.
A common assumption in the geological analysis of modern reefs is that coral community zonation seen on the surface should also be found in cores from the reef interior. Such assumptions not only underestimate the impact of tropical storms on reef facies development, but have been difficult to test because of restrictions imposed by narrow‐diameter cores and poor recovery. That assumption is tested here using large‐diameter cores recovered from a range of common zones across three Campeche Bank reefs. It is found that cores from the reef‐front, crest, flat and rubble‐cay zones are similar in texture and coral composition, making it impossible to recognize coral assemblages that reflect the surface zonation. Taphonomic signatures imparted by variations in encrustation, bioerosion and cementation, however, produce distinct facies and delineate a clear depth zonation. Cores from the reef‐front zone (2–10 m depth) are characterized by sections of Acropora palmata cobble gravel interspersed with sections of in‐place (but truncated) A. palmata stumps. Upper surfaces of truncated colonies are intensely bioeroded by traces of Entobia isp. and Gastrochaenolites isp. and encrusted by mm‐thick crustose corallines before colony regeneration and, therefore, indicate punctuated growth resulting from a hurricane‐induced cycle of destruction and regeneration. Cores from the reef crest/flat (0–2 m depth) are also characterized by sections of hurricane‐derived A. palmata cobble‐gravels as well as in‐place A. palmata colonies. In contrast to the reef front, however, these cobble gravels are encrusted by cm‐thick crusts of intergrown coralline algae, low‐relief Homotrema and vermetids, bored by traces of Entobia isp. and Trypanites isp. and coated by a dense, peloidal, micrite cement. Cores from the inter‐ to supratidal rubble‐cay zone (+0–5 m) are only composed of A. palmata cobble gravels and, although clasts show evidence of subtidal encrustation and bioerosion, these always represent processes that occurred before deposition on the cay. Instead, these gravels are distinguished on the basis of their limited bioerosion and marine cements, which exhibit fabrics formed in the intertidal zone. These results confirm that hurricanes have a major influence on facies development in Campeche Bank reefs. Instead of reflecting the surface coral zonation, each facies records a distinctive, depth‐related set of taphonomic processes, which reflect colonization, alteration and stabilization following the production of new substrates by hurricanes.  相似文献   

19.
Paleogene sediments of the inner fold belt, Naga hills, have very well preserved ichnofossils. 16 ichnospecies have been documented among 13 ichnogenera such as Arenicolites isp., Chondrites targionii, Cylindrichnus isp., Diplocraterion parallelum, Gyrochorte isp., Ophiomorpha annulata, O. irregulaire, O. nodosa, O. rudis, Palaeophycus tubularis, Planolites beverleyensis, Scolicia palaeobullia, Skolithos linearis, Trypinites weisei, Thalassinoides horizontalis and Zoophycos isp. The ichnofossil assemblages comprise mostly domichnia and fodinichnia benthos of the Skolithos and Cruziana ichnofacies. A shallow marine nearshore to offshore marine environment with fluctuating energy condition has been envisaged.  相似文献   

20.
Five radiolitid rudist species are described from the Turonian sequence of Abu Roash area. They are recognized in three rudist biostromes that occur in two informal members of Abu Roash Formation; the Rudist- and the Actaeonella-bearing limestone–marl members. The three biostromes show autochthonous and parautochthonous fabrics and moderate to high packing potential. The first rudist biostrome at the base of the Rudist-bearing limestone–marl member (Middle Turonian) contains Durania gaensis, Praeradiolites ponsianus and Bournonia fourtaui. The second biostrome in the same member consists of Bournonia roashensis. The third biostrome that recognized in the Actaeonella-bearing limestone–marl member (Late Turonian) consists of Durania arnaudi. Rudist biostromes in the Rudist-bearing limestone–marl member were deposited on subtidal rudist shoals with moderate to high energy versus that of the Actaeonella-bearing limestone–marl member that deposited in low to moderate energy on deeper part of subtidal rudist shoals. The exposed Turonian succession at Abu Roash area could be divided into three depositional sequences bounded by three sequence boundaries (paleosols and angular unconformity).The first rudist biostrome in the Rudist-bearing limestone–marl member represents the lower part of the transgressive systems tract of the first depositional sequence. The deepening upward trend of the transgressive systems tract is due to increase of accommodation space in transgressive context during relative sea-level rise episode. On the other hand, the second rudist biostrome in the Rudist-bearing limestone–marl member and the third rudist biostrome in the Actaeonella-bearing limestone–marl member are in shallowing-upward set sequence forming the highstand systems tract of the first and third depositional sequences. This indicates that, the accommodation space was being filled more rapidly than was being created during the highstand stage.  相似文献   

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