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1.
During the latest Cretaceous, distinct dinosaur faunas were found in Laurasia and Gondwana. Tyrannosaurids, hadrosaurids, and ceratopsians dominated in North America and Asia, while abelisaurids and titanosaurians dominated in South America, India, and Madagascar. Little is known about dinosaur faunas from the latest Cretaceous of Africa, however. Here, a new abelisaurid theropod, Chenanisaurus barbaricus, is described from the upper Maastrichtian phosphates of the Ouled Abdoun Basin in Morocco, North Africa on the basis of a partial dentary and isolated teeth. Chenanisaurus is both one of the largest abelisaurids, and one of the youngest known African dinosaurs. Along with previously reported titanosaurian remains, Chenanisaurus documents the persistence of a classic Gondwanan abelisaurid-titanosaurian fauna in mainland Africa until just prior to the end-Cretaceous mass extinction. The animal is unusual both in terms of its large size and the unusually short and robust jaw. Although it resembles South American carnotaurines in having a deep, bowed mandible, phylogenetic analysis suggests that Chenanisaurus may represent a lineage of abelisaurids that is distinct from those previously described from the latest Cretaceous of South America, Indo-Madagascar, and Europe, consistent with the hypothesis that the fragmentation of Gondwana led to the evolution of endemic dinosaur faunas during the Late Cretaceous. 相似文献
2.
Xing Lid Ariana Paulina-Carabajal Philip J. Currie Xu Xing Zhang Jianping Wang Tao Michael E. Burns Dong Zhiming 《《地质学报》英文版》2014,88(6):1653-1664
The neuroanatomy of the mid-sized theropod Sinosaurus triassicus from the Lower Jurassic Lufeng Formation, Lufeng Basin in Yunnan Province, China was studied using X-ray computed tomography. The braincase is characterized by a large supraoccipital knob that is capped by a posterior projection of the parietal and two external foramina for the caudal middle cerebral vein, which is completely enclosed by the supraoccipital. The basicranium has well defined, short basipterygoid processes that project ventral to the basal tubera. The basisphenoid is expanded, projects posteroventrally, and is pierced by four pneumatic recesses. The endocranial morphology resembles that observed in other basal theropods—in particular some allosauroids—and has a strongly marked pontine flexure and a large dorsal expansion. The inner ear morphology is also similar to that observed in other basal theropods, with slender semicircular canals. The anterior semicircular canal is 20% larger than the posterior semicircular canal, and the angle formed between them is less than 90° when seen in dorsal view. 相似文献
3.
A well preserved skeleton of a new abelisaurid is reported here. The holotype of Viavenator exxoni was found in the outcrops of the Bajo de la Carpa Formation (Santonian, Upper Cretaceous), northwestern Patagonia, Argentina. This new taxon belongs to the South American clade of abelisaurids, the brachyrostrans. The current phylogenetic analysis places it as the basalmost member of a new clade of derived brachyrostrans, named Furileusauria, characterized by distinctive cranial, axial and appendicular anatomical features. The Santonian age of Viavenator allows filling the stratigraphic gap exhibited between the basal brachyrostrans of Cenomanian–Turonian age, and the derived forms from the Campanian-Maastrichtian. The evolution of abelisaurids during the Late Cretaceous, faunal replacements, and the adaptive radiation that occurred during that period of time in South America are discussed. 相似文献
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R.V. Dingle 《Cretaceous Research》1982,3(4):367-389
One hundred and thirty-six species, representing 67 genera have been recorded from the late Jurassic-Maastrichtian marine sediments of South Africa. The faunas show a major dichotomy across a regionally-developed late Cenomanian-early Coniacian hiatus with the Portlandian-Cenomanian Cytheruridae/Progonocytheridae/Schizocytheridae dominated faunas being replaced in the Coniacian by Trachyleberididae/Brachycytheridae/Schizocytheridae dominated faunas. Comparison with other Gondwanide localities shows that the two South African basins from which ostracods have been described (Outeniqua and Natal/Zululand) formed part of a Callovian-Cenomanian South Gondwana ostracod province that stretched from the Neuquen Basin of Argentina to Madagascar/Tanzania/Kutch and west Australia. The most characteristic and cosmopolitan forms within this province belong to the Majungaella/Amicytheridea/Progonocythere group, along with Arculicythere in the Aptian-Cenomanian.In Tanzania, (the only locality of the old South Gondwana province where the succession is complete) these assemblages are replaced in the Turonian by the influx of Brachycythere, and Cythereis and various other trachyleberids. Changes of a similar nature are seen whenever marine sedimentation resumed after the local “mid” Cretaceous hiatus (South Africa, India, Argentina). Argentina differs in not having Brachycythere, whose rapid appearance in the West Indian Ocean basin soon after its earliest record in Brazil, is attributed to the destruction of the barrier at the eastern end of the Walvis Ridge/Rio Grande Rise in late Cenomanian or early Turonian times. Despite this common element with Brazil and West Africa, the South African Coniacian to Maastrichtian faunas are closer to those of Tanzania and Australia than they are to either Argentina or Brazil/West Africa. In Zululand they show evidence of a steady increase in water depth, leading to the establishment of progressively more diverse cytheracean populations, with a particularly large increase across the Santonian/Campanian boundary. 相似文献
6.
《Cretaceous Research》2012,33(6):781-785
The terrestrial lizard fossil record of Gondwana is very scarce. Few lizards, from mostly fragmentary fossils, have been identified in Madagascar, Tanzania, Morocco, South Africa, India and South America. Among the South American specimens there are basal Squamata forms (Olindalacerta and Tijubina) and a possible iguanid (Pristiguana) form Brazil. In Argentina gondwanian terrestrial lizards are represented by a putative iguanid and a small, poorly preserved dentary. This last specimen, recovered from the Anacleto Formation (Neuquén Group, Río Colorado Subgroup) in vicinity of the locality of Cinco Saltos (Río Negro Province), is here described formally. Comparisons with extant lizards indicate more affinity of the fossil with the Scincomorpha (non Scincophidia) than with any other group of lizards. This finding suggests that lizards were probably better represented and more diverse in the Mesozoic of South America than previously thought, although the fossil record is, at the moment, much poorer than in Laurasia. 相似文献
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New theropod remains with abelisaurid affinities from the Upper Cretaceous (Anacleto Formation, lower Campanian), NW Patagonia, Argentina, are here described. The specimen (MPCN-PV 69) consists of a partial premaxilla, fragmentary vertebrae, proximal portion of both humeri, distal portion of the pubis, and an incomplete pedal ungual. Characters linking with Abelisauridae are a premaxilla with a subquadrangular body, externally ornamented, and paradental plates with a striated surface; and humerus with bulbous proximal head, conical internal tuberosity, and reduced greater tubercle. The humerus is similar to those of Carnotaurus and Aucasaurus, due to the presence of a bulbous head and a discontinuity between the head and the internal tuberosity, but also differs from both taxa in the more distal location of the greater tubercle with respect to the internal tuberosity. Aucasaurus also comes from Anacleto Formation, but differences in the humeri suggest that MPCN-PV 69 is a different taxon. The phylogenetic analysis performed supports the affiliation to Abelisauridae, but fails to determinate a more precise relationship with others abelisaurids. However, a majority rule consensus of the analysis shows a position within Brachyrostra. Despite being fragmentary, MPCN-PV 69 probably represents a new abelisaurid from the Anacleto Formation, thus increasing the knowledge and diversity of Late Cretaceous South American abelisaurids. 相似文献
9.
《Gondwana Research》2013,23(3-4):1141-1150
Herein we report on a new anuran genus and species, Uberabatrachus carvalhoi, from exposures of the Marília Formation (Maastrichtian) at the well-known fossil site “Ponto 1 do Price” near Peirópolis, Uberaba, Brazil. The holotype consists of an articulated partial skeleton characterized by a broad, well-ossified skull lacking dermal ornamentation, eight procoelous presacral vertebrae, and posteriorly deflected, cylindrical sacral diapophyses; these features, together with the possible presence of a discrete palatine, are consistent with neobatrachian affinities. It clearly differs from the neobatrachian Baurubatrachus, also from the Marília Formation, in having a skull lacking exostosis and with a different temporal architecture, among other features. It also differs from the still unnamed neobatrachian recovered from the Adamatina Formation in the presence of paired frontoparietals with narrow supraorbital flanges in the anterior orbital region and maxilla-squamosal contact. The new frog is not referable to any of the neobatrachian taxa known from the Aptian–Albian Crato Formation owing to different cranial features (e.g., position of the jaw articulation, relative width of the braincase, and morphology of the frontoparietal). With respect to the extant neobatrachians in the context of recent phylogenetic hypotheses, the cylindrical sacral diapophyses of Uberabatrachus contrast with the dilated diapophyses occurring in australobatrachian hyloids and microhyloid ranoids, whereas the procoelous presacral column is unlike the diplasiocoelous condition of most other ranoids. This evidence together with the combination of features of Uberabatrachus suggests a placement among nobleobatrachian hyloids, although this combination is not shared with any extant or extinct species for which skeletal information is known. From the foregoing we conclude that Uberabatrachus represents a new nobleobatrachian taxon, a record that reinforces the hypothesis of a diversified neobatrachian fauna in South America by the Cretaceous and is consistent with the previous proposal that neobatrachian early diversification is related to the breakup of Gondwana. 相似文献
10.
A new abelisaurid theropod from the Cenomanian of Candeleros Formation (Neuquén Province, NW Patagonia, Argentina) is described. It includes an isolated frontal, partial pelvis, two fused sacral centra, femoral shaft, rib fragments, partial metatarsal II and shaft of metatarsal III. A paleohistological analysis shows that MMCh-PV 69 was somatically mature at time of death. The prevalence of relatively ordered intrinsic fibers in the primary bone indicates a relatively lower growth rate of MMCh-PV 69 compared with other abelisauroids, such as Aucasaurus garridoi. Phylogenetic analysis found MMCh-PV 69 as a basal abelisaurid, sister group of the node formed by Majungasaurinae and Brachyrostra. We estimated the body mass of MMCh-PV 69 in 240 kg, which makes it one of the smallest abelisaurids recorded. This new form adds to the extensive theropod fauna of the Candeleros Formation, probably the most diverse association of meat-eating dinosaurs recorded in a Cretaceous Formation from South America. 相似文献
11.
Fossil plants from Upper Jurassic sediments belonging to the Gymnosperms s.l. have been studied in the Manamana massif from the southwestern part of Madagascar. The levels are Upper Oxfordian in age on the basis of the palaeontological content, especially with ammonites. The majority of fossil plants are devoided of organic matter but in some cases collodion peels have been made to present epidermal characters. The cuticle of Brachyphyllum madagascariense nov. sp. has been observed with scanning electron microscope (SEM). This new flora is constituted by Pteridospermales and Coniferales. Fragmentary fronds of Pteridospermales have been collected of Pachypteris sp. A and Pachypteris sp. B. Some epidermal characters are described. Coniferales are represented by leafy stems. The diagnosis of Brachyphyllum madagascariense nov. sp. is given with morphological and epidermal characters. Others conifers have been determined: Brachyphyllum sp. A, Brachyphyllum sp. B, Elatocladus sp., Cupressinocladus sp., Cyparissidium sp.. Araucarites sp. remains are probably isolated fertile scales of a female cone. Complete cones of Conites sp. are also present.Comparisons of the systematic composition are made with other Gondwana floras of the same age from Antarctica, South America, Madagascar, India, South Africa, New Zealand and Australia. This flora shows xerophytic adaptations. With the Filicales described by Appert (1973a, b) from the same localities, it is now the most diverse flora from the Upper Jurassic of Madagascar, despite the lack of Cycadales, Bennettitales and Angiosperms. 相似文献
12.
The terrestrial lizard fossil record of Gondwana is very scarce. Few lizards, from mostly fragmentary fossils, have been identified in Madagascar, Tanzania, Morocco, South Africa, India and South America. Among the South American specimens there are basal Squamata forms (Olindalacerta and Tijubina) and a possible iguanid (Pristiguana) form Brazil. In Argentina gondwanian terrestrial lizards are represented by a putative iguanid and a small, poorly preserved dentary. This last specimen, recovered from the Anacleto Formation (Neuquén Group, Río Colorado Subgroup) in vicinity of the locality of Cinco Saltos (Río Negro Province), is here described formally. Comparisons with extant lizards indicate more affinity of the fossil with the Scincomorpha (non Scincophidia) than with any other group of lizards. This finding suggests that lizards were probably better represented and more diverse in the Mesozoic of South America than previously thought, although the fossil record is, at the moment, much poorer than in Laurasia. 相似文献
13.
Carlos Roberto A. Candeiro Agustín G. Martinelli Leonardo S. Avilla Thomas H. Rich 《Cretaceous Research》2006,27(6):923-946
An updated, annotated list of all tetrapods from the Adamantina, Uberaba and Marília formations (Bauru Group), which constitute some of the best studied Upper Cretaceous units in Brazil, is presented. Tetrapod diversity in the Bauru Group is remarkable, including an admixture of typically austral Gondwanan taxa (e.g., abelisaurids, notosuchians) and boreal Gondwanan forms (e.g., carcharodontosaurids). Of note is the absence of Laurasian taxa in the upper portion of the Bauru Group. With the exception of some turtles, an anuran, mesoeucrocodylians and one titanosaur, most taxa from the Bauru Group are based on fragmentary and isolated bones, and as such many specimens can be identified only to a higher taxonomic level. Fishes, turtles, anurans, mesoeucrocodylians, dinosaurs, birds and mammals from the Adamantina and Marília formations resemble the latest Late Cretaceous vertebrate faunas from southern South America, except for the absence of ornithischian dinosaurs. 相似文献
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Alfredo E. Zurita Alfredo A. Carlini David Gillette Rodolfo Sánchez 《Journal of South American Earth Sciences》2011,31(2-3):178-185
Knowledge of the main aspects of the Great American Biotic Interchange (GABI) concerning the glyptodontine Glyptodontidae (Xenarthra) is very scarce. A bidirectional dispersal process was recently proposed for this clade, with the presence of the North American genus Glyptotherium Osborn recognized in latest Pleistocene sediments of northern South America (Venezuela and Brazil). However, the earliest stages of this paleobiogeographical process remain poorly understood, mainly because of the limited fossil record on this clade in late Pliocene sediments. The goals of this contribution are: a) to present and describe the first record of a glyptodontine glyptodontid from the late Pliocene of northern South America, tentatively assigned to a new species of Boreostemma Carlini et al. (Boreostemma? sp. nov); and b) to analyze its paleobiogeographical implications with respect to the GABI. This new material was recovered from the San Gregorio Formation (late Pliocene, prior the GABI) in northern Venezuela, where it is represented by several osteoderms of the dorsal carapace. A comparison among the three known late Pliocene glyptodontine glyptodontids of a) southern South America (Paraglyptodon), b) northern South America (Boreostemma), and c) southern North America ("Glyptotherium"), reveals a series of shared characters between (b) and (c), not present in (a). The most important of these shared characters in (b) and (c) are: all the osteoderms present a great development of the central figure, which is always larger than the peripherals; the sulcus that delimits the central and peripheral figures is narrower and shallower; and all the osteoderms present are relatively thin. This evidence suggests that the lineage of Glyptodontinae which participated in the GABI and subsequently diversified in North America originated in northern South America. Moreover, the evident morphological differences between these glyptodontines with respect to the southern South American forms show a significant separation of both lineages since at least latest Miocene-early Pliocene. 相似文献
16.
Sections and fusulinids of the Bolorian (presumably) and Kubergandian (lower part) stages in the Sanandaj-Sirjan tectonic zone are described. Two fusulinid assemblages are distinguished in a most complete section near Sirjan. The lower one is represented by Skinnerella, Paraleeina, and relatively primitive Misellina forms, whereas Armenina, Kubergandella, and Yangchienia species appear in the upper assemblage and suggest its early Kubergandian age. Accordingly, the lower assemblage is attributed to the Bolorian Stage, although it is lacking fusulinids typical of this stage except for the primitive Misellina morphotypes. Fusulinids from the Tange-Darchaleh section near Qomsheh (Shahreza) are typical of the Kubergandian Stage. The described three new species of the genus Skinnerella are close to morphotypes known from younger (Murgabian) deposits and represent their ancestral forms most likely. 相似文献
17.
In this study, we erect Sauroniops pachytholus gen. et sp. nov., a large-bodied theropod dinosaur from the Cenomanian (Upper Cretaceous) of Morocco, on the basis of an almost complete frontal showing a unique combination of features including a naso–frontal suture extended along 40% of the frontal length, a thick dome in the anterolateral corner of the dorsal surface, a trapezoidal prefrontal facet that is restricted to the anterodorsal margin of the lateral surface of the frontal with no participation in the orbital roof and separated from the lacrimal facet by a narrow vertical lamina, a hypertrophied ‘D-shaped’ lacrimal facet that is four times the anterior depth of the postorbital facet, and a raised posteromedial margin of the dorsal surface describing a saddle with the anterolateral dome and confluent with a series of anteromedial rugosities. Phylogenetic analysis found robust support for placing Sauroniops among the basal carcharodontosaurids and related to Eocarcharia, showed that some of the unusual features of the new theropod were convergently acquired by abelisaurids, and revealed a mosaic pattern in the evolution of the carcharodontosaurid skull table. The frontals of Sauroniops and Carcharodontosaurus, both from the ‘Kem Kem compound assemblage’ of Morocco, show comparable size but differ in the extent of the naso–frontal articulation, the shape and disposition of the prefrontal and lacrimal articulations, the development of dorsal ornamentation and the morphology of the supratemporal fossa. Among carcharodontosaurids, the skull table developed unique configurations among each lineage and appears diagnostic at the species-level. The dome-like frontal in Sauroniops may indicate head-butting behaviour in this taxon or evolved for visual display. 相似文献
18.
Choristoderes are a group of extinct diapsid reptiles that once occupied the freshwater systems in the Northern Hemisphere from the Middle Jurassic through the Miocene. The Early Cretaceous monjurosuchid Philydrosaurus from western Liaoning, China, represents a transitional morphotype between a broad-snouted (crocodile-like) and a narrow-snouted (gavial-like) skull during the evolution of Choristodera. New specimens of the taxon from the Early Cretaceous Jiufotang Formation include an incomplete braincase and a nearly complete juvenile skeleton; the study of these specimens provides new information on the braincase and ontogenetic change of the skeleton of this transitional monjurosuchid. In the braincase floor, the foramen internus canalis caroticus externus opens in a groove lateral to the constricted lateral aspect of the parasphenoid, and ventral exposure of this foramen differs from Champsosaurus, in which parasphenoid/pterygoid fusion completely encloses the canal. On the occiput, the vagus nerve foramen penetrates the basioccipital, while the two foramina for the hypoglossal nerve open at the exoccipital/basioccipital suture. Comparative study of the new juvenile specimen with adult Philydrosaurus reveals previously unknown developmental changes of the cranial and postcranial skeleton of this monjurosuchid. Ontogenetically, Philydrosaurus underwent a dramatic change of skull proportions, including elongation of the antorbital and postorbital regions, and elongation of the jaws, with a great increase of the number of marginal teeth. The lower temporal fenestra is entirely closed in early ontogeny, as in large, fully-grown adults. 相似文献
19.
Proterozoic terrains in South India and Madagascar provide important clues in understanding the Gondwanaland tectonics, especially the assembly of this mega-continent during the Pan-African period. The Archaean terrains in both Madagascar and India are characterized by N-S trending greenstone belts occurring within gneissose granitic rocks in the northern part. Extensive development of K-rich granitic rocks of ca. 2.5 Ga is also characteristic in both areas. Such a broad age zonation of younger Dharwar (ca 2.6–3.0 Ga) in the north and the older Sargur (ca 3.0–3.4 Ga) in the south as in South India remains to be identified in future studies from Madagascar. The occurrence of greenschist facies rocks in the northeastern part and higher grade rocks in most of other parts in the north-central terrain of Madagascar is comparable with the general tendency of increasing metamorphic grade from northwestern to southern areas ranging from greenschist to granulite facies in South India. The Proterozoic crystalline rocks in both continents show pronounced lithological similarity with the wide occurrence of graphite-bearing khondalite in association with charnockitic rocks. While the Archaean-Proterozoic boundary is well defined in southern India by the Palghat-Cauvery or the KKPT shear zones as recently identified, this boundary is ill-defined in Madagascar due to extensive Pan-African overprinting, as well as the development of the Proterozoic cover sequence, the Itremo Group. There is also a possible general correlation between the Mesoproterozoic cover sequences in Madagascar and India, such as between the Itremo Group of west-central Madagascar and the Kaladgi and Cuddapah sequences of South India. The Pan-African granulite facies metamorphism of ca. 0.5 Ga extensively developed in both India and Madagascar is generally comparable in intensity and extent. P-T conditions and P-T-t paths also appear comparable, with the general range of ca. 700–1000°C and 6–9 kb, and near-isothermal decompressional paths. A-type granite plutons and alkaline rocks including anorthosites and mafic plutonic rocks of ca. 500–800 Ma develop in both terrains, provide strong basis for the correlation of both terrains, and define a Pan-African igneous province within East Gondwanaland. Major shear zones in both continents are expected to play a critical role in the correlation, albeit are still poorly constrained. Detailed elucidation of the tectonic history of the shear zones, and the timing of various events along the shear zones would provide important constraints on the correlation of the two continental fragments. 相似文献