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1.
A sample of fallen blocks of fluvial sandstone from the Lower Cretaceous Jiaguan Formation of Sichuan Province yielded an assemblage of dinosaur and pterosaur tracks preserved as natural impressions and casts. Collectively the assemblage reveals 132 tracks representing at least 30 trackways of tridactyl and didactyl theropods, sauropods, ornithopods and pterosaurs. Ichnotaxonomically, the trackways of small tridactyl theropods (pes lengths 7–18 cm) are indeterminable, whereas the trackway of a small didactyl dromaeosaur (pes length up to 7.5 cm) is tentatively assigned to cf. Velociraptorichnus. The sauropod trackways are assigned to cf. Brontopodus based on the medium to nearly wide-gauge pattern. Other characteristics are the U-shaped manus and strong heteropody. One sauropod trackway shows a peculiar pattern with a lack of left manus imprints, and an unusual position and rotation of right manus imprints. Different scenarios and explanations for this phenomenon are discussed. Ornithopod trackways are the most abundant in the sample and characterized by pes imprints of a small biped that are assigned here to the new ichnospecies Caririchnium liucixini. It is characterized by an unusual broad shape and weak mesaxony. Bivariate analysis of different Caririchnium ichnospecies reveals increasing mesaxony toward the larger forms, a trend that is the reverse of typical theropod ichnotaxa, where large imprints have weak mesaxony. Three isolated, small pterosaur tracks (two manus, one pes) are visible on a single surface. They show strong similarities to the widespread ichnogenus Pteraichnus. This is the ninth report of tetrapod tracks from the Jiaguan Formation in recent years and represents one of the most diverse assemblages recorded to date. It is also rare evidence of typical didactyl dromaeosaur tracks and the co-occurrence of sauropod and ornithopod tracks in a fluvial depositional environment representing arid climate conditions. 相似文献
2.
记述了云南省禄丰县腰站乡下侏罗统禄丰组2个恐龙足迹,命名了棋盘张北足迹（Changpeipus pareschequier ichnosp. nov.）一新种。这是在禄丰组中首次发现恐龙足迹化石,重新观察并修订了张北足迹一属的属征。滦平张北足迹（Changpeipus luanpingeris）应为石炭张北足迹（Changpeipus carbonicus）的亚成年个体,为同物异名。此前被认为是石炭张北足迹（Changpeipus carbonicus）前足迹的IVPP V2472 2a,应为其未成年体的后足迹。张北足迹与卡岩塔足迹（Kayentapus）类似。综合种种特征,棋盘张北足迹（新种）的造迹恐龙很可能为禄丰组的腔骨龙类（Coelophysoidea） 相似文献
3.
An unusual theropod track assemblage from the Cretaceous of the Zhucheng area, Shandong Province, China 总被引：3，自引：0，他引：3
More than 125 footprints of theropods from the Cretaceous Longwangzhuang Formation have been mapped in a preliminary study at a site in the Zhucheng region of China. The tracks represent at least three morphotypes. The largest morphotype is a large theropod (footprint length ∼30 cm) represented by a single trackway and an isolated natural cast. At least 10 trackways assigned to the new ichnospecies Corpulentapus lilasia represent a medium-sized biped (footprint length ∼13 cm) with very short, wide, robust, ‘tulip-shaped’ tracks and long steps (∼5 × footprint length), and a short central digit (III) indicating weak mesaxony. Corpulentapus trackways are narrow and theropod-like even though track morphology is convergent with the footprints of some ornithopods. The third morphotype, made by a medium-sized grallatorid track maker (ichnogenus Paragrallator), is about the same size (∼13 cm) as the robust morphotype, but far more elongate and gracile, with an elongate central digit (III) indicating strong mesaxony. This ichnotaxon requires detailed comparison with Grallator sensu stricto. The contrast in morphology between the two common morphotypes is striking and demonstrates that two distinct medium-sized taxa of presumed theropod affinity frequented the same habitat in significant numbers. 相似文献
4.
Here we report a large dinosaur tracksite from an extensive fluvial sandstone surface in the Lower Cretaceous Jiaguan Formation of Sichuan Province, China. The site contains over 250 individual tracks comprising at least 18 recognizable trackways, including the longest theropod trackway (cf. Eubrontes) known from China. This exceptional theropod trackway consists of 81 successive footprints covering a distance of 69 m. The tracks are well-preserved and are expressed both as true tracks on the main “upper” surface and as transmitted undertracks on a locally exposed “lower” bed. Also recorded are six other theropod trackways, including small Grallator-like ichnites, eight sauropod trackways (cf. Brontopodus), and three small ornithopod (cf. Ornithopodichnus) trackways with a parallel orientation, which may indicate gregarious behavior. Several trackways of a larger theropod trackmaker show pes imprints with elongated traces of the metatarsals, suggesting extramorphological (substrate-controlled) variation and/or plantigrade posture, which is here interpreted as indicating a change in gait assumed in response to deep and soft sediment. The assemblage indicates a diverse dinosaur fauna in the Lower Cretaceous Sichuan Basin with variously sized theropods, sauropods, and ornithopods. The late occurrence of footprints of the Grallator-Eubrontes plexus in Lower Cretaceous strata is further evidence of the extended stratigraphic range of this morphotype and the distinct palaeobiogeographic distribution of these trackmakers in East Asia. 相似文献
5.
The Marwar Supergroup of the Bikaner-Nagaur Basin is composed of sediments deposited from the late Neoproterozoic (Ediacaran) to Upper Cambrian. The Nagaur Sandstone Formation of the Nagaur Group (uppermost division of the Marwar Supergroup) preserves trace fossils significant for establishing Early Cambrian biostratigraphic zones and depositional facies. Fifteen ichnospecies (and eight ichnogenera) identified in the Nagaur Sandstone Formation include “Treptichnus” pedum, Cruziana cf. tenella, Cruziana isp., Diplichnites ispp. A, B, and C, Gyrophyllites isp., Lockeia isp., Merostomichnites isp., Monomorphichnus gregarius isp. nov., Monomorphichnus isp., Planolites isp., Psammichnites isp., Rusophycus bikanerus isp. nov., Rusophycus cf. carbonarius, Rusophycus isp. and radial trace fossils.These trace fossils belong to ethological categories pascichnia, repichnia, cubichnia, and fodinichnia and represent arthropod and worm-like burrowing biota. The assemblage and a regional comparison with contemporaneous trace fossils in the eastern Gondwanan realm suggest that the sequence in the study area belongs to the Cruziana tenella Ichnozone and to Stage 2 (upper part of Terreneuvian), however the Middle Cambrian is not excluded. The trace fossil assemblage belongs to the archetypal Cruziana ichnofacies. Cross bedded sandstone, mud cracks and rainprints in the ichniferous strata of the Nagaur Sandstone Formation indicate deposition in an intertidal sand flat with channels that was exposed episodically. 相似文献
6.
The John Martin Reservoir tracksites from the Dakota Group of Bent County, in southeastern Colorado form part of the mid Cretaceous Dinosaur Freeway characterized by abundant ornithopod footprints (Caririchnium). Over 350 tracks (331 Caririchnium, 1 Magnoavipes, 22 crocodiles, and a few pterosaurs) were discovered at 10 new tracksites. All tracks were found as natural casts, including Caririchnium trackways that were still in-situ, parallel and regularly spaced, suggesting gregarious behavior. Most crocodile tracks are swimming tracks that consist of three or four scratch marks also with parallel orientations in many cases. A few pterosaur tracks consist of pes footprints and scratch marks suggesting swimming or floating activity in shallow water. This is the first pterosaur evidence from the Dakota Group. The Caririchnium size structure from John Martin Reservoir is consistent with the size structure tendency of the whole Dinosaur Freeway, which shows larger track size in the north. This tendency could be interpreted as evidence for more than one ornithopod species spread out across the Dinosaur Freeway. The alternative, that there was one ornithopod species that migrated north and south seasonally is less likely. The John Martin paleoecosystem is interpreted from the track evidence as a well-vegetated coastal plain environment with many ornithopods and a few theropods on land, crocodiles in the water, and pterosaurs in the skies and on the water. 相似文献
7.
Imad Bouchemla Mohamed Bendella Madani Benyoucef Olev Vinn Bruno Ferré 《Proceedings of the Geologists' Association. Geologists' Association》2021,132(2):207-226
The Upper Emsian to Frasnian Ia-Ib strata of the Marhouma area (or “km 30” outcrop), exposed in the Ougarta Range (SW Algeria) belong to the Chefar El Ahmar Formation. On the basis of distinct lithological and palaeontological features, this formation is subdivided into three members (Lower Marly Limestones Member, Middle Marly Limestones Member, and Upper Marly Limestones Member). The studied beds show low to moderate diversity of trace fossil assemblage which contains thirteen ichnotaxa: Chondrites intricatus, Chondrites isp., Chondrites cf. targionii, Circulichnis cf. montanus, Cochlichnus isp., Neonereites biserialis, Neonereites multiserialis, Nereites isp., Palaeophycus isp., Planolites isp., Thalassinoides isp., Zoophycos aff. cauda-galli, and Zoophycos isp. A. The two latter ichnotaxa are the most common trace fossils in the assemblage and occur at three different levels showing different bioturbation intensities. The first Zoophycos-bearing level (Zl 1) is characterised by an overall high bioturbation intensity reflecting a very high oxygenation rate and nutrient supply, allowing the development of large and dense Zoophycos specimens. The second Zoophycos-bearing level (Zl 2) has a considerable reduction of bioturbation intensity as compared to the previous level, with an abundance of Chondrites, which is probably due to radical palaeoecological changes that suggests dysoxic and stressful conditions. The third Zoophycos-bearing level (Zl 3) is characterised by an overall moderate bioturbation intensity. The distribution of trace fossils was influenced by lithology, sedimentation rate, energy level (storm events), bottom oxygenation, and nutrient supply. The lithofacies and trace fossils of the Chefar El Ahmar Formation both indicate a depositional environment fluctuating from the lower shoreface to lower offshore zone. 相似文献
8.
The poor preservation and apparent monospecifity of Permian tetrapod footprints from eolian paleoenvironments have thus far hampered their reliable interpretation. This study clarifies how this is due to distinct and repeated ichnotaphonomic effects on trackway pattern and footprint morphology on originally inclined planes. Once these effects are excluded, the anatomy-consistent ichnotaxobases useful for ichnotaxonomy can be recognized. Several nomina dubia are identified, among these the ichnogenus Chelichnus, here considered a taphotaxon. The eolian ichnoassociations from the Lopingian of Scotland and Germany include six different ichnotaxa: cf. Capitosauroides isp. (?eutheriodont therapsid), Dicynodontipus geinitzi (cynodont therapsid), Dolomitipes isp. (dicynodont therapsid), Pachypes loxodactylus n. comb. (pareiasaurian parareptile), Procolophonichnium isp. (small parareptile) and Rhynchosauroides isp. (non-archosauriform neodiapsid). This is completely different from the interpretations of the last 20 years, which postulated that these paleoenvironments comprised monospecific associations of synapsid tracks. These ichnoassociations are instead moderately diverse, similar to low-latitude marginal marine to floodplain ichnoassociations and belong to the Lopingian Paradoxichnium footprint biochron. The Cornberg Formation of Germany, being constrained between the Illawarra reversal and the mid-Wuchiapingian Kupferschiefer at the Rotliegend/Zechstein transition, constitutes the earliest evidence of Lopingian tetrapod faunas at low paleolatitudes and the first evidence of low-paleolatitude faunal turnover related to the end-Guadalupian mass extinction from both the skeleton and the track record. This suggests a global extension of the dinocephalian extinction event, which occurred at high-mid (South Africa and Russia) and low (Western Europe) paleolatitudes of Pangea about 259–260 Ma and was probably triggered by the eruption of the Emeishan Large Igneous Province of SW China, which considerably changed global environmental conditions in both marine and continental settings. 相似文献
9.
Basal iguanodontians (“iguanodontids”) were a successful group of ornithopod dinosaurs that attained a near global distribution by the late Early Cretaceous. Despite their body fossils being known in abundance from the uppermost Jurassic and Lower Cretaceous of the western United States, their remains have never been found in western Canada. With its extensive, terrestrial sedimentary record for the whole of the Early Cretaceous, the expectation is that these dinosaurs would have been present in western Canada. This paper reports the finding of a fossil footprint from the Gladstone Formation in southwestern Alberta that is interpreted to have been made by an iguanodontid. This identification is based on the late Barremian age of the hosting rock, gross footprint details that match the skeletal foot structure of Iguanodon bernissartensis, and similarities to Early Cretaceous dinosaur footprints found elsewhere in the world that are attributed to iguanodontids. This finding fills a noticeable gap in the iguanodontian fossil record, and is consistent with the clade's attainment of global distribution during the Early Cretaceous. 相似文献
10.
In the locality of Rambla de los Gavilanes (Betic Chain Murcia, Spain), six new outcrops with dinosaur footprints are reported from an uppermost Cretaceous succession of coastal marine deposits. Four of these sites are found in the upper part of the lower Maastrichtian and the other two in the upper Maastrichtian, these latter located only a few meters below the Cretaceous-Paleocene boundary (K/Pg). Traces appear as impressions or as natural casts, on tabular limestone beds consisting of micritic carbonate facies, these deposited in coastal marine areas that include a variety of environments (coastal lakes, tidal flats, salt marshes, and shallow coastal lakes). Two main types of traces have been recognized, respectively attributed to sauropod footprints and large ornithopod footprints with remarkable pointed hoof marks. The Maastrichtian sites with large ornithopod footprints in Europe are reported from Spain, Poland and Romania. Sauropod (titanosaurian), theropod (Irenesauripus and other undetermined theropods), and ornithopod footprints have been found at sites located up to 20 m below the K/Pg boundary. The Spanish dinosaur ichnites known closest to the K/Pg boundary are in the Pyrenean upper Maastrichtian, and are tridactyl, theropod and ornithopod (Hadrosauropodus and Amblydactylus) footprints. In this work, the assignation of the Rambla de los Gavilanes ornithopod footprints has been made following the recommended synthesis in several recent papers which reduce the large ornithopod ichnogenus to four. 相似文献
11.
Newly discovered tracks of a large quadrupedal ornithopod and a theropod with trail trace, and a bird track from the Cretaceous Saniri Formation of Yeongdong area are described. One ornithopod trackway is very unusual in having large (length width ∼15 and >15 cm respectively) clover-leaf-like manus tracks situated anteromedial to the pes tracks with atypical negative, inward rotation of 45°. Ornithopod pes tracks are quadripartite with three separated, elongate-oval, nearly parallel sided, wide digit impressions and a separate suboval heel impression (mean length and width about 41 and 36 cm respectively: l/w ratio 1.13). Manus track morphotypes are a clover-leaf-shaped configuration of three digit impressions, representing digits II–IV, in triangular configuration and registered just in front of (anteromedial to) pes track digits II and III. The pes morphotype is typical of Caririchnium, but the manus morphotype is quite distinct from previously described ichnotaxa, thus justifying a new ichnotaxon: Caririchnium yeondongensis ichnosp. nov., probably representing a facultatively-quadrupedal Iguanodon-like trackmaker.Theropod tracks are composed of three tapered pes digit impressions with interdigital angles between digits II and IV 45°. Length and width about 22.8 cm and 15.5 cm, respectively. Pace, stride, and pace angles are about 51 cm, 101 cm, and 170°, respectively. Theropod tracks are characteristically associated with a nearly continuous tail trace, which is up to 360 cm in length, 4.5 to 6 cm in width, and broad “U” shaped in cross section. Expulsion rims and dragging striae occur intermittently. In addition, dinosaur skin impressions, poorly preserved large sauropod tracks, a bird track, invertebrate and plant fossils are found from the lake margin deposits also containing rain drop impressions and desiccation cracks. Dinosaur tracks of the Yeongdong area represent the oldest (Valanginian-Hauterivian) dinosaur tracks of Korea. 相似文献
12.
Three-dimensional tracks provide unique insights into the locomotor mechanics of their track makers. An isolated, large hadrosauriform print attributable to Caririchnium lotus from the “mid”-Cretaceous Lotus track site (Jiaguan Formation) in China permits reconstruction of the footfall, weight-bearing, and kick-off phases of the step cycle. Large-scale modifications of the pes during the step cycle indicate C. lotus trackmakers were capable of locomotory modifications in response to substrate consistency beyond the “expected” shift between bipedal and quadrupedal postures. An unusual curvature to the trace of one of the outer digits indicates substantial transverse mobility. The remaining digits demonstrate lesser degrees of transverse movement accompanied by extension of the digits during footfall. The absence of overprinted scale-scratch marks and toe drags are consistent with a vertical kick-off of the pes and concomitant flexion of the digits. This track suggests that pedal mobility in C. lotus track makers was greater than previously suspected and has implications for reconstructions of hadrosauriform locomotion. 相似文献
13.
《Proceedings of the Geologists' Association. Geologists' Association》2021,132(5):529-536
Abundant Trypanites isp. associated with Gastrochaenolites isp. and Petroxestes pera are evidenced in the first found hardground of the late Albian marine transgressive series, overlying the Continental Intercalaire in the Ouled Nail Range, eastern Saharan Atlas. This hardground is encrusted by oyster shells belonging to Ostrea falco Coquand. The studied hardground developed in warm environment and displays a lateral extension of at least 6 km. It is interpreted as an erosional transgressive surface. The ichnogenus Petroxestes is reported for the first time from Africa. This ichnoassemblage is attributed to the Trypanites ichnofacies. 相似文献
14.
记述了辽宁省四合屯下白垩统义县组三个半恐龙足迹,归入似鹬龙足迹（亦译为跷脚龙足迹）一未定种（Grallator isp.）。这是义县组恐龙足迹化石的首次描述。该行迹至少由3个造迹者所造。从足迹推断恐龙体长1.51m,属于义县组兽脚类恐龙较为普遍的体长范围。重建了尾羽鸟(Caudipteryx)和中华龙鸟(Sinosauropteryx)的足部,前者的足迹轮廓与似鹬龙足迹未定种的吻合度超过后者。根据化石记录,似鹬龙足迹类型可能广泛存在于义县组的各种中小型兽脚类（驰龙类与伤齿龙类除外）中。 相似文献
15.
An ichnofossil assemblage is preserved in sediments of the Middle Bhuban Formation,Surma Group,of Miocene age from the Durtlang and Chanmari sections,Aizawl district,Mizoram,northeastern India.The studied sections comprise a thick,bioturbated and thinly bedded sandstone–silty shale facies.The preserved ichnofossils show high density and low diversity with the assemblage represented by feeding and resting traces,i.e.,Psilonichnus upsilon Frey et al.,1984,Ophiomorpha isp.,Teichichnus spiralis Mikulá.1990,Skolithos isp.,Palaeophycus isp.,and other horizontal burrows.Sedimentary structures associated with these traces are mainly cross bedding,flaser bedding and slump structures.Considering the distribution of the Psilonichnus ichnogenus across an integrated ichnological–sedimentological framework,the stratigraphic interval of the Middle Bhuban Formation,in which they occur,are interpreted to have been deposited under a shallow,marginal-marine channel complex dominated by tidal channels developed in quiet,brackish-water portions of a delta plain. 相似文献
16.
A total of more than 40 tridactyl and didactyl tracks were preserved as natural casts on four fallen blocks of sandstone representing the Lower Cretaceous Jiaguan Formation of Gulin County in southeastern Sichuan Province, China. While several trackways can be distinctly followed, others are isolated imprints only. All have been flattened by overburden pressures. Tridactyl tracks are present with three size-classes being <10 cm, 10–20 cm and >20 cm in length. Morphologically they are similar to the ichnogenus Eubrontes, considering the relatively weak mesaxony. Eight of the tracks on one of the blocks are clearly didactyl and are here interpreted as representing large and medium sized dromaeosaurids. The largest track is about ∼30 cm long and comparable in size to the type of Dromaeopodus (∼28 cm), from the Lower Cretaceous of Shandong Province, which was the largest dromaeosaurid track previously reported. This report adds new data to the growing number of dromaeosaurid tracksites reports from China, and from the Jiaguan Formation, suggesting that this theropod group had a preference for fluvial paleoenvironments. 相似文献
17.
记叙了中国陕西省商洛市邵涧村下白垩统东河群2个大型兽脚类恐龙足迹,并将其归于Megalosauripus isp.。
东河群的时代与热河生物群同期,邵涧Megalosauripus isp. 可能可与热河生物群的大型兽脚类恐龙之足部形态做简
单对比。此外,邵涧Megalosauripus isp.造迹者的体长约为7.4m,这意味着早白垩世的陕西中南部亦存在着大型的
兽脚类恐龙。邵涧Megalosauripus isp.的发现,不仅为中国早白垩世恐龙足迹增加了新的成员,而且为将来在该地
区下白垩统中发现大型兽脚类恐龙的骨骼化石提供了确凿的依据。 相似文献
18.
Joshua Hedge Anthony P. Shillito Neil S. Davies Richard J. Butler Ivan J. Sansom 《Proceedings of the Geologists' Association. Geologists' Association》2019,130(1):103-111
The late Carboniferous–early Permian was a period of major environmental change, with the rainforests that covered the equatorial zone during the Carboniferous disappearing due to increasing aridification. This environmental transition had significant impacts on the terrestrial biota, including a major extinction event among plant and vertebrate groups. A rich and unique ichnofauna from the Alveley Member (Moscovian: Westphalian D) of the Salop Formation at Alveley in Shropshire (England) has yielded important insights into late Carboniferous terrestrial communities. However, research to date has focused entirely on the vertebrate footprints. Abundant invertebrate ichnofossils also occur at Alveley, typically on the rippled upper surfaces of beds assigned to a floodplain facies that preserve the vertebrate tracks in hyporelief on their base. We provide the first detailed examination of the invertebrate ichnofauna from Alveley, identifying six ichnospecies within five ichnogenera (Diplichnites, Gordia, Paleohelcura, Palmichnium, Protichnites), including a new species of the common arthropod ichnogenus Diplichnites. This moderately diverse invertebrate ichnofauna is dominated by arthropod repichnia. There is no evidence of infaunal bioturbation and the single example of Gordia indicates limited sediment grazing activity. The Alveley ichnofauna is typical of Euramerican continental ichnoassemblages from the very latest Carboniferous, and indicates a moderate diversity of arthropods (crustaceans, arachnids and chelicerates) living alongside the marginal freshwater to terrestrial assemblage of temnospondyl amphibians, and basal synapsid, diadectomorph and captorhinomorph amniotes. 相似文献
19.
The new non-marine bivalve species Nippononaia (Martinsonella) tamurai sp. nov. is described from the Upper Formation of the Mifune Group in Kumamoto Prefecture, Southwest Japan. The specimens originally were identified as Plicatounio (Plicatounio) B sp. by Tamura (1990). The subgenus Nippononaia (Martinsonella) previously was only reported from China, and this is the first record from Japan. The Upper Formation of the Mifune Group is of Late Cenomanian to Early Turonian age, as indicated by ammonites. “Nippononaia” (?) obsoleta Hase, 1960 from the Shiohama Formation of Yamaguchi, Japan, and Plicatounio (s.l.) A sp. of Tamura (1990) from strata northeast of Geoncheonri, South Korea, are re-assigned to Nippononaia (Martinsonella). These occurrences may be of significance for the inter-regional correlation of non-marine Cretaceous strata. 相似文献
20.
豫西济源地区早三叠世和尚沟组湖相遗迹化石及遗迹组构 总被引：2，自引：0，他引：2
豫西济源地区早三叠世和尚沟组滨浅湖沉积中动物遗迹化石共鉴定出9个遗迹属10个遗迹种。包括Arenicolites isp., Beaconites coronus, Cylindricum isp., Palaeophycus heberti, Palaeophycus tubularis, Planolites isp., Psilonichnus isp., Scoyenia gracilis, Skolithos linearis和Taenidium barretti等。根据遗迹化石分布特征及沉积环境分析,可识别出三种遗迹组构1）Scoyenia遗迹组构,反映了干旱气候条件下的滨湖沉积环境;2）Planolites—Taenidium遗迹组构,属于湖泊水体逐渐变浅的浅湖沉积环境;3）Psilonichnus遗迹组构,代表了水动力由弱到强的湖泊三角洲沉积环境。 相似文献