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1.
We describe two new species of extinct biting midges in the fossil genus Protoculicoides: Protoculicoides hispanicus Szadziewski and Arillo, sp. nov. and Protoculicodes sanjusti Szadziewski and Arillo, sp. nov., from Albian (101–113 Ma) amber of San Just, Spain. Atriculicoides Remm, 1976 is recognized as a new junior synonym of Protoculicoides Boesel, 1937 (NEW SYNONYM). Devalquia Choufani et al., 2013 is regarded here as a new junior synonym of the extant genus Culicoides Latreille, 1809 (NEW SYNONYM). Metahelea roggeroi Choufani et al., 2013 from Upper Cretaceous French amber, is excluded from the tribe Heteromyiini and placed in the extant genus Stilobezzia Kieffer, 1911, tribe Ceratopogonini: Stilobezzia roggeroi (Choufani et al., 2013), comb. nov. The extinct tribe Atriculicoidini Szadziewski, 1996 is regarded as a new subfamily, Atriculicoidinae, stat. nov., that includes the fossil genus Protoculicoides, with 13 species reported from Albian, Turonian, Cenomanian, Coniacian, Santonian and Campanian ambers (78–113 Ma). Protoculicoides skalskii Szadziewski & Arillo, 1998 from Lower Cretaceous amber of Álava, Spain, Protoculicoides succineus Szadziewski, 1966 from Lower Cretaceous Lebanese amber and Protoculicoides burmiticus Szadziewski & Poinar, 2005 from Upper Cretaceous Burmese amber are transferred to the extinct genus Archiculicoides Szadziewski, 1996, comb. nov. Protoculicoides krzeminskii Choufani et al., 2014 from Lower Cretaceous Lebanese amber is transferred to the extinct genus Archiaustroconops Szadziewski, 1996, comb. nov. The Cretaceous subfamily Atriculicoidinae forms an unresolved trichotomy with the extant subfamilies Forcipomyiinae and Dasyheleinae, both of which date to the Eocene. The fossil record indicates that wings with macrotrichia in biting midges evolved during the mid-Cretaceous greenhouse climate. We suggest that this was an evolutionary adaptation to new atmospheric conditions with higher levels of CO2 in order for the antennal Johnston's organs of males to receive the vibrational sex signals produced by females during flight. 相似文献
2.
Novelaria, a new genus of rhagionid of late Albian age with three new species, is the first record of this family from Charentes amber (southwestern France). The new genus is probably closely related to the recent genus Chrysopilus. However its relationship with the other fossils in amber is discussed. A key for separation of the new species is provided and the diversity of the family during the Cretaceous is also briefly discussed. 相似文献
3.
Diverse new material of mantises found in the Cretaceous amber-bearing deposits from Lebanon (Barremian), Spain (Albian), and Myanmar (Albian–Cenomanian) are described and figured. The Lebanese and Spanish forms are nymphs; while the one from Myanmar is an adult specimen. The Lebanese nymph corresponds to a new specimen of Burmantis lebanensis Grimaldi, 2003 while the adult Burmese (Myanmar) specimen belongs to the new species Burmantis zherikhini. The Spanish specimen represents a new genus and species and is established as Aragonimantis aenigma, but is considered family incertae sedis. The Spanish specimen is the first record of Mesozoic mantises from western-European amber deposits. A revised phylogenetic hypothesis for Cretaceous mantises is proposed. 相似文献
4.
A new biting midge Archiculicoides andersoni sp. nov. from Upper Cretaceous Burmese amber is described and illustrated. An unknown male of Leptoconops myanmaricus Szadziewski, 2004 is described and an undetermined female of the genus Archiaustroconops and Austroconops in the collection of National Museums Scotland is reported. A key for the determination of 10 named species in 6 genera of biting midges reported from Burmese amber is also provided. 相似文献
5.
Andy Gale 《Proceedings of the Geologists' Association. Geologists' Association》2019,130(6):659-672
Diverse thoracican cirripedes from the Hauterivian of the Hannover district of northern Germany are described, including seven species, belonging to five genera. Of these, a new genus belonging to the Scalpellidae, Jaegerscalpellum, includes one Hauterivian species, J. elegans sp. nov., an Aptian species, J. comptum (Withers, 1910) and an Albian species, J. politum (Darwin, 1851) are also referred to it. A new Cretiscalpellum, C. mutterlosei sp. nov. is described from the Hauterivian, and C. matrioni sp. nov. is described from the Middle Albian of France. The oldest record of the Unilatera Gale, 2018, Pedupycnolepas pulcher sp. nov. is described from the Hauterivian; this displays typical shell structure of the group, retained by living Verrucidae. Finally, four species of Zeugmatolepadidae, subfamily Martillepadinae, are recorded from the Hauterivian, including Martillepas hausmanni (Koch and Dunker, 1836), M. decoratus sp. nov., M. auriculum sp. nov. and Etcheslaepas borealis (Collins, 1990). The Hauterivian fauna from Hannover shows affinities both with Late Jurassic and later Cretaceous (Aptian-Cenomanian) forms, and includes the earliest scalpellids, unilateran (Pedupycnolepas) and Cretiscalpellum species known. It constrains the age of the Cretaceous cirripede evolutionary radiation to the earliest Cretaceous. 相似文献
6.
A diverse fauna of wasps of the extinct parasitoid family Stigmaphronidae (Ceraphronoidea) are recorded in Early Cretaceous (Lower Albian) amber from Spain. Seven new species in five genera are described and figured based on 51 specimens, representing more material than in all the world’s other amber deposits combined. New species include: Elasmophron mari sp. nov., Libanophron sugaar sp. nov., Hippocoon basajauni sp. nov., Burmaphron jentilak sp. nov., B. sorginak sp. nov., B. iratxoak sp. nov., and Tagsmiphron olentzero sp. nov. The significance of the fauna is discussed and compared with that of other Cretaceous amber deposits, in particular the tremendous richness of the Spanish fauna is contrasted with the complete absence of stigmaphronids in the slightly younger and nearby French amber. Whether this stark difference represents particularly favorable conditions for these parasitoids, or their hosts, in the Cretaceous Spanish archipelago, or whether it is owing to taphonomic factors is discussed. 相似文献
7.
Situated at the easternmost area of the Late Mesozoic Jehol Biota, China, abundant fossils, including invertebrates, vertebrates and plants, were collected from the Early Cretaceous Yingzuilazi Formation of the Baishan Basin, southeastern Jilin. The faunal remains from this formation distinctly belong to the Early Cretaceous Lycoptera–Ephemeropsis–Eosestheria assemblage of the Jehol Biota. Based on the gross leaf morphology and epidermal structures, two new species and one indeterminate species of genus Solenites(Czekanowskiales), S. baishanensis sp. nov., S. gracilis sp. nov. and S. sp. are described in this paper. This is the first fossil plant of Czekanowskiales reported from this new fossil locality of the easternmost area of the Late Mesozoic Jehol Biota, China. The new discovery contributes to improve our knowledge for understanding of leaf morphology, epidermal characters and diversity of this genus during the Early Cretaceous time, and extends geologic and geographic distribution of Solenites in northern China and Eurasia. Along with consideration of associated plant and faunal assemblages and sedimentology, we suggest that the regional climate was warm and moderately humid with seasonal temperature and precipitation variations in warm–temperate zone during the Early Cretaceous. 相似文献
8.
Marcelle K. BouDagher-Fadel 《Cretaceous Research》1996,17(6):767-771
Tunisia, Ukraine, Russia, the North Sea area and elsewhere have yielded knowledge that the Early Cretaceous planktonic foraminifera are predominantly members of the Praehedbergellidae. The taxonomy and phylogeny of this family have been revised and refined. New generic and specific taxa have been proposed, new phylogenetic lineages have been described, and many type specimens have been imaged by scanning electron microscopy for the first time.Gorbachikella(Hauterivian–Early Aptian) gave rise toPraehedbergella(Late Hauterivian–Late Aptian), evolvingBlefuscuiana,Lilliputianella, and the planospiralBlowiellain the Barremian. The last gave rise toGlobigerinelloidesin the Aptian, from which the macroperforateAlanlordella(and its descendantPlanomalina) evolved in the Albian.Blowiellawas ancestral to taxa with radially elongated chambers (Claviblowiella, Leupoldina, Schackoina). 相似文献
9.
The Eutheiini includes over 90 extant species classified in seven genera and distributed predominantly in the Northern Hemisphere. So far only one extinct genus and species unambiguously placed in this tribe has been known, Archeutheia, from Albian of Spain. We report the discovery of Eutheia, a member of the largest extant genus of Eutheiini, in Santonian of northern Siberia. Extant species of Eutheia are primarily defined on the basis of male genital characters, and the specimen discovered in Taimyr amber is a female; consequently it is described as Eutheia sp. The new finding remarkably extends the known range of Eutheiini during Cretaceous over the area of about 6 thousand kilometers, from the Iberian Plate to northern Siberia. A long morphological stasis in Eutheia suggests that this genus was associated with stable mesic microhabitats of the upper soil layers or rotten wood for at least 83 my. 相似文献
10.
Mirovia 《Cretaceous Research》2002,23(6):761
The Lower–Middle Albian coaly clay bed of the Escucha Formation, which is exposed at Rubielos de Mora (eastern Iberian Ranges, Spain), contains a diverse fossil plant assemblage. Among the taxa present in this layer, Mirovia gothanii Gomez sp. nov. differs from other species of the genus by its greater leaf length, margins typically overhanging the depressed stomatal groove, a single short, blunt, papilla borne by each subsidiary cell, non-stomatal cells inside the groove and margins, and a higher number of resin ducts in the mesophyll. Morphological study of the well-preserved cuticles demonstrates that the species also occurs in Lower Cretaceous coals of Santa Maria de Meià (Pyrenees, Spain) where Gothan (1954) described it as Sciadopitytes sp. Both localities constitute the southernmost extent of the genus in Laurasia when the family was likely to have reached its climax in terms of abundance and diversity. 相似文献
11.
<正>One new genus and five new species of the family Evaniidae are described from the Early Cretaceous (Albian)Spanish amber of Penacerrada-Ⅰ(Province of Burgos),San Just and Arroyo de la Pascueta(both in the Province of Teruel):Cretevania alonsoi sp.nov.,C.montoyai sp.nov.,C.alcalai sp.nov.,C.rubusensis sp.nov.,and Iberoevania roblesi gen.and sp.nov.Taxonomic changes include Cretevania pristina(Zhang and Zhang,2000) comb.nov.,C.exquisita(Zhang,Rasmtsyn,Wang and Zhang,2007)comb.nov.,C.vesca(Zhang,Rasnitsyn,Wang and Zhang,2007)comb.nov.,and C.cyrtocerca(Deans,2004)comb.nov.,as a result of the reinterpretation of the genera Procretevania and Eovernevania.The new well preserved specimens of the genus Cretevania,together with the characters shown by the type specimens of the synonymized genera,give new information about their anatomical characters of taxonomical importance,and the genus Cretevania Rasnitsyn,1975 is re-diagnosed.The holotypes of the Russian species in amber have been revised.A cladistic analysis of fossil and extant groups of the superfamily Evanioidea is included.Cretevania had a wide palaeogeographic distribution,with the highest diversity known from Spain.The 13 known Cretevania species show a high interspecific variation mainly in wing characteristics,and a wide range of body and wing size. 相似文献
12.
A new family of thrips, Moundthripidae, is described on the basis of a new genus and species, Moundthrips beatificus, from Early Cretaceous Lebanese amber. This taxon has plesiomorphic prognathous mouthparts, a unique type of wing venation and the main apomorphies of the Thysanoptera of the legs and mouthpart structures, suggesting that they were acquired very early during the evolution of this order. 相似文献
13.
14.
Three new species within the stigmaphronid genus TagsmiphronEngel and Grimaldi, 2009, and one new species within the megaspilid genus ConostigmusDahlbom, 1858 are described from Upper Cretaceous (Campanian) amber originating at the Grassy Lake locality in Alberta, Canada. New taxa include Tagsmiphron spiculum sp. nov., Tagsmiphron leucki sp. nov., Tagsmiphron exitorum sp. nov., and Conostigmus cavannus sp. nov. The new Conostigmus species is a rare discovery. It is the third megaspilid species to be found in Cretaceous amber, with the two specimens described herein effectively doubling the number of known Mesozoic exemplars for the family. We provide the first comprehensive report of known Ceraphronoidea within Canadian amber, and contrast this against other Cretaceous amber assemblages, discussing the potential palaeobiogeographic and palaeoenvironmental implications of the Canadian amber assemblage. 相似文献
15.
New Mesozoic Mesopsychidae(Mecoptera)from Northeastern China 总被引：2，自引：2，他引：0
<正>The Mesozoic family Mesopsychidae Tillyard,1917 presently consists of seven described genera and 13 species from the mid-Triassic to the Early Cretaceous of Australia,South Africa and Eurasia.In the present paper one new genus and three new species of fossil mesopsychids are described that add significant distributional and stratigraphic extensions to the family.This finding documents the first formal record of fossil Mesopsychidae in China.Both Lichnomesopsyche gloriae gen.et sp.nov.and L.daohugouensis gen.et sp.nov. were found from the Middle Jurassic Jiulongshan Formation of Inner Mongolia,whereas Vitimopsyche kozlovi sp. nov.,of mid Early Cretaceous age,was collected from the Jehol Biota of northern Hebei Province.This new,well-preserved material from China reveals complete and previously unknown body features,including head, antennae,mouthparts,legs and abdomen.The delicate and long proboscides of these new taxa indicate that they were feeding on externally exposed,nutrient-rich fluids of gymnospermous ovulate fructifications,and incapable of piercing surface epidermis-attributable principally to the absence of stylets.These proboscides originated, perhaps multiply,among basal Mecoptera and are functionally and structurally convergent with equivalent mouthparts borne by fossil and extant Diptera,Lepidoptera,Neuroptera and Coleoptera. 相似文献
16.
A new genus, Cretaproscolia, and three new species of scoliid wasps, Archaeoscolia hispanica, Cretoscolia montsecana and Cretaproscolia josai, are described from the Lower Cretaceous of Spain and Brazil. The species representing the new genus is attributed to the plesiomorphic extant subfamily Proscoliinae, while the two other new species are assigned to two previously described genera in the extinct, archaic subfamily Archaeoscoliinae. The Brazilian species is the first Mesozoic scoliidid to have been described from the New World. 相似文献
17.
The Family Afrograptidae is a ‘conchostracan’ group with multiple radial costae reaching to the umbo on their carapaces. It comprises four described genera: Afrograpta, Camerunograpta, Congestheriella and Graptoestheriella with a total of thirteen described species which are occasionally reported from the Jurassic and the Cretaceous in Africa, Europe and South America (i.e. Afrograpta from the Upper Cretaceous of Cameroon; Camerunograpta from the Jurassic to Cretaceous of Cameroon; Congestheriella from the Jurassic to Upper Cretaceous of the Congo Basin, Brazil, Bulgaria, Venezuela and Argentina; and Graptoestheriella from the Upper Jurassic to Lower Cretaceous of Brazil). A new genus and a new species, Surreyestheria ockleyensis gen. et sp. nov., belonging to the Family Afrograptidae from the Lower Cretaceous (lower Barremian) Upper Weald Clay Formation of Ockley Village, Surrey County, southern England is described in this paper. The new genus mainly differs from the other four genera by the special reticulate ornamentation on its carapace. It indicates that the Family Afrograptidae was more diverse and more widely distributed in the late Mesozoic than previously supposed. Afrograptidae is a special branch of Estheriellina the latter originating in the late Palaeozoic and the former in the early Mesozoic. Afrograptids, as a whole had been widespread across Pangea in the Early Jurassic. 相似文献
18.
The diversity of serphitid wasps (Proctotrupomorpha: Serphitoidea) in Early Cretaceous (Albian) amber from Spain is described. Four new species have been found representing the genera Serphites Brues 1937, Aposerphites Kozlov and Rasnitsyn 1979, and Microserphites Kozlov and Rasnitsyn 1979. From the Peñacerrada I (Moraza) outcrop two species are described as Aposerphites angustus Ortega-Blanco, Delclòs, Peñalver and Engel, new species and Serphites lamiak, new species. A single species was found at the San Just (Teruel) outcrop and is described as S. silban, new species. Another single specimen was found in El Soplao (Cantabria) outcrop, described as Microserphites soplaensis, new species. This last specimen is especially interesting in sharing typical serphitid and mymarommatoid characters, giving additional support to the apparent close relationship of both groups. 相似文献
19.
The third Cretaceous Aeshnoptera in amber is described from Myanmar. It represents a new family Burmaeshnidae fam. nov., genus and species Burmaeshna azari gen. et sp. nov. Its exact affinities remain uncertain but it is probably the sister group of the Late Cretaceous family Enigmaeshnidae. This discovery supports the hypothesis of an intense period of appearance of many aeshnopteran subclades during the late Early Cretaceous and the Late Cretaceous. 相似文献
20.
The Cretaceous hetero diadematid echinoids Heterodiadema Cotteau, 1862 and Trochodiadema de Loriol, 1900 are revised and provided with amended diagnoses. Of six species previously referred to Heterodiadema, only three are now retained in this genus, namely Heterodiadema libycum (Desor in Agassiz & Desor, 1846), Heterodiadema ouremense de Loriol, 1884 and Heterodiadema buhaysensis Smith, 1995. Similarly, seven species were previously assigned to Trochodiadema, but here only Trochodiadema abramense de Loriol, 1900 is left in this genus. Based on material previously referred to Trochodiadema, two new genera and one new species are established: Loriolidiadema gen. nov. from the Lower Cretaceous of the Middle East, and Arabicodiadema gen. nov. and Arabicodiadema alii sp. nov from the Albian and Cenomanian of the Arabian Peninsula. Trochodiadema isidis Fourtau, 1921 is transferred to Aegyptiaris Abdelhamid & Abdelghany, 2015. New morphological data on primary and secondary spines and perignathic girdle are presented; these improve our understanding of the evolution and relationships of the family Heterodiadematidae Smith & Wright, 1993. 相似文献