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1.
With the aim of studying the interspectific differences in the bioaccumulation of a series of elements in six species of estuarine macrophytes, we measured the concentrations of various metals (Co, Cr, Cu, Fe, Mn, Ni, Pb, and Zn), as well as C and N, in several species of macroalgae and one species of seagrass in estuaries in northwest Spain. To facilitate the interpretation of the results, we also determined the concentrations of the same metals in sediments. Contamination by particulate metal did not appear to be important. The levels of Cu in the different species were generally high in comparison with those found in similar studies; in contrast, the concentrations of Pb and Zn were relatively low. The metal concentrations showed greater variability and a distibution that was further from normal than those of C and N. In general the highest concentrations of elements were found inEnteromorpha spp. andZostera noltii, and the lowest occurred inUlva spp. There were several significant positive correlations among tissue concentrations of Cr, Cu, and Ni. There were also significant correlations between tissue concentrations of N and Zn, suggesting possible synergetic effects between them. Cluster analysis clearly revealed that the chemical composition ofZ. noltii differed from that of the macroalage. Macroalgae were not grouped according to their taxonomic position, therefore it appears that the concentrations of the elements depended more on the ecological traits of the macroalgal species rather than on any possible, physiological differences among different taxonomic groups. Of the species studied,Ascophyllum nodosum andEnteromorpha spp. appeared to be the best biomonitors of metal contamination.  相似文献   

2.
The Hercynian mountain ranges were islands of mountain glaciation and alpine tundra in a Central European ice‐free corridor during the Late Pleistocene. Today they are notable areas of glacial landforms, alpine‐forest free areas, peatlands and woodlands. However, our knowledge of the Lateglacial and early Holocene environmental changes in this region is limited. We present a new multi‐proxy reconstruction of a mid‐altitude environment in the Bohemian Forest spanning this period. A core (5.2 m length) in the ?erné Lake cirque (1028 m a.s.l.) was subjected to lithological, geochemical, pollen and macrofossil analysis supplemented by two optically stimulated luminescence (OSL) and 10 AMS radiocarbon dates. We determined the impact of regional and supraregional climate changes on the environment. The two most significant changes in sedimentation during the Lateglacial (17.6 and 15.8–15.5 cal. ka BP) were synchronous with regional glacial chronostratigraphy. Unlike Central European mountain ranges, in the Bohemian Forest the Younger Dryas was not coincident with glacier re‐advance, but was a dry, cold episode with low lake levels, which prevailed until the early Preboreal. Plant macrofossils indicate local establishment of Betula nana and Betula pendula/pubescens at 15.4–13.4 cal. ka BP. Comparison with Holocene records from Central Europe shows a similar immigration history of vegetation at mid and higher altitudes. The tree line exceeded an altitude of ~1000 m a.s.l. around 10.5 cal. ka BP and coincided with rapid geochemical changes in the sediment. The 8.2 ka BP event did not have any response in the sedimentary record, but corresponded to stabilization of the Picea abies population and expansion of Fagus. Fagus colonized the Bohemian Forest earlier than other Hercynian mid‐mountains, but never predominated in the composition of the forest at higher elevations. Abies alba was the last tree species that immigrated to the study area.  相似文献   

3.
Previously only three terrestrial interglacial periods were known from southern Scandinavia, all of which could be relatively easily correlated within the central European stratigraphical framework. Here, we present a new interglacial–interstadial pollen, plant macrofossil and charcoal record from Trelde Klint, Denmark, and analyse its biostratigraphy, correlation with other European records, vegetation development, fire dynamics and absolute dating. Except for a slight truncation of the early part of the record, the pollen stratigraphy exhibits a full interglacial succession, including temperate trees (Quercus, Ulmus and Tilia) during its mesocratic stage. Macrofossil analysis allowed identification to species level for Quercus robur, Picea abies and two mosses. Conifers (Pinus and Picea) dominate the pollen record of the interglacial sequence, and the occurrence of Larix pollen in the top part of the interglacial record as well as in the interstadial sediments is especially indicative of this interglacial. The overall diversity of tree genera is rather low. These biostratigraphical features suggest that Trelde Klint is unique among Danish records, but it is similar to records from northern Germany. Numerical analyses (REVEALS and DCA) indicate that forests during the temperate stage were dense and that vegetation openness increased only towards the end of the interglacial, accompanied by increased fire occurrence. A short interstadial sequence with a dominance of Pinus and Betula and the presence of Larix is present above the interglacial deposit. We argue that lack of attention to differences in fire regimes may hamper understanding of between‐site correlations of interglacial pollen records. OSL dating, using a novel feldspar technique, yields an average age of 350±20 ka for the sandy sediments above the interglacial layers at Trelde Klint, suggesting that the whole interglacial–interstadial succession belongs to Marine Isotope Stage 11.  相似文献   

4.
Although body fossils of shorebirds and shorebird-like species are extremely rare from the Cretaceous, rapid increase in the discovery of bird footprint sites provides valuable alternate evidence to help fill gaps in the story of the early evolution of shorebird-like species. Newly discovered bird tracks from the Albian-Cenomanian Dakota Formation in northeastern Utah represent the first report of the ichnogenus Koreanaornis from North America and only the second report of bird tracks from this formation. These tracks are not attributable to Aquatilavipes as previously claimed. Three well-preserved trackways are described and provisionally referred to Koreanaornis cf. hamanensis (Kim). However, a review of the ichnotaxonomy of shorebird ichnites reveals that this ichnotaxon also closely resembles the Miocene ichnospecies Avipeda sirin (Vyalov). This latter comparison points to the need for a thorough evaluation of the similarity between Mesozoic and Cenozoic avian ichnotaxa, which may be over-split in some cases and under-differentiated in others.The new material helps distinguish ichnogenus Koreanaornis from the larger bird track Aquatilavipes, which is more abundant and widespread in North America. In some cases Aquatilavipes has been incorrectly used as a catch-all ichnogenus both in North America and Asia. The Dakota Formation stratigraphy at the tracksite indicates that the track makers lived in a marginal marine paleoenvironment. However, despite the widespread distribution of such facies, often replete with dinosaur tracks, the bird track record of the Dakota Formation, and the Cretaceous of the western USA remains relatively sparse in comparison with other areas such as east Asia.  相似文献   

5.
The orthophragminids in lower Bartonian Reineche Limestone member, a fossiliferous shallow-marine unit exposed in Cap Bon peninsula in Tunisia, are represented by 17 species assigned to Discocyclinidae Galloway 1928 and Orbitoclypeidae Brönnimann 1946. These taxa, associated with nummulitids and alveolinids, belong to the lineages of Discocyclina Gümbel 1870, Nemkovella, 1987, Orbitoclypeus Silvestri 1907, and Asterocyclina Gümbel 1870, described for the first time from north Africa lying at the southern margin of Tethyan ocean during Paleogene. We identified Nemkovella evae, previously not recorded in upper Lutetian/lower Bartonian and younger Eocene deposits of northern Tethyan platforms, and erected a new subspecies, N. evae reinechensis n. ssp. A comparison of Reineche orthophraminids, assigned to orthophragmines zone (OZ) 12 and shallow benthic zone (SBZ 17), to the well-described coeval assemblages at northern Tethyan platforms in Italy, Hungary, Turkey, and to those in Kutch Basin in the Indian subcontinent suggests that some species are confined to certain paleogeographic domains. Orbitoclypeus haynesi, the only orbitoclypeid and the most abundant orthophragminid in lower Bartonian deposits in Kutch, appears to be the most common orbitoclypeid in Reineche Limestone. In Europe, this species is not known and is replaced by Orbitoclypeus varians, the most common orbitoclypeid in middle Eocene of central Europe. Both species occur in varying proportions in marine successions in Turkey. Asterocyclina sireli, identified so far only in Turkey, occurs in Reineche Limestone and in lower Bartonian deposits in Kutch. This species is recorded for the first time in the Indian subcontinent. Relying on present study, as well as our recent studies in Kutch Basin, we conclude that the generic and specific diversity of orthophragminids decreases eastward from the peri-Mediterranean region to Indian subcontinent and to the western Pacific.  相似文献   

6.
A single confidently dated species of cephalopod is so far known in the Tremadocian of the southern Central Andean Basin (NW Argentina and southern Bolivia). This species belongs to the Eothinoceratidae and has a strong affinity mainly with Avalonia. During the Floian, a notable increase in diversity took place, with the appearance of a variety of families represented by several genera, in particular, within the Family Eothinoceratidae. In addition to the previously described species from southern Bolivia, we evaluate the other records of that family from the Central Andean Basin, and propose the following new taxa: Saloceras sikus sp. nov., Saloceras quena sp. nov., Mutveiceras gen. nov., and Mutveiceras cienagaensis sp. nov. We also describe Margaritoceras diploide, Margaritoceras sp., and Mutveiceras sp. From a palaeogeographic perspective, the cephalopod fauna shows affinities mainly with those of England, Wales, and the Montagne Noire (cold water Gondwana and peri‐Gondwana). As with other cephalopod faunas of mid to high palaeolatitudes, eothinoceratids occur along with other cephalopods forming assemblages of low morphological diversity. We interpret the forms described here as demersal with a subvertical poise, but capable of making rapid buoyancy changes, living in a wide spectrum of shallow offshore to shoreface settings. Copyright © 2014 John Wiley & Sons, Ltd.  相似文献   

7.
We present a database of late-Quaternary plant macrofossil records for northern Eurasia (from 23° to 180°E and 46° to 76°N) comprising 281 localities, over 2300 samples and over 13,000 individual records. Samples are individually radiocarbon dated or are assigned ages via age models fitted to sequences of calibrated radiocarbon dates within a section. Tree species characteristic of modern northern forests (e.g. Picea, Larix, tree-Betula) are recorded at least intermittently from prior to the last glacial maximum (LGM), through the LGM and Lateglacial, to the Holocene, and some records locate trees close to the limits of the Scandinavian ice sheet, supporting the hypothesis that some taxa persisted in northern refugia during the last glacial cycle. Northern trees show differing spatio-temporal patterns across Siberia: deciduous trees were widespread in the Lateglacial, with individuals occurring across much of their contemporary ranges, while evergreen conifers expanded northwards to their range limits in the Holocene.  相似文献   

8.
We reconstruct the Holocene range fluctuations of the European pond turtle (Emys orbicularis) for northern and Central Europe. Based on 529 subfossil records and radiocarbon dating of critical finds, we provide evidence that E. orbicularis experienced a rapid range expansion with Holocene warming. Phylogeographic data argue for a colonization of northern and Central Europe from a glacial refuge located in the south-eastern Balkans. Already in the Late Preboreal (9100–8600 cal BC) the species occurred in northern Central Europe and was widespread in Denmark and southern Sweden (Skåne) during the Boreal (8600–7100 cal BC). The maximum range extension occurred during the Holocene climatic optimum (Atlantic: 7100–3750 cal BC), with records in southern England, middle Sweden, and northern Estonia. After the early Subboreal (3750–1750 cal BC) northern populations collapsed, coinciding with a climatic cooling and oceanization. Northern populations were most likely established via long distance dispersal (rafting using rivers debouching in the North and Baltic Seas); after northern Europe was reached, a minimum dispersal of 50 km per 100 years may be hypothesized for the Boreal.  相似文献   

9.
In statistically optimised P–T estimation, the contributions to overall uncertainty from different sources are represented by ellipses. One source, for a diffusion‐controlled reaction at non‐equilibrium, is diffusion modelling of the reaction texture. This modelling is used to estimate ratios, Q, between free‐energy differences, ΔG, of reactions among mineral end‐members, to replace the equilibrium condition ΔG = 0. The associated uncertainty is compared with those already inherent in the equilibrium case (from end‐member data, activity models and mineral compositions). A compact matrix formulation is introduced for activity coefficients, and their partial derivatives governing error propagation. The non‐equilibrium example studied is a corona reaction with the assemblage Grt–Opx–Cpx–Pl–Qtz. Two garnet compositions are used, from opposite sides of the corona. In one of them, affected by post‐reaction Fe, Mg exchange with pyroxene, the problem of reconstructing the original composition is overcome by direct use of ratios between chemical‐potential differences, given by the diffusion modelling. The number of geothermobarometers in the optimisation is limited by near‐degeneracies. Their weightings are affected by strong correlations among Q ratios. Uncertainty from diffusion modelling is not large in comparison with other sources. Overall precision is limited mainly by uncertainties in activity models. Hypothetical equilibrium P–T are also estimated for both garnet compositions. By this approach, departure from equilibrium can be measured, with statistical uncertainties. For the example, the result for difference from equilibrium pressure is 1.2 ± 0.7 kbar.  相似文献   

10.
In different-aged sand-fixing zones and shifting sand dunes, the seasonal variations of foliar carbon isotope discrimination (Δ), potassium (K) concentration and ash content were investigated in three dominated deserted plants (Artemisia ordosica, Hedysarum scoparium and Caragana korshinskii) during growth season in Shapotou, which is in the southeast margin of the Tengger Desert. The correlations of foliar Δ with foliar K concentration and ash content were examined to evaluate the foliar K and ash content as surrogates of Δ in those deserted plants. Results showed that there were significant effects of plant species, micro-habitation and growth season on foliar Δ, K concentration and ash content. Foliar Δ of C. korshinskii was significantly lower than those of A. ordosica and H. scoparium, and K concentrations in A. ordosica were 2.14 and 2.36 times those of C. korshinskii and H. scoparium. At the same time, micro-habitation and the conditions in growth seasons had significant effects on foliar Δ, K concentration and ash content. Ash content and K concentration were positively correlated to Δ in A. ordosica and H. scoparium, while there was significantly negative relationship between foliar K concentration and Δ in C. korshinskii. Thus, those findings suggest that foliar ash content and K concentration can serve as surrogates of carbon isotope discrimination (Δ) in A. ordosica and H. scoparium, while they do not in C. korshinskii. This result implied that the correlations of foliar Δ with ash content and K concentration were various due to the physiological features of plant species, and species differences should be fully considered when evaluating the surrogates of carbon isotope discrimination in plants.  相似文献   

11.
During the late Pleistocene and early Holocene, Bison was widely dispersed across North America and occupied most regions not covered by ice sheets. A dietary study on Bison paleopopulations from Alaska, New Mexico, Florida, and Texas was performed using two methods that relate dental wear patterns to diet, mesowear analysis and microwear analysis. These data were compared to a mixed sample of extant Bison from the North American central plains, extant wood Bison from Alberta (Canada) and a variety of other modern ungulates. Mesowear relates macroscopic molar facet shape to levels of dietary abrasion. The mesowear signature observed on fossil Bison differs significantly from the hyper-abrasive grazing diet of extant Bison. Tooth microwear examines wear on the surface of enamel at a microscopic scale. The microwear signal of fossil samples resembles to modern Bison, but the fossil samples show a greater diversity of features, suggesting that fossil Bison populations regularly consumed food items that are texturally inconsistent with the short-grass diet typical of modern plains Bison. Mesowear and microwear signals of fossil Bison samples most closely resemble a variety of typical mixed feeding ungulates, all with diets that are substantially less abrasive than what is typical for modern plains Bison. Furthermore, statistical tests suggest significant differences between the microwear signatures of the fossil samples, thus revealing geographic variability in Pleistocene Bison diets. This study reveals that fossils are of value in developing an understanding of the dietary breadth and ecological versatility of species that, in recent times, are rare, endangered, and occupy only a small remnant of their former ranges.  相似文献   

12.
The biochronology of Cenomanian-early Turonian ammonite faunas from three key stratotype areas (north-west Europe, central Tunisia and the Western Interior of North America) has been analysed and revised by utilizing the unitary association method. This review is prompted by the huge amount of biostratigraphic data published during recent decades and by a taxonomic homogenisation of the ammonite faunas from these key areas. The Cenomanian and lower Turonian of Tunisia comprise twenty-four Unitary Association zones and the middle Cenomanian-lower Turonian of the Western Interior Basin twenty-three such zones. The unitary association method means a two-fold increase in resolution of these ammonite zonations compared to the standard, empirical schemes. Central Tunisia and the Western Interior are correlated with north-west Europe by constructing a zonation including all taxa common to these areas. These correlations highlight the variable completeness and resolution of the faunal record through space and time, and reveal a significant number of diachronous taxa between the three areas. These correlations enable the designation of a new global marker for the middle/upper Cenomanian boundary, which is characterised by the disappearance of the genera Turrilites, Acanthoceras and Cunningtoniceras and by the appearance of Eucalycoceras, Pseudocalycoceras and Euomphaloceras. The only synchronous datum known is the last occurrence of Turrilites acutus, which may thus be used as a marker for the middle/upper Cenomanian boundary, provided that it does not turn out to be diachronous in the light of any new data.  相似文献   

13.
Well-preserved Ginkgo pollen organs are analyzed from the Middle Jurassic Xishanyao Formation of the Turpan–Hami Basin, Xinjiang Uygur Autonomous Region, northwestern China, and are described as a new species, Ginkgo hamiensis Z.X. Wang et B.N. Sun sp. nov. The immature male cones are cylindrical and catkin-like, with two longitudinal stripes on the stalk. The pollen sacs are shaped like a long oval with two pollen sacs fused together for each microsporophyll, and the microsporophyll tip is a triangular cystidium. The pollen grains are oblong or fusiform and monocolpate; both ends are blunt or sharp. By comparison with previously reported fossil records of Ginkgo plants, we determined that the current fossils are different from all other reported species; thus, the present fossil is referred to as a new species of Ginkgo. The reproductive organs of the Ginkgo fossils described herein can provide valuable information for the study of Ginkgo plants. Further, there are two probable evolutionary trends in the Ginkgo pollen cones. One trend is that the number of pollen sacs changed from three or four during the Jurassic and Cretaceous to two at the present day; the other is that the number of pollen sacs has remained two from the Middle Jurassic to the present day. In addition, the pollen cones described herein are similar to the pollen cones of the extant Ginkgo, which strongly indicates that the morphology of Ginkgo plants may have remained highly conserved over millions of years.  相似文献   

14.
A broad-scale survey of seagrass species composition and distribution along Florida's central Gulf Coast (known as the Big Bend region) was conducted in the summer of 2000 to address growing concerns over the potential effects of increased nutrient loading from adjacent coastal rivers. Iverson and Bittaker (1986) originally surveyed seagrass distribution in this region between 1974–1980. We revisited 188 stations from the original survey, recording the presence or absence of all seagrass species. Although factors such as accuracy of station relocation, differences in sampling effort among studies, and length of time between surveys preclude statistical comparisons, several interesting patterns emerged. While the total number of stations occupied by the three most common seagrass species,Thalassia testudinum, Syringodium filiforme, andHalodule wrightii, was similar between the two time periods, we observed a change in the number of records of each species as well as changes in distribution with depth.T. testudinum andHalophila engelmanni occurrence declined in the deepest areas of the region, while the number of stations occupied byS. filiforme andH. wrightii increased in nearby areas. We observed several localized areas of seagrass loss, frequently associated with the mouths of coastal rivers. These results suggest that increased nutrient loading to coastal rivers that discharge into the Big Bend area may be affecting seagrasses by increasing phytoplankton abundance in the water column, thus changing water clarity characteristics of the region.  相似文献   

15.
The Agenian is the earliest Neogene European Land Mammal Age. It encompasses the mammalian zones MN1 (23.03–22.7 Ma) and MN2 (22.7–20.0 Ma) and roughly coincides with the Aquitanian standard age. Agenian mammalian assemblages from Western Europe encompass a mixture of rhinocerotid taxa of Oligocene affinities and of Miocene newcomers, mostly recorded in France, Germany, Switzerland, and to a lesser extent, Spain. Rhinocerotidae are documented by seven species referred to five genera (Pleuroceros pleuroceros, Protaceratherium minutum, Plesiaceratherium aquitanicum, Mesaceratherium paulhiacense, Diaceratherium lemanense, D. asphaltense, and D. aginense), further attesting to a low suprageneric diversity. Their systematics, morphology, ecology, stratigraphical and geographical ranges are detailed in the present article. Occurrences and geographical ranges of all seven rhinocerotid species are illustrated on palaeogeographical maps of the circum-Mediterranean region at 23 Ma (MN1) and 21 Ma (MN2). The richest Agenian localities (Paulhiac, MN1; Laugnac, MN2) record a specific diversity similar to that of Orleanian rhinocerotid assemblages, with up to five/six associated species. All Agenian rhinocerotid species from Western Europe are endemic to the concerned region, which is consistent with the complete geographic isolation of Western Europe by earliest Miocene times. However, all five genera are documented by twin species in coeval localities of South and Central Asia, which implies (1) vicariant speciation events by latest Oligocene times and (2) the existence of intermittent pathways for terrestrial megamammals such as rhinocerotids during the concerned interval.  相似文献   

16.
Core P1‐003MC was retrieved from 851 m water depth on the southern Norwegian continental margin, close to the boundary between the Norwegian Current (NC) and the underlying cold Norwegian Sea Deep Water. The core chronology was established by using 210Pb measurements and 14C dates, suggesting a sampling resolution of between 2 and 9 yr. Sea‐surface temperature (SST) variations in the NC are reconstructed from stable oxygen isotope measurements in two planktonic Foraminifera species, Neogloboquadrina pachyderma (d.) and Globigerina bulloides. The high temporal resolution of the SST proxy records allows direct comparison with instrumental ocean temperature measurements from Ocean Weather Ship (OWS) Mike in the Norwegian Sea and an air temperature record from the coastal island Ona, western Norway. The comparison of the instrumental and the proxy SST data suggests that N. pachyderma (d.) calcify during summer, whereas G. bulloides calcify during spring. The δ18O records of both species suggest that the past 70 yr have been the warmest throughout the past 600 yr. The spring and summer proxy temperature data suggest differences in the duration of the cold period of the Little Ice Age. The spring temperature was 1–3°C colder throughout most of the period between ca. AD 1400 and 1700, and the summer temperature was 1–2°C colder throughout most of the period between ca. AD 1400 and 1920. Fluctuations in the depth of the lower boundary of the NC have been investigated by examining grain size data and benthic foraminiferal assemblages. The data show that the transition depth of the lower boundary of the NC was deeper between ca. AD 1400 and 1650 than after ca. AD 1750 until present. Copyright © 2003 John Wiley & Sons, Ltd.  相似文献   

17.
We characterized the abundance and species composition of sessile and mobile epifaunal assemblages in the York River, a tributary of the Chesapeake Bay, U.S., during the summer hypoxia seasons in 1996 and 1997. We collected communities on artificial substrates in two areas of the river that have historically experienced different exposure to hypoxia. Despite frequent hypoxic stress, epifauna formed dense communities in both areas. Dominant species comprised a range of phyla and included the polychaetesPolydora cornuta andSabellaria vulgaris, the bryozoansMembranipora tenuis andConopeum tenuissimum, the tunicateMolgula manhattensis, the barnacleBalanus improvisus, the anemoneDiadumene leucolena, and the hydroidsEctopleura dumortieri andObelia bicuspidata. Common mobile species included the nudibranchsCratena kaoruae andDoridella obscura, the amphipodsMelita nitida andParacaprella tenuis, the polychaeteNereis succinea, and the flatwormStylochus ellipticus. We found few differences in species composition between the two areas, even though one area usually experienced lower oxygen concentrations during hypoxic events, suggesting that hypoxia does not exclude any epifaunal species, in the York River. We did find differences between the two study areas in percent cover and abundance of some species. While tunicates, hydroids, and anemones were equally abundant in both areas during both study years, bryozoans and the polychaeteS. vulgaris were more abundant in the area with gereally higher oxygen, suggesting that they may be less tolerant of hypoxic stress. The polychaeteP. cornuta was more abundant in the area that usually had lower oxygen. These results suggest that many epifaunal species have high hypoxia tolerance, and most epifaunal species found in the lowr York River are able to survive in hypoxic areas. Epifaunal species are not necessarily more susceptible to hypoxia than infaunal species in the York River. Epifaunal communities in areas with brief hypoxic episodes and moderate hypoxia (0.5–2 mg O2 I−1) can persist with little change in species composition, and with few changes in abundance, as oxygen concentrations fall.  相似文献   

18.
Pleistocene faunas from south China are difficult to subdivide based on the long temporal ranges of many taxa and a reduced number of genera in comparison to faunas from temperate north China. In south China, the Ailuropoda–Stegodon fauna is a very general one and includes a relatively stable suite of genera that apparently persisted for long periods of time. These attributes have made constraining its time range difficult. Application of electron spin resonance (ESR) dating of tooth enamel constrains the ages well where uranium uptake was minor. Where uranium uptake into teeth was significant, an approach combining ESR and 230Th/234U isotopic analysis also yields excellent ages. Previous estimates of early, middle and late Pleistocene time ranges previously determined by biostratigraphic seriation for the Ailuropoda–Stegodon fauna are confirmed in all cases but are made more precise with our approach, including specific time ranges for certain archaic taxa. Absolute dating also yields an extended time range for Gigantopithecus blacki of 1200 to 310 ka.  相似文献   

19.
It has been hypothesized that refugia of thermophilous tree species were located in Northern Italy very close to the Alps, though, this hypothesis has yet to be tested thoroughly. In contrast to Central and Southern Italy with its relative wealth of data, only a few fragmentary records are currently available from Northern Italy for the last Glacial (Würm, Weichselian). Our new study site Lago della Costa lies adjacent to the catchment of the megafans of the Alpine forelands and the braided rivers of the Northeastern Po Plain that have so far inhibited the recovery of continuous Glacial and Late-Glacial records. We analyze pollen, plant macrofossils, charcoal and ostracods to reconstruct the vegetation, fire and lake history for the period 33,000–16,000 cal. BP. We compare our data with Glacial records from Southern Europe to discuss similarities and dissimilarities between these potential refugial areas. A comparison with independent paleoclimatic proxies allows to assess potential linkages between environmental and climatic variability. New macrofossil and pollen data at Lago della Costa unambiguously document the local persistence of boreal tree taxa such as Larix decidua and Betula tree species around the study site during the last Glacial. The regular occurrence of pollen of temperate trees in the organic lake sediments (fine-detritus calcareous gyttja) suggests that temperate taxa such as Corylus avellana, Quercus deciduous, Tilia, Ulmus, Fraxinus excelsior, Carpinus, Abies alba and Fagus sylvatica, most likely survived the Last Glacial Maximum (LGM) at favorable sites in the Euganean Hills. The percentage values of temperate trees are comparable with those from Southern Europe (e.g. Monticchio in Southern Italy). We conclude that the Euganean Hills were one of the northernmost refugial areas of temperate taxa in Europe. However, the relative and absolute abundances of pollen of temperate trees are highly variable. Pollen-inferred declines of temperate tree communities (e.g. Quercetum mixtum) and low ostracod-inferred water levels at Lago della Costa correspond to the cold Heinrich events H-2 (LGM; 23,000–19,000 cal. BP) and H-3 (around 28,000 cal. BP), as recorded in the marine sediments of the North Atlantic. Similar patterns of significant temperate tree population collapses during cold Heinrich events are recorded at southern Mediterranean sites (e.g. Monticchio and the Alboran Sea). These findings suggest close linkages between Northern Atlantic and South-Central European climates during the past Glacial.  相似文献   

20.
Rapid rates of species range extension during the Holocene represent seed dispersal distances of at least 6 to 8 km per generation for North American species of Fagus, Quercus, and Carya, taxa whose fruits are heavy nuts. Occasional seed dispersal by biotic seed predators is necessary for these dispersal distances. One likely agent for dispersal across long distances and habitat discontinuities was the extinct (since 1914 A.D.) passenger pigeon (Ectopistes migratorius). This bird's abundance, capacity for delayed digestion, and nomadic habits strongly suggest an occasional seed dispersal role, although a coevolutionary seed dispersal relationship cannot be inferred from available evidence. The capacity of some heavy seeds for dispersal distances greater than those of most light, wind-dispersed seeds has biogeographic and genetic implications.  相似文献   

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