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1.
The mandible of a polar bear (Ursus maritimus Phipps) found in about 1920 at Kjul Å, North Jutland, and described by Nordmann & Degerbol in 1930. has been l4C dated to 11.100 ± 160 B.P. It is so far the only find of polar bear in Denmark. Comparison with recent 14C datings of Swedish and Norwegian polar bears shows that the Danish specimen was a member of a southern Scandinavian Late Weichselian population. The contemporaneous Zirphaeu sea deposits can be regarded as the boreal-arctic shallow water equivalent of the arctic Upper Saxicava sand deposits from northern Jutland. The polar bear mandible, however, was deposited on land, as was the metacarpal bone of a brown bear ( Ursus arctos ) from the nearby Nr. Lyngby locality of Allerød age. The overall picture of the Late Weichselian mammal fauna in Denmark shows a mixed composition of different ecotypes. Their sympatric occurrence points at a unique environment not comparable to any now existing, and probably related to the very low latitude of the Weichselian ice sheet.  相似文献   

2.
The skeleton of a young prime adult cave bear, Ursus spelaeus, was found in Chiostraccio Cave (Siena, Tuscany, central Italy), only slightly buried under rock falls. The specimen was dated yielding a conventional age of 24,030 ± 100 14C yr BP (29,200–28,550 cal yr BP), which makes it the latest known representative of the species in Italy. The skeleton was accompanied by the remains of wolf (Canis lupus), wild boar (Sus scrofa), aurochs (Bos primigenius), red deer (Cervus elaphus), roe deer (Capreolus capreolus), bat (Vespertinus murinus), and crow (Corvus monedula). The site seems confirming that the latest Italian U. spelaeus populations shared the risk of intrusion. The association of the cave bear with other animals suggests that the assemblage is an attritional palimpsest of remains of different species not originally associated in life. Cave bears were probably more vegetarian than brown bears and possibly became extinct when plant productivity dropped at the onset of MIS 2. Central and southern Italy may have offered isolated and sheltered refugia for cave bears.  相似文献   

3.
This review provides an up-to-date synthesis of the matrilineal phylogeography of a uniquely well-studied Holarctic mammal, the brown bear. We extend current knowledge by presenting a DNA sequence derived from one of the earliest known fossils of a polar bear (dated to 115 000 years before present), a species that shares a paraphyletic mitochondrial association with brown bears. A molecular clock analysis of 140 mitochondrial DNA sequences, including our new polar bear sequence, provides novel insights into the times of origin for different brown bear clades. We propose a number of regional biogeographic scenarios based on genetic data, divergence time estimates and paleontological records. The case of the brown bear provides an example for researchers working with less well-studied taxa: it shows clearly that phylogeographic models based on patterns of modern genetic variation alone can be substantially improved by including data on historical patterns of genetic diversity in the form of ancient DNA sequences derived from accurately dated samples and by using an approach to divergence-time estimation that suits the data under analysis. Using such approaches it has been possible to (i) establish that the processes shaping modern genetic diversity in brown bears acted recently, within the last three glacial cycles; (ii) distinguish among hypotheses concerning species’ responses to climatic oscillations in accordance with the lack of phylogeographic structure that existed in brown bears prior to the last glacial maximum (LGM); (iii) reassess theories linking monophyletic brown bear populations to particular LGM refuge areas; and (iv) identify vicariance events and track analogous patterns of migration by brown bears out of Eurasia to North America and Japan.  相似文献   

4.
Biostratinomic analysis (processes acting between death and burial) of Lateglacial mammal bone assemblages from three caves in northern England demonstrates the value of re‐examining archived assemblages. With AMS radiocarbon dating of key specimens, these assemblages shed light on the ecology of a region at the northern limit of Lateglacial human activity in Britain. During the Lateglacial Interstadial bears, wolves and humans expanded into the region, bears by around 12 500 14C yr BP, and the earliest evidence for human presence is around 12 300 14C yr BP. At Victoria Cave, wolf activity included predation and scavenging of large ungulates and scavenging bear carcasses apparently resulting from hibernation deaths. The scavenging of bear carcasses is possibly confined to the first part of the Lateglacial Interstadial, whereas evidence for wolf scavenging large ungulates increases later in the Interstadial, after about 11 800 14C yr BP, perhaps reflecting changes in the productivity of the Lateglacial ecosystem, and in human subsistence patterns. The assemblage from Sewell's Cave is wolf den debris from the very end of the Lateglacial Interstadial around 10 800 14C yr BP, whilst that from Kinsey Cave is dominated by large‐bodied carnivores, and is argued to have a quite different taphonomic history. Copyright © 2007 John Wiley & Sons, Ltd.  相似文献   

5.
Paleobiologists generally agree that within the past 10,000 yr North American black bears (Ursus americanus) have decreased in body and tooth size. Some researchers infer that diminution was gradual and continuous; thus, one might infer that a specimen is old if it is larger than an average-size modern bear. Ursid remains recovered in the 1950s from Lawson Cave, Missouri, that are larger than some modern bears have been reported to date to the late Pleistocene, but association with modern taxa, taphonomic considerations, and a radiocarbon date of 200 yr B.P. indicate that they are modern. Modern specimens from Lawson Cave and other parts of the American Midwest are relatively large compared to modern North American black bears from other areas, suggesting that many supposed late Pleistocene bears from the area might be modern also.  相似文献   

6.
The Namur area in Belgium is useful to study brown (Ursus arctos) and cave bears (Ursus spelaeus) as the assemblage contains little temporal and no geographical variation. Here, we aim to assess ontogenetic allometry within cave bears, as well as ecomorphological differences between adult brown bears (n = 9), adult cave bears (n = 5) and juvenile cave bears (n = 3). Landmarks for 3D digitization of the mandible were chosen based on the taphonomical damage of the specimens. Extant brown bears and extinct Pleistocene brown and cave bears were digitized with a Microscribe G2. Generalized Procrustes superimposition was performed on the coordinates. Allometry was studied using regression analysis. Principal component analysis (PCA) was conducted to assess ecomorphological differences between the groups. 61% of the shape variance within juvenile and adult cave bears was predicted by size (n = 8, p < 0.01). The juvenile cave bears have relatively deep horizontal rami. In adult cave bears, the horizontal ramus is much narrower dorsoventrally. Juvenile cave bears have a small masseteric fossa and a short coronoid process, whereas both are larger, relative to mandible size, in adult cave bears. This made juvenile cave bears likely less effective masticators than fully grown cave bears. In the PCA, principal component (PC) 1 accounts for 45.0% of the total variance and PC2 accounts for 27.6%. Fossil U. arctos from Namur fall within the 95% confidence interval of modern North American U. arctos on both PCs, but are more similar to cave bears than the average extant brown bear. From the similarity of fossil and modern brown bears, it can be deduced that the diet of fossil brown bears was probably also within the range of their modern North American conspecifics, although they might have been more efficient at masticating plant matter.  相似文献   

7.
The Saltville Valley of southwestern Virginia contains an abundance of extinct Late Pleistocene megafauna. Recent excavations in the valley produced two particularly notable discoveries, the remains of a scavenged mammoth ( Mammuthus sp.) and a giant short-faced bear ( Arctodus simus ). The bear is the first carnivoran record from Saltville, and its large dental dimensions, dentary proportions, and exceptionally short-face, evince an exceedingly powerful bite for this individual. Mammoth remains are relatively abundant from Saltville, but the material described here preserves extreme examples of scavenging by carnivores. Based on comparisons with the feeding behavior and bone alteration patterns of extant hyenas, cats, wolves, and bears, we conclude that at least two types of large carnivorans scavenged this mammoth carcass. AMS radiocarbon dates of 14 510 ± 55 14C yr BP on the Mammuthus and 14 853 ± 55 14C yr BP on the A. simus demonstrate that they are nearly the same age. Very few direct radiocarbon dates on A. simus have been reported from the contiguous United States. The Saltville A. simus represents the most eastern date for the species, as well as the only date from the Appalachian Highlands.  相似文献   

8.
An isotopic investigation of upper Pleistocene mammal bones and teeth from Scladina cave (Sclayn, Belgium) demonstrated the very good quality of collagen preservation. A preliminary screening of the samples used the amount of nitrogen in whole bone and dentine in order to estimate the preserved amount of collagen before starting the extraction process. The isotopic abundances of fossil specimens from still-extant species are consistent with their trophic position. Moreover, the15N isotopic abundance is higher in dentine than in bone in bears and hyenas, a phenomenon already observed in modern specimens. These results demonstrate that the isotopic compositions of samples from Scladina cave can be interpreted in ecological terms. Mammoths exhibit a high15N isotopic abundance relative to other herbivores, as was the case in Siberian and Alaskan samples. These results suggest distinctive dietary adaptations in herbivores living in the mammoth steppe. Cave bears are clearly isotopically different from coeval brown bears, suggesting an ecological separation between species, with a pure vegetarian diet for cave bear and an omnivorous diet for brown bear.  相似文献   

9.
Upper Pleistocene remains of the Ice Age steppe lion Panthera leo spelaea (Goldfuss, 1810) have been found in the Perick Caves, Sauerland Karst, NW Germany. Bones from many hyenas and their imported prey dating from the Lower to Middle Weichselian have also been recovered from the Perick Cave hyena den. These are commonly cracked or exhibit deep chew marks. The absence of lion cub bones, in contrast to hyena and cave bear cub remains in the Perick Caves, and other caves of northern Germany, excludes the possibility that P. leo spelaea used the cave for raising cubs. Only in the Wilhelms Cave was a single skeleton of a cub found in a hyena den. Evidence of the chewing, nibbling and cracking of lion bones and crania must have resulted from the importation and destruction of lion carcasses (4% of the prey fauna). Similar evidence was preserved at other hyena den caves and open air sites in Germany. The bone material from the Perick and other Central European caves points to antagonistic hyena and lion conflicts, similar to clashes of their modern African relatives.  相似文献   

10.
Qagnaxˆ Cave, a lava tube cave on St. Paul Island in the Pribilofs, has recently produced a mid-Holocene vertebrate faunal assemblage including woolly mammoth, polar bear, caribou, and Arctic fox. Several dates on the mammoth remains converge on 5700 14C yr BP. These dates, ~ 2300 yr younger than mammoth dates previously published from the Pribilof Islands, make these the youngest remains of proboscideans, and of non-extinct Quaternary megafauna, recovered from North America. Persistence of mammoths on the Pribilofs is most parsimoniously explained by the isolation of the Pribilofs and the lack of human presence in pre-Russian contact times, but an additional factor may have been the local existence of high-quality forage in the form of grasses enriched by nutrients derived from local Holocene tephras. This interpretation is reinforced by stable carbon and nitrogen isotope values obtained from the mammoth remains. The endpoint of mammoth survival in the Pribilofs is unknown, but maybe coterminous with the arrival of polar bears whose remains in the cave date to the Neoglacial cold period of ~ 4500 to 3500 14C yr BP. The polar bear record corroborates a widespread cooling of the Bering Sea region at that time.  相似文献   

11.
Doklady Earth Sciences - New finds of brown bear (Ursus arctos L., 1758) fossil remains from the territory of Yakutia, namely, skulls and mandibular bones, have been investigated. The new finds are...  相似文献   

12.
《Quaternary Science Reviews》2007,26(3-4):436-440
Four caves on the SW flank of Mt. Carmel, es Skhul, el Wad, el Jaml and et Tabun, were first excavated in the 1930s by a team led by Dorothy Garrod. They yielded human remains whose age and evolutionary status remain controversial partly because the complexity of the cave deposits invites conflicting interpretations. The abrasion of artefacts and pebbles in el Wad and es Skhul, which was originally ascribed to spring flow within the caves, is explained here by wave action, with the implication that during part of the Middle Palaeolithic the caves were on the shoreline rather than being separated from it—as they now are—by several kilometres of coastal plain and a height difference of some 45 m. U-series, thermoluminescence (TL) and electron spin resonance (ESR) dating suggests that this occurred about 120,000 years ago, when sea level in the eastern Mediterranean stood 5–6 m above its present position. It follows that Mt. Carmel has subsequently undergone some 40 m of uplift. During the period of maximum submergence, the coastal route between Africa and the northern Mediterranean would have been partly blocked, but the loss of the coastal plain for transit and as a source of animal food was offset by easier access from the caves to marine resources.  相似文献   

13.
Tens of thousands of palaeontological and archaeological remains were collected by William Pengelly during 19th century excavations of Kents Cavern, but are now widely dispersed between museums. This has previously precluded spatial analysis. We have now assembled available museum records into a single database, and, using our previously-reconstructed Pengelly excavation map as a base, we have been able to exploit the unique Pengelly location code to set up a GIS mapping system. This allows, for the first time, the analysis of spatial patterns. In addition, the GIS serves to highlight potential problems of recording or curation in the original data. Here we report on the construction of the GIS system and its first use in the analysis of spatial distribution of bear remains. The maps demonstrate that Ursus deningeri entered the cave through a now-sealed High Level Chamber entrance at the back of the cave in the middle Pleistocene, whereas Ursus arctos accessed the cave in the late Pleistocene through the now-sealed Northeast Gallery entrance. The denning areas are reconstructed as Labyrinth/Bear's Den for U. deningeri and Vestibule/Great Chamber for U. arctos. Considerable post-mortem re-distribution of the remains of both species is indicated.  相似文献   

14.
The cave bear ( Ursus spelaeus ) was one of several spectacular megafaunal species that became extinct in northern Eurasia during the late Quaternary. Vast numbers of their remains have been recovered from many cave sites, almost certainly representing animals that died during winter hibernation. On the evidence of skull anatomy and low δ15N values of bone collagen, cave bears appear to have been predominantly vegetarian. The diet probably included substantial high quality herbaceous vegetation. In order to address the reasons for the extinction of the cave bear, we have constructed a chronology using only radiocarbon dates produced directly on cave bear material. The date list is largely drawn from the literature, and as far as possible the dates have been audited (screened) for reliability. We also present new dates from our own research, including results from the Urals. U. spelaeus probably disappeared from the Alps and adjacent areas – currently the only region for which there is fairly good evidence – c . 24 000 radiocarbon years BP ( c . 27 800 cal. yr BP), approximately coincident with the start of Greenland Stadial 3 ( c . 27 500 cal. yr BP). Climatic cooling and inferred decreased vegetational productivity were probably responsible for its disappearance from this region. We are investigating the possibility that cave bear survived significantly later elsewhere, for example in southern or eastern Europe.  相似文献   

15.
Four Quaternary volcanic ash zones in the southern Norwegian Sea have been investigated in core P57-7 from the Iceland Plateau. Both the geochemical composition and morphological variation of each ash layer have been studied. The four volcanic ash zones appear in the light oxygen isotope stages 1, 5, 7 and 11. The ash zones are composed of transparent platy grains, light brown transparent grains, brown blocky and black blocky grains and white/transparent pumice, and each zone shows a distinct stratigraphic evolution. The geochemical results show a mixture of basaltic and silicic grains in each ash zone, and that each zone contains grains from more than one eruption. The geochemical investigations strongly suggest that all the ashes are derived from Iceland. The youngest ash zone includes two layers, which based on their geochemical composition and stratigraphic position are correlated with the 14C-dated Vedde Ash (10 600 yr BP) and Saksunarvatn Ash (9 100 yr BP). Possible sources on Iceland for these layers are discussed.  相似文献   

16.
甘肃庆阳地区沿河谷萨拉钨苏组分布普遍,在庆阳附近和环工子该组底部砂支和上部粘土层陶片。同一层位产哺乳动物化石,其地质时代属晚更新世。^14C测年资料也反映它是一套距今约三万年前的堆积,与根据哺乳动物化石所确定的地质年代相一致。这一发现把陶器出现的年代提前了大约2万年,其意义重大。  相似文献   

17.
The Pinturas Formation is a continental succession, characterized by eolian sediments (mainly tuffaceous) and paleosols, originated in an upland setting of west central Patagonia. The main intraformational erosive surfaces and lithological changes define three sequences. The Formation bears a rich mammalian association (Florentino Ameghino's Astrapothericulan fauna), whose age in relation to the Santacrucian SALMA (middle Miocene) is still controversial. Recent collections from the Pinturas Formation, performed with stratigraphic control, allow differentiation of two distinct hystricognath rodent associations. The lower and middle sequences bear a particular combination of Colhuehuapian (early Miocene) and Santacrucian genera, mostly represented by species exclusively known to the Pinturas Formation (‘Pinturan’ association). The upper sequence bears typical Santacrucian species, more derived than its Pinturan counterparts. According to the rodent record, the lower and middle sequences of the Pinturas Formation are older than the base of the Santa Cruz Formation exposed at Monte Observación and Monte León, and the upper sequence may be correlated with the lowermost levels of the Santa Cruz Formation and deposits exposed at Karaiken that bear Ameghino's ‘Notohippidian’ fauna. These correlations agree with more recent radiometric dates and other biostratigraphic evidence, supporting Ameghino's original hypothesis. The Pinturan rodent assemblage of the lower and middle sequences suggests the presence of humid forests, in accordance with other faunal components and palynological data. Sedimentologic, paleopedologic, and ichnologic evidence, however, suggest environments dominated by herbaceous vegetation. This seeming contradiction is interpreted as the result of a marked environmental gradient due to the paleotopography and/or climatic fluctuations. The mammal record corresponds to the more humid intervals, which have less representation in the sedimentary record. The Pinturas case reveals that more detailed analyses are necessary to understand the relationship between the stressed abiotic Patagonian scenarios and the evolution of their Tertiary vertebrate communities.  相似文献   

18.
Activity ratios of 234U/238U, 230Th/234U, and 230Th/232Th have been determined for calcite, gypsum and halite speleothems from caves of the Nullarbor Plain, mostly in the area N and NW of Mundrabilla Station, for the purpose of U-series dating. All calcite speleothems contain adequate amounts of uranium for dating, but some show an excess of 230Th. Stratigraphic relationships indicate that there were at least three phases of calcium carbonate deposition in the Nullarbor caves. The calcite samples, with one possible exception, have ages in excess of ca. 400000 yrs BP. This suggests that no significant amounts of calcium carbonate deposition have taken place during the last 400ka. At present, active deposition of speleothems is restricted almost entirely to gypsum and halite. The only gypsum speleothem dated was found to have a finite age of ca. 185 ka. Six dates on a small halite speleothem containing insect and arachnid remains indicate that it formed rapidly during Holocene time.  相似文献   

19.
Ninety-four prespawning adult Atlantic sturgeon (Acipenser oxyrinchus) were sampled in the Hudson River for age, sex, body size, gonad weight, fecundity, mature oocyte size, and plasma concentrations of gonadotropins, sex steroids, and vitellogenin during the spring spawning migrations in 1992 and 1993. In males, the age and total length ranged from 12 yr to 19 yr and from 133 cm to 204 cm and in females from 14 yr to 36 yr and from 197 cm to 254 cm. The majority of males were 13–16 yr old, and females were 16–20 yr old. Some females had residual atretic ovarian bodies, presumably remaining from a previous spawning and indicating iteroparity. Pre-ovulatory condition was recognized by migration of the germinal vesicle or by germinal vesicle breakdown and by significantly elevated plasma gonadotropins, progesterone, and vitellogenin. All pre-ovulatory females were captured upriver from Hudson River kilometer 136. Individual fecundity ranged from 0.4 million to 2.0 million eggs and oocyte diameter from 2.4 mm to 2.9 mm, and both characters exhibited a significant (p<0.05) positive relationship with female body size. Iteroparous females, tentatively identified by the presence of atretic bodies remaining in the ovary from a previous spawning, had significantly (p<0.05) higher fecundity and produced larger eggs, compared with females spawning presumably for the first time.  相似文献   

20.
Gypsum (CaSO4·2H2O), alunite (KAl3(SO4)2(OH)6), and rare phosphate–sulphate sanjuanite Al2(PO4)(SO4)(OH) 9(H2O) and rossiantonite (Al3(PO4)(SO4) 2(OH)2(H2O)14) have recently been identified as secondary mineral deposits in different quartz‐sandstone caves in the Gran Sabana region, Venezuela. Due to the extended time scale required for speleogenesis in the hard and barely soluble quartz‐sandstone lithology, these caves are considered to be as old as 20 to 30 My. The study of these peculiar secondary mineral deposits potentially reveals important insights for understanding the interaction between deep, superficial and atmospheric processes over thousands to perhaps millions of years. In this study, chemical and petrographic analyses of potential host rock sources, sulphur and oxygen isotope ratios, and meteorological, hydrological and geographical data are used to investigate the origin of sulphates and phospho–sulphates. The results suggest that the deposition of sulphates in these caves is not linked to the quartz‐sandstone host rock. Rather, these mineral deposits originate from an external atmospheric sulphate source, with potential contributions of marine non‐sea salt sulphates, terrestrial dimethyl sulphide and microbially reduced H2S from the forests or peatbogs within the watershed. Air currents within the caves are the most plausible means of transport for aerosols, driving the accumulation of sulphates and other secondary minerals in specific locations. Moreover, the studied sulphate minerals often co‐occur with silica speleothems of biological origin. Although this association would suggest a possible biogenic origin for the sulphates as well, direct evidence proving that microbes are involved in their formation is absent. Nonetheless, this study demonstrates that these quartz‐sandstone caves accumulate and preserve allogenic sulphates, playing a yet unrecognized role in the sulphur cycle of tropical environments.  相似文献   

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