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1.
A new CA-ID TIMS U–Pb age of 130.39 ± 0.16 Ma is presented here from the Pilmatué Member of the Agrio Formation, lower Hauterivian of the Neuquén Basin in west-central Argentina. This high precision radioisotopic new age, together with the two former ones from the upper Hauterivian Agua de la Mula Member of the Agrio Formation and modern cyclostratigraphic studies in the classical sections of the Mediterranean Province of the Tethys indicate that the Hauterivian Stage spans some 6 Ma, starting ca. 132 Ma and ending ca. 126 Ma. These radioisotopic ages are tied to ammonite biostratigraphy and calcareous nannofossil bioevents and biozones recognized in the Neuquén Basin which in turn are correlated with the Mediterranean standard zones. A new geological time scale for the Valanginian–Hauterivian stages in the Mediterranean region integrating astrochronological and radiochronological data differs with the current official geological time scale which still maintains poorly constraint absolute ages for the Berriasian-Aptian interval.  相似文献   

2.
In the last decades, the Argentinian ornithopod record has been increased with new and diverse bone remains found along all the Upper Cretaceous. Most of them are very incomplete and represent taxa of different size. As result, the studies about the palaeobiodiversity of the Ornithopoda clade in South America are complex. In this paper, new postcranial remains of an indeterminate medium-sized ornithopod from the Santonian Bajo de la Carpa Formation (Rincón de los Sauces, Neuquén province) are presented. They present diagnostic features of the Ornithopoda clade, and several characters that relate them with other Argentinian ornithopods, especially with the medium-sized members of the Elasmaria clade sensu Calvo et al. (2007) (Macrogryphosaurus and Talenkauen). The postcranial material allows to identificate at least three different ontogenetic stages: adult, subadult more immature and subadult. These bones are the first record of Ornithopoda for the Bajo de la Carpa Formation and one of the very scarce Santonian records of this clade in South America. The diversity of the Late Cretaceous South American ornithopods presents two clear distributions: the Cenomanian-Santonian was characterized by small and medium euiguanodonts and elasmarian; and the Campanian–Maastrichtian by the medium sized elasmarian and large sized hadrosaurids.  相似文献   

3.
The following note on a new method for extracting pollen from fossil pollen grains summarizes two articles in French, published in Annales de la Société Géologique de Belgique (Juvigné, 1973a; 1973b).The new method proposed has three important parts: (1) Elutriation of the sediment in water; (2) the specific gravity of the heavy liquid must be between 2.25 and 2.5; and (3) the dispersion of the sediment in the heavy liquid is made with a high-speed mixer.  相似文献   

4.
We present a new species of Steinmanella Crickmay from the Valanginian of the Neuquén Basin, west-central Argentina, and at the same time update a Valanginian–earliest Barremian bivalve zonation which is integrated into the local, Chilean and European standard ammonoid zonations. Steinmanella caicayensis sp. nov. presents a moderately inflated shell, a subtriangular outline, well-developed carinae and a straight to moderately convex anterior margin. Steinmanella is very abundant in the Tithonian–Barremian interval in the basin and thus provides an ideal opportunity to perform detailed taxonomic and stratigraphic studies; a former zonation based on these trigonioids is here revised and expanded with a thorough revision of Valanginian occurrences based on new material. Two new zones are proposed, namely the S. quintucoensis and S. caicayensis zones, encompassing Valanginian times. The proposed bivalve zonation encompassing the Valanginian–earliest Barremian time interval can be useful in correlating with other southwestern Gondwanan regions, such as Perú, Chile, South Africa, Tanzania, and India, where Steinmanella has been recorded.  相似文献   

5.
A new long-necked sauropod dinosaur, Yunmenglong ruyangensis gen. et sp. nov., is erected on the basis of an incomplete skeleton from the late Early Cretaceous Haoling Formation of the Ruyang Basin, Henan Province. The characters of the anterior cervical vertebrae, the shape of the neural canal of the dorsal vertebra and the ball-shaped distal end of the neural spine of caudal vertebrae with coarse surfaces differ from other long-necked sauropod dinosaurs. The new genus has characters in common with both Euhelopus and Erketu; it represents the first long-necked sauropod dinosaur recorded from central China to date. The rod-like, well-developed epipophyses and the pleurocoels on the cervical vertebrae indicate that it may be close to Euhelopus, an observation also confirmed by a phylogenetic analysis, which shows that Erketu, Yunmenglong and Qiaowanlong form a clade, and are more derived than Euhelopus.  相似文献   

6.
A new species of nerineoid gastropod, Eunerinea mendozana, is described from the top of the Agrio Formation, Lower Cretaceous, at Lomas Bayas, Mendoza Province, west-central Argentina. The significance of this record lies in that it extends the Early Cretaceous distribution of the genus to the Southern Hemisphere and also may point to the occurrence of subtropical conditions in the northern part of the Neuquén Basin close to the Hauterivian/Barremian boundary. Individuals of E. mendozana are found forming large monospecific assemblages immersed in carbonate sediments. The relationships of E. mendozana with other South American Early Cretaceous nerineoids are considered, together with a re-evaluation of some of those records.  相似文献   

7.
Lirainosaurus astibiae was originally described by Sanz and collaborators in 1999 on the basis of a skull fragment, isolated teeth, several vertebrae (e.g. the holotypic anterior caudal vertebra) and appendicular bones from the Late Cretaceous of Laño (northern Spain). A review of all the vertebral remains, including new material (cervical, dorsal and caudal vertebrae, dorsal ribs, haemal arch), provides additional information about the axial skeleton of Lirainosaurus. A study of the laminae and fossae shows interesting variations in these structures in the axial series, especially concerning the prezygapophyses and diapophyses: e.g. the X-shaped morphology of the centroprezygapophyseal lamina only in the posterior dorsal vertebrae, and the division of the postzygodiapophyseal fossa into two fossae in the posterior dorsal vertebrae and the proximal caudal vertebrae. Two vertebral characters are here considered to be autapomorphic for L. astibiae: the presence of a lamina in the interzygapophyseal fossa in the most proximal caudal vertebrae (a postzygodiapophyseal lamina that separates the ventral postzygapophyseal centrodiapophyseal fossa and the dorsal postzygapophyseal spinodiapophyseal fossa), and the spinopostzygapophyseal structure not posteriorly projected in the posterior caudal vertebrae. The combination of characters present in the axial remains of Lirainosaurus astibiae supports the idea that it is a derived lithostrotian close to Saltasaurinae.  相似文献   

8.
This study is the first detailed account of freshwater to restricted marine molluscs from the Upper Cretaceous sedimentary rocks in the northern sector of the Neuquén Basin. The fossils are from the Campanian–Maastrichtian Loncoche Formation in southern Mendoza, west-central Argentina, which records the initial connection of the Neuquén Basin to the Atlantic Ocean. Six species of bivalves (Diplodon bodenbenderi, Pleiodon? sp., Isognomon? mechanquilensis, Mactridae? indet., Panopea? sp., and Laternula sp.) and three of gastropods (Paleoanculosa macrochilinoides, Paleoanculosa ameghiniana, and a possible cerithioidean) are described. Specimens were collected from fine to coarse sandstones, which may be massive or with planar stratification, planar-cross stratification or trough-cross stratification, and a few from bioclastic limestones and mudstones. Although the sections are from 50 to 300 m thick, the specimens are found only in the lower 120 m. Molluscs represent autochthonous/parautochthonous assemblages composed of mostly non-broken gastropods and articulated bivalve specimens, some of which show signs of postmortem transport; however, they were not removed far from their original habitat. Review of the habitats of living genera supports the inference of dominantly freshwater palaeoenvironments in the lower and middle part of the Loncoche Formation, with restricted marine influence in the southernmost localities studied where there are a few samples that contain specimens belonging to predominantly marine groups (e.g., Laternula, Panopea).  相似文献   

9.
Four gastropod species are described from dysaerobic biofacies of the lower part of the Agua de la Mula Member of the Agrio Formation, Neuquén Basin: Metacerithium turriculatum (Forbes, 1845), Nerineopsis acutecarinatum (Behrendsen, 1891) and the new species Ampullina pichinka and Mesalia? kushea. They were active epifaunals and possibly deposit-feeders and/or browsers, and epifaunal to semi-infaunal possibly suspension- or deposit-feeder in the case of M.? kushea, thriving in soft to firm substrates. These gastropods, together with two aporrhaid species, are the dominant components of a typically small-sized, low-diversity macrobenthic fauna that occurs in dark, organic-rich shales. The facies in which this fauna is recorded are thought to represent a transition from lower to upper outer ramp marine settings, in the context of a second-order TST and early stages of a HST. They record a transition from lower dysoxic conditions in the lowest part of the Agua de la Mula Member to upper dysoxic conditions upwards. Two distinct gastropod biofacies were recognized corresponding to the two identified stages: biofacies A, dominated by Protohemichenopus neuquensis and N. acutecarinatum, more tolerant to dysoxic conditions, and biofacies B, dominated by M. turriculatum and Mesalia? kushea, less tolerant to oxygen deficiency.  相似文献   

10.
The marine Oligo-Miocene units of western Taurides, deposited under different tectonic regimes (in Bey Da?lar? platform in foreland and coeval sequences in hinterland), were studied to establish a high-resolution biostratigraphic framework. Biometric study of the full spectrum of larger foraminifera in a regional scale allowed us correlating them with the shallow benthic zonation (SBZ) system introduced by [Cahuzac, B., Poignant, A., 1997. Essai de biozonation de l’Oligo-Miocène dans les bassins européens à l’aide des grands foraminifères néritiques. Bulletin de la Société géologique de France 168, 155–169], and to determine the ages of these sites on zonal precision for the first time. In correlating these assemblages to standard shallow benthic zones, planktonic data were also used whenever possible. Taxa, classified under the genera Nummulites, Miogypsina, Miolepidocyclina, Nephrolepidina, Eulepidina, Heterostegina, Operculina and Cycloclypeus (?) and their assemblages, closely resemble to the fauna described from European basins. These groups characterize the SBZ 22B to 25 zones referring to a time interval from early Chattian to Burdigalian. However, a main gap in late Chattian (SBZ 23) and in early part of the Aquitanian (SBZ 24) is also recorded in the platform succession. In the meantime, rare Eulepidina in the Burdigalian levels suggest a clear Indo-Pacific influence. Based on the discovery of early Chattian (SBZ 22B) deposits (previously mapped under Eocene/Miocene units), the Oligo-Miocene stratigraphy of the Bey Da?lar? platform is also revised. A more precise chronology for regional Miocene transgression is presented based on the miogypsinid evolutionary scale.  相似文献   

11.
The complete osteology of the abelisaurid Viavenator exxoni, from the Bajo de la Carpa Formation (Santonian, Upper Cretaceous) is described. Viavenator is characterized by a series of autapomorphies, including: transversely compressed parietal depressions on both sides of the supraoccipital crest; ventral edges of the paraoccipital processes located above the level of the dorsal edge of the occipital condyle; basioccipital-opisthotic complex about two and a half times the width and almost twice the height of the occipital condyle, in posterior view; well-developed crest below the occipital condyle, diverging ventrally and defining the subcondylar recess; deeply excavated and sub-circular basisphenoidal recess, with its major axis transversely oriented; basipterygoid processes horizontally placed with respect to the cranial roof and located slightly dorsally to the basal tubera; mid and posterior cervical centra with slightly convex lateral and ventral surfaces; hyposphene-hypanthrum articular complex present from dorsal 2 onward; presence of an interspinous accessory articular system in middle and posterior dorsal vertebrae; presence of a pair of pneumatic foramina within the prespinal fossa in anterior caudal vertebrae; distal end of the scapular blade posteriorly curved. Particularly, Viavenator shows plesiomorphic cranial characters, i.e. flat frontals lacking domes or horns, combined with derived postcranial characters, e.g. the interspinous accessory joint system of dorsal vertebrae. This combination between plesiomorphic and derived traits suggests that Viavenator is a transitional form, which is an idea supported by its intermediate stratigraphic and phylogenetic placement between the basal and older (e.g. Skorpiovenator, Ilokelesia) and derived and younger members of the clade (e.g. Aucasaurus, Carnotaurus).  相似文献   

12.
The first Podocarpaceae wood record is described from the Mulichinco Formation (Valanginian, Lower Cretaceous), Neuquén Basin, Argentina. The specimen was directly associated with a middle caudal vertebra of a diplodocid sauropod dinosaur. A new species – Podocarpoxylon prumnopityoides – is proposed based on features that include the presence of abietinean wood type (tracheid radial pitting), plus podocarpoid (cupressoid type) and some dacrydioid (taxodioid type) cross-field pits, diffuse axial parenchyma and low rays. This combination of anatomical characters is comparable to both Prumnopitys and Podocarpus, whereas the type of pits in the cross-fields resembles some members of the extant Prumnopitys. This is the first unequivocal record of the Family Podocarpaceae in the Valanginian of South America and confirms the hypothesis that the divergence between the “Podocarpoid-Dacrydioid” and “Prumnopityoid” clades occurred earlier than the Early Cretaceous.  相似文献   

13.
Four isolated cervical vertebrae from the Kimmeridge Clay Formation (Upper Jurassic, Kimmeridgian) of Abingdon, Oxfordshire, England are identified as from a pliosaurid plesiosaurian sauropterygian on account of their shortness relative to width and height, their near platycoelous nature and the location of tall rib facets on the centrum body. They are noteworthy for their size, with a maximum width of 269 mm, maximum height of 222 mm and maximum length of 103 mm. Simple scaling and comparisons with cervical vertebrae of Mid Jurassic pliosaurs Peloneustes and Liopleurodon, and the Early Cretaceous Stenorhynchosaurus and Sachicasaurus suggest a total body length of between ~ 9.8 m and 14.4 m for the Abingdon Kimmeridge Clay pliosaur. Likely the true length was towards the higher end of this range.A genus and species cannot be confidently determined on the basis of the described material, but they likely belong to Pliosaurus sp. or a similar animal, for which a precise neck length is not known. We estimate a neck length of 0.77 m for Pliosaurus ?brachyspondylus based on the average cervical lengths provided for specimen CAMSM J.35991.  相似文献   

14.
This paper reports the occurrence of a Toarcian–Aalenian (Early–Middle Jurassic) radiolarian fauna in the Los Molles Formation, Neuquén Basin, Argentina, as well as comments on its paleobiogeographic affinities. The micropaleontologic analysis was carried out in fine-grained rocks from a turbiditic section of the Los Molles Formation. These samples were first chemically processed using only hydrogen peroxide (H2O2), and afterward treated with acetic (CH3COOH) and hydrofluoric (HF) acids. The first chemical procedure permitted the recovery of only few spongy spumellarians, while the second one enabled to recover more diversified radiolarian assemblages. In general, the studied fauna presents low diversity and abundance, with a strong dominance of spumellarians over nassellarians. The fauna is composed by the genera Paronaella, Homoeoparonaella, Praeconocaryomma, Archaeocenosphaera, Orbiculiformella, Praeparvicingula, and some unidentified spumellarians and nassellarians. According to paleobiogeographic models, the studied Toarcian–Aalenian fauna presents a mid to high latitude affinity. It is possible to infer from those data a bipolar distribution of some taxa, such as Praeparvicingula and probably Praeconocaryomma, between the Northern and Southern hemispheres since the Early Jurassic (Toarcian).  相似文献   

15.
Cervical vertebrae of elasmosaurids (Sauropterygia, Plesiosauroidea) have proven to be informative even at family level. The present paper uses a previously developed methodology based on bivariate graphic analysis that evaluates the proportion indices of cervical vertebrae in elasmosaurids, which is applied to a well-preserved postcranial specimen of this group recovered from upper Maastrichtian beds of central Chile. The plotting of such indices groups the Chilean material with other specimens from the Late Cretaceous of Argentina and Antarctica, the latter referred to the genus Aristonectes. This result allows us to identify the first relatively complete individual of the genus based on non-cranial characters. The identification of the Chilean specimen as Aristonectes sp. provides important new data on the postcranial skeleton of the genus, that has remained poorly known until now.  相似文献   

16.
Variations in the sedimentary organic matter were documented throughout the Agua de la Mula Member (late Hauterivian) of the Agrio Formation, at a combined section in the type area of the Agrio Formation; the base at Agrio del Medio and the middle to top at Bajada del Agrio. A main organic-rich interval was identified in the basal Agua de la Mula Member, dominated by marine-derived Amorphous Organic Matter (AOM), coinciding with the highest Total Organic Carbon (TOC) content, between 1.4 and 3.8 wt.%, suggesting dysoxic conditions. The rest of the Member is predominantly characterized by terrestrially-derived organic matter, mainly phytoclasts, with low TOC values, around 1% or lower, indicating predominantly oxic depositional settings. By integrating stacking pattern and shell beds analysis, four depositional sequences SQ1–SQ4 were recognized. The organic-rich, finely laminated bituminous black shales of the Spitidiscus riccardii zone constitute a mayor and rapid inundation defining a Transgressive System tract (TST), related to a third order asymmetrical mesosequence dominated by a thick High System Tract (HST). It represents the most widespread and important flooding episode within the Neuquén Basin during the late Hauterivian. Inside sequences of higher frequency (SQ1, SQ2, SQ3, SQ4) of probably fourth order were recognized and analysed including several ammonids zones (Spitidiscus riccardii, Crioceratites schlagintweiti, C. diamantensis and Paraspiticeras groeberi). With the exception of the oxygen-controlled, basinal and outer ramp settings indicated for the TST1, which is equivalent to the TST of a lower order sequence, and the lower TST2, respectively, the prevalence of well oxygenated, inner to middle ramp depositional environments, is suggested for the rest of the sedimentary succession and emphasized in HST of SQ2, SQ3 and SQ4. Thus, a shallowing-upward trend with improved oxygenation is recorded through the Agua de la Mula Member, reflected by decreasing TOC, AOM content and preservation state of the organic matter. Based on the whole rock fluorescence analysis of the two selected organic-rich intervals from the Spitidiscus riccardii and the lowermost Crioceratites diamantensis zones, the presence of hydrocarbons suggests a very early in situ generation.  相似文献   

17.
A new abelisaurid theropod from the Cenomanian of Candeleros Formation (Neuquén Province, NW Patagonia, Argentina) is described. It includes an isolated frontal, partial pelvis, two fused sacral centra, femoral shaft, rib fragments, partial metatarsal II and shaft of metatarsal III. A paleohistological analysis shows that MMCh-PV 69 was somatically mature at time of death. The prevalence of relatively ordered intrinsic fibers in the primary bone indicates a relatively lower growth rate of MMCh-PV 69 compared with other abelisauroids, such as Aucasaurus garridoi. Phylogenetic analysis found MMCh-PV 69 as a basal abelisaurid, sister group of the node formed by Majungasaurinae and Brachyrostra. We estimated the body mass of MMCh-PV 69 in 240 kg, which makes it one of the smallest abelisaurids recorded. This new form adds to the extensive theropod fauna of the Candeleros Formation, probably the most diverse association of meat-eating dinosaurs recorded in a Cretaceous Formation from South America.  相似文献   

18.
Saltasaurine titanosaurs are characterized by their relatively small size compared to other sauropods, extreme postcranial pneumaticity, and dermal armour covering the body. This group has been reported in the Upper Cretaceous of the Lecho, Allen, and Anacleto formations of Argentina. We describe here a new saltasaurine specimen (MACN-Pv RN 233) from the Campanian of the Angostura Colorada Formation (Río Negro Province) that is represented by eight caudal vertebrae and six osteoderms. This specimen is described in detail and its phylogenetic relationships with the other three known saltasaurines, as well as its implications to the knowledge of caudal vertebra and osteoderm anatomy, are discussed. Our results place MACN-Pv RN 233 more closely related to Saltasaurus loricatus and Rocasaurus muniozi than to Neuquensaurus australis. MACN-Pv RN 233 possesses a combination of features that differ from other saltasaurines, but because of the fragmentary nature of the specimen we decided for the sake of taxonomic stability to not erect a new taxon. This specimen shows the first unambiguous evidence of chevron pneumatisation for a sauropodomorph, implying a broader osteological invasion of the diverticula from the abdominal air sac than previously thought for this group of dinosaurs. MACN-Pv RN 233 preserves two osteoderm morphotypes, one similar to those reported for Neuquensaurus australis and Saltasaurus loricatus. This new specimen expands the distribution of the group to a new geological unit and increases the dinosaur diversity known for the Angostura Colorada Formation.  相似文献   

19.
Two-dimensional analytical models of pumping induced drawdown and stream depletion account for the streambed properties and stream geometry. Using data from three pumping tests performed under various hydrologic conditions, the following hypotheses were tested: (1) a partially penetrating stream at a certain distance from the pumping well can be represented by the streambed leakage term in the analytical models; (2) in the streambed leakage term, the streambed conductance coefficient λ=(WK′)/m′ accounts for the average stream width W, the streambed thickness m′, and hydraulic conductivity K′. The first hypothesis was tested by comparing results from tests under flow and no-flow stream conditions. The second hypothesis was tested by comparing results from two tests under low and high streamflow conditions. Similar estimates of the hydraulic conductivity and storativity for all tests indicate the validity of the streambed leakage term. The drawdown data of the test under low- and high-flow conditions (varying W) do not follow the predictions of the analytical models, which results in inconsistent λ estimates. Thus, for different hydrologic conditions, λ cannot be scaled solely by the stream width W. One possible explanation for this result is streambed dynamics caused by the changes in the stream stage.
Résumé Les rabattements dans les nappes et cours d’eau générés par des modèles analytiques de pompages en deux dimensions dépendent des propriétés du lit du cours d’eau considéré et de sa géométrie. En utilisant les données de trois essais de pompages effectués en conditions hydrologiques contrastées, les hypothèses suivantes ont été testées: (1) un ruisseau à pénétration partielle situé à une certaine distance du puits de pompage peut être exprimé par le terme de drainance dans les modèles analytiques; (2) dans le terme de drainance du lit du cours d’eau, le coefficient de conductivité du lit λ=(WK′)/m′ représente la largeur moyenne du cours d’eau W, l’épaisseur du lit m′, et la perméabilité K′. La première hypothèse a été testée en comparant les résultats des tests en conditions d’assèchement et d’écoulement. Pour éprouver la seconde hypothèse, les résultats de deux tests en conditions d’étiage et de hautes eaux ont été comparés. Sur l’ensemble des tests, la concordance entre les perméabilités calculées a permis de valider le terme de drainance du cours d’eau. Les rabattements observés en conditions d’étiage et de hautes eaux (W variable) ne suivent pas les prédictions des modèles analytiques, fournissant au final des estimations discordantes de λ. Aussi, dans le cas de conditions hydrologiques différentes, la seule largeur du cours d’eau ne suffit pas à estimer λ. Une explication possible de ce résultat réside dans la dynamique du lit causée par les variations de niveau du cours d’eau.
Resumen Modelos analíticos en dos dimensiones de descenso inducido por bombeo y de agotamiento de la corriente del río representan las propiedades del lecho del río y la geometría del mismo. Usando datos de tres ensayos de bombeo llevados a cabo bajo condiciones hidrológicas variadas, se probaron las siguientes hipótesis: (1) puede representarse un río parcialmente penetrante a una cierta distancia del pozo de bombeo por el término de goteo a través del lecho del río en los modelos analíticos; (2) en el término de goteo a través del lecho del río, el coeficiente de conductancia del lecho del río λ=(WK′)/m′ representa el ancho promedio del río W, la potencia del lecho del río m′, y la conductividad hidráulica K′. La primera hipótesis fue comprobada comparando los resultados de pruebas bajo condiciones de flujo y no flujo de corriente. La segunda hipótesis fue probada comparando los resultados de dos pruebas bajo condiciones de corriente altas y bajas. Estimaciones similares de la conductividad hidráulica y el almacenamiento para todas las pruebas apuntan hacia la validez del término de goteo a través del lecho del río. Los datos de descensos de la prueba bajo condiciones de flujo altos y bajos (variando W) no siguen las predicciones de los modelos analíticos, lo que resulta en estimaciones inconsistentes de λ. Así, para condiciones hidrológicas diferentes, λ no puede ser escalada solamente con el ancho de la corriente W. Una posible explicación para este resultado es la dinámica del lecho del río causada por los cambios en el nivel del río.
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20.
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