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Turbulence measurements in fine-scale phytoplankton layers (∼1 to ∼10 m) in the Gulf of Aqaba (Red Sea) were used to evaluate mechanisms of layer formation, maintenance, and breakdown. Simultaneous profiles of chlorophyll a (Chl a) fluorescence and temperature microstructure were measured in the upper 40 m of a 430 m water column over a 16-d period, using a Self Contained Autonomous MicroProfiler (SCAMP). Layers of concentrated phytoplankton were identified in 95 of the 456 profiles. The layers were situated in density stratified regions between 15 and 38 m depth and were characterized by intensities of 0.1 to 0.35 μg Chl a L−1 (as much as two times background concentrations) and an average thickness of 10 m. We show that turbulent mixing and isopycnal displacements associated with internal waves modulated the thickness of the layers. Variations in mixing rates within layers were connected to the vertical structure of the stratified turbulence and the stage of layer development. The breakdown of a persistent phytoplankton layer was tied to strong turbulent mixing at the base of the surface mixed layer, which encroached on the layer from above. Hydrographic observations and scaling analysis suggest that the layers most likely formed in horizontal intrusions from the adjacent coastal region. The cross-shore propagation of phytoplankton-rich intrusions may have important implications for the trophic state of offshore planktonic communities.  相似文献   

3.
The distribution of dissolved lead in the coastal waters of the East China Sea was investigated seasonally. The average concentrations in surface waters during the spring and autumn were 0.52 nM and 0.27 nM, respectively. In the spring, the concentration of dissolved Pb in the surface waters and bottom waters ranged from 0.13 to 1.86 nM and from 0.15 to 0.94 nM, respectively. For both the surface water and the bottom water, the highest values were observed at the Yangtze River Estuary. Seasonal variability of D-Pb between spring and autumn in the ECS was observed. These results suggested that riverine inputs and atmospheric inputs may be the main sources of lead in this area, while adsorption and co-precipitation on suspended particles at the river estuary and biological process may be the major sinks.  相似文献   

4.
We investigate mixing processes under stratified conditions on the Northwest European Continental shelf using a numerical model (POLCOMS). Our results indicate that convection induced by vertical shearing of horizontal density gradients (‘shear-induced convection’) is a regularly occurring feature in the bottom and surface boundary layers in this open shelf-sea situation. Two types of turbulence models are investigated to study their capability for reproducing the observed location of tidal mixing fronts, and the physical processes occurring in seasonally stratified waters. The first model is a one-equation variant of the Mellor–Yamada model, whereas the second model combines a more recent second-momentum closure with a two-equation model. It is found that generally mean frontal positions (as estimated from ICES data) are predicted more accurately by the two-equation model. The one-equation model reproduces the mean frontal locations to 18.1 km (<3 grid spacings) and the two-equation model to 17.1 km; although in the Celtic Sea the accuracy is ∼33 and ∼12 km, respectively. Comparison with historical tide gauges, current metres, CTD stations, and thermistor chain data from the North Sea Project all show an improvement in accuracy when the two-equation model is used. This is particularly apparent in the model's ability to reproduce the spring–neap variability during stratification. We find that in the presence of shear-induced convection the routinely applied clipping of the turbulent length-scale, previously thought to be a minor ingredient in a turbulence model, has a dramatic effect on the results: if the length-scale clipping is not applied, substantial over-mixing is observed to occur. The causes and possible remedies of this effect are investigated. Overall our results demonstrate a sensitivity to the details of the turbulence model that is significantly greater than previously thought.  相似文献   

5.
The interaction of multiple inflows entering a reservoir (Thomson Reservoir, Australia) is investigated with a field experiment and three-dimensional numerical model simulations. The focus of the study is the mixing and transport patterns within the reservoir of the inflowing water. Data from the field experiment showed the existence of multiple inflows intruding horizontally into the reservoir immediately below the thermocline and selectively propagating into the reservoir at a depth determined by the individual density of each inflow. The results serve to show that inflows slot into a stratified reservoir in an orderly fashion, their depth dependent on the separation of inflow densities and reservoir bed, with minimal mixing between intrusions. Inflows thus do not take their nutrient load into the reservoir as a whole, but rather slot their load into an intrusion layer at an appropriate depth. Nutrients brought in via inflows become accessible only when vertical mixing, either directly at the base of the surface layer, or indirectly via the benthic boundary layer flux, transports the intrusion waters into the surface layer.  相似文献   

6.
Nutrients from the Mississippi/Atchafalaya Rivers greatly stimulate biological production in the ‘classical’ food web on the inner shelf of the northern Gulf of Mexico. Portions of this production, especially large diatoms and zooplankton fecal pellets, sink and decompose in the bottom water, consuming oxygen and contributing to the annual development of an extensive zone of bottom water hypoxia, typically >15,000 km2 since 1993. The microbial food web is also active in the Mississippi River plume, but consists of small organisms that sink slowly. This ‘recycling’ food web has not been considered as a significant contributor to vertical flux and hypoxia. However, gelatinous zooplankton, especially pelagic appendicularians such as Oikopleura dioica, mediate the conversion of microbial web organisms to organic particles with high sinking rates. When pelagic appendicularians are abundant in coastal regions of the northern Gulf of Mexico, they stimulate the rapid vertical transfer of microbial web productivity in the surface layer, which is only 5–15 m thick in the coastal hypoxic region, to the sub-pycnocline layer that becomes hypoxic each summer. In this paper we present results from two studies examining the significance of this pathway. In both 2002 and 2004, we observed high production rates of appendicularians in coastal waters. Discarded gelatinous houses and fecal pellets from the appendicularian populations often provided more than 1 g m−2 d−1 of organic carbon for the establishment and maintenance of hypoxia in the northern Gulf of Mexico. This source of organic matter flux is especially important in regions far from the river plumes and during periods of low river discharge. Autotrophic elements of this food web are primarily supported by recycled inorganic nutrients originating in the Mississippi and Atchafalaya Rivers. Sources of dissolved organic matter (DOM) supporting the heterotrophic components of this microbial food web may include in situ production, the Mississippi/Atchafalaya Rivers, and Louisiana's coastal wetlands. If significant, the latter source provides a possible link between Louisiana's high rates of coastal land loss and the large hypoxic zone observed along the coast during summer. Both of the latter DOM sources are independent of phytoplankton production stimulated by inputs of riverine inorganic nutrients.  相似文献   

7.
8.
Vertical distributions of chlorophyll in deep, warm monomictic lakes   总被引:1,自引:0,他引:1  
The factors affecting vertical distributions of chlorophyll fluorescence were examined in four temperate, warm monomictic lakes. Each of the lakes (maximum depth >80 m) was sampled over 2 years at intervals from monthly to seasonal. Profiles were taken of chlorophyll fluorescence (as a proxy for algal biomass), temperature and irradiance, as well as integrated samples from the surface mixed layer for chlorophyll a (chl a) and nutrient concentrations in each lake. Depth profiles of chlorophyll fluorescence were also made along transects of the longest axis of each lake. Chlorophyll fluorescence maxima occurred at depths closely correlated with euphotic depth (r 2 = 0.67, P < 0.01), which varied with nutrient status of the lakes. While seasonal thermal density stratification is a prerequisite for the existence of a deep chlorophyll maximum (DCM), our study provides evidence that the depth of light penetration largely dictates the DCM depth during stratification. Reduction in water clarity through eutrophication can cause a shift in phytoplankton distributions from a DCM in spring or summer to a surface chlorophyll maximum within the surface mixed layer when the depth of the euphotic zone (z eu) is consistently shallower than the depth of the surface mixed layer (z SML). Trophic status has a key role in determining vertical distributions of chlorophyll in the four lakes, but does not appear to disrupt the annual cycle of maximum chlorophyll in winter.  相似文献   

9.
Although large-scale tidal and inertial motions dominate the kinetic energy and vertical current shear in shelf seas and ocean, short-scale internal waves at higher frequencies close to the local buoyancy frequency are of some interest for studying internal wave breaking and associated diapycnal mixing. Such waves near the upper limit of the inertio-gravity wave band are thought to have relatively short O (102–103 m) horizontal scales and to show mainly up- and downward motions, which contrasts with generally low aspect ratio large-scale ocean currents. Here, short-term vertical current (w) observations using moored acoustic Doppler current profiler (ADCP) are presented from a shelf sea, above a continental slope and from the open ocean. The observed w, with amplitudes between 0.015 and 0.05 m s−1, all span a considerable part of the water column, which is not a small vertical scale O(water depth) or O (100–500 m, the maximum range of observations), with either 0 or π phase change. This implies that they actually represent internal waves of low vertical modes 1 or 2. Maximum amplitudes are found in layers of largest stratification, some in the main pycnocline bordering the frictional bottom boundary layer, suggesting a tidal source. These ‘pycnocline-w’ compose a regular train of (solitary) internal waves and linearly decrease to small values near surface and bottom.  相似文献   

10.
In this paper SST imagery and a three-dimensional numerical model of a river plume were employed to detect upwelling induced by tidal straining in the Rhine ROFI (region of fresh water influence). Previous studies have shown that the Rhine ROFI in the North Sea exhibits strong cross-shore density gradients that compete with tidal and wind mixing to establish stratification. During neap periods with low mixing energy an area measuring 30 km offshore by 100 km alongshore becomes stratified. When the ROFI is stratified strong cross-shore currents are observed, with surface currents rotating anti-cyclonically and bottom currents rotating cyclonically. The cross-shore currents interact with the cross-shore density gradients to produce a semi-diurnal cycle of stratification. Due to continuity requirements imposed by the proximity of the coast, the offshore-directed surface currents and onshore-directed bottom currents should lead to coastal upwelling.  相似文献   

11.
The sedimentology of the floor of the Amvrakikos Gulf, a river influenced, semi-enclosed relatively shallow-silled embayment, lying along the northeastern Hellenic coast of the Ionian Sea (eastern Mediterranean Sea), is investigated with respect to its origin (terrigenous and/or biogenic), the prevailing oceanographic conditions and human interference. Nearshore (water depths approximately <10 m) sediments, especially along the northern margin of the Gulf, consist mostly of biogenic sands, as the result of water exchange between the freshwater lagoonal waters and the surface waters of the Gulf. An exception to this is the mouth area of the Arachthos River, which is dominated by the terrigenous riverine sediment influx. The offshore (water depths >10 m) bottom surficial sediments are fine-grained (silty and clayey) of terrigenous origin (>70%); this is attributed to the inter-seasonal, strong two-layer stratification of the water column in the Gulf which restricts benthic productivity by inhibiting the downward flux of surface eutrophic waters and the development of nearbed disoxic conditions in water depths >40 m. River damming has reduced also the riverine terrigenous sediment supply; this is more profound in the case of the Arachthos River where not only the deltaic evolution has been affected, but also the textural character of the seabed sediments of the mouth area has been altered; this is expected to influence the benthic communities of prodeltaic surficial sediment.  相似文献   

12.
The basal depth of the outer layer with internal magnetic sources was calculated from magnetic data available within a roughly 500 km wide and 1200 km long area, running from central Germany to southern Italy. The dataset, deriving from different aeromagnetic surveys, is reduced to the reference altitude of 3000 m a.s.l. and a reference year of 1980.0. The adopted method, which transforms the spatial data into the frequency domain, provides a relationship between the two-dimensional spectrum of the magnetic anomalies and the top and centroid depths of the magnetic sources. The magnetic layer bottom depth (MLBD) thus obtained is 29-33 km deep in the stable areas (central Europe Variscan units, Corsica-Sardinia Variscan block) and corresponds to the Moho, having an average temperature of 560 °C. From the Alps to the Apennines, MLBD ranges between 22 and 28 km and is clearly shallower than the Moho. In these units, the wide variation of MLBD appears to be compatible with the presence of shallow magnetised bodies, consisting of lower crustal rocks (Ivrea-Verbano zone), ophiolitic units (Penninic zone and Voltri Massif) and intrasedimentary basic volcanic bodies (Po Basin). An average value of 25 km can be attributed to MLBD, which corresponds to a temperature of 550 °C. In the peri-Tyrrhenian zone and the Ligurian Sea, MLBD is below the Moho, which ranges from 17 to 20 km depth, and it has a temperature matching approximately to the Curie temperature of magnetite (580 °C).  相似文献   

13.
Bio-physical glider measurements from a unique process-oriented experiment in the Eastern Alboran Sea (AlborEx) allowed us to observe the distribution of the deep chlorophyll maximum (DCM) across an intense density front, with a resolution (~ 400 m) suitable for investigating sub-mesoscale dynamics. This front, at the interface between Atlantic and Mediterranean waters, had a sharp density gradient (Δρ ~ 1 kg/m3 in ~ 10 km) and showed imprints of (sub-)mesoscale phenomena on tracer distributions. Specifically, the chlorophyll-a concentration within the DCM showed a disrupted pattern along isopycnal surfaces, with patches bearing a relationship to the stratification (buoyancy frequency) at depths between 30 and 60 m. In order to estimate the primary production (PP) rate within the chlorophyll patches observed at the sub-surface, we applied the Morel and Andrè (J Geophys Res 96:685–698 1991) bio-optical model using the photosynthetic active radiation (PAR) from Argo profiles collected simultaneously with glider data. The highest production was located concurrently with domed isopycnals on the fresh side of the front, suggestive that (sub-)mesoscale upwelling is carrying phytoplankton patches from less to more illuminated levels, with a contemporaneous delivering of nutrients. Integrated estimations of PP (1.3 g C m?2d?1) along the glider path are two to four times larger than the estimations obtained from satellite-based algorithms, i.e., derived from the 8-day composite fields extracted over the glider trip path. Despite the differences in spatial and temporal sampling between instruments, the differences in PP estimations are mainly due to the inability of the satellite to measure DCM patches responsible for the high production. The deepest (depth > 60 m) chlorophyll patches are almost unproductive and probably transported passively (subducted) from upper productive layers. Finally, the relationship between primary production and oxygen is also investigated. The logarithm of the primary production in the DCM interior (chlorophyll (Chl) > 0.5 mg/m3) shows a linear negative relationship with the apparent oxygen utilization, confirming that high chlorophyll patches are productive. The slope of this relationship is different for Atlantic, mixed interface waters and Mediterranean waters, suggesting the presence of differences in planktonic communities (whether physiological, population, or community level should be object of further investigation) on the different sides of the front. In addition, the ratio of optical backscatter to Chl is high within the intermediate (mixed) waters, which is suggestive of large phytoplankton cells, and lower within the core of the Atlantic and Mediterranean waters. These observations highlight the relevance of fronts in triggering primary production at DCM level and shaping the characteristic patchiness of the pelagic domain. This gains further relevance considering the inadequacy of optical satellite sensors to observe DCM concentrations at such fine scales.  相似文献   

14.
The seasonal ecological response of microzooplankton in the southeastern Arabian Sea is presented. During the spring intermonsoon period, stratification and depletion of nitrate in the surface waters (nitracline was at 60 m depth) cause low integrated chlorophyll a (av. 19±11.3 mg m−2) and primary production (av. 164±91 mgC m−2 d−1). On the other hand, nutrient enrichment associated with coastal upwelling and river influx during the onset and peak summer monsoon resulted in high integrated chlorophyll a (av. 21±6 mg m−2 and av. 29±21 mg m−2, respectively) and primary production (av. 255±94 mgC m−2 d−1 and av. 335±278 mgC m−2 d−1, respectively). During all three periods, diazotropic cyanobacterium Trichodesmium erythraeum dominated in the nutrient depleted surface waters. A general increase in abundance of larger diatoms was evident in the surface waters of the inshore region during monsoon periods. The microzooplankton abundance was found to be significantly higher during the spring intermonsoon (av.241±113×103 ind m−2) as compared to onset of summer monsoon (av. 105±89×103 ind m−2) and peak summer monsoon (av.185±175×103 ind m−2). Microzooplankton community during the spring intermonsoon was numerically dominated by ciliates while heterotrophic dinoflagellate was the dominant ones during the monsoon periods. The high abundance of ciliates during the spring intermonsoon could be attributed to the stratified environmental condition prevailed in the study area which favors high abundance of smaller phytoplankton and cyanobacteria, the most preferred food of ciliates. On the other hand, the dominance of heterotrophic dinoflagellates during the monsoon periods could be linked to their ability to graze larger diatoms which were abundant during the monsoon periods. The overall results show low abundance of microzooplankton in the eastern Arabian Sea during the monsoon periods mainly due to a decline in ciliates abundance. This decline during the monsoon periods could be the result of (a) low abundance of smaller phytoplankton and (b) high stock of mesozooplankton predators (av. 245 ml 100 m−3).  相似文献   

15.
Zhao J  Cao W  Yang Y  Wang G  Zhou W  Sun Z 《Marine pollution bulletin》2008,56(10):1795-1801
A moored optical buoy was deployed in the Pearl River estuarine waters for a 15-day period. A four-day algal bloom event occurred during this study period. Both chlorophyll a concentration and algal cell density (a proxy for biomass) changed dramatically before and after the event. The chlorophyll concentration at a 2.3 m depth rose from 5.15 mg/m−3 at 15:00 h on August 19 to 23.62 mg/m−3 at 9:00 h on August 21, and then decreased to 3.24 mg/m−3 at 15:00 h on August 24. The corresponding cell density ranged from 1.57 × 105 to 1.76 × 106 cells/L. We used normalized fluorescence line height (NFLH) and normalized fluorescence intensity (NFI) in order to determine fluorescence activity. Combined with the in situ sampling dataset, we were able to correlate natural fluorescence (NFLH and NFI) with chlorophyll a concentrations, and found correlation coefficients of 0.72 and 0.75, respectively. We also found correlations between natural fluorescence and cell density, with correlation coefficients of 0.71 and 0.65, respectively. These results indicate that applying continuous time series of natural fluorescence can reflect changes in biomass. This technique will prove extremely useful for in situ and real-time observations using an optical buoy. Although there are still problems to solve in the real-time observation of natural fluorescence in algal bloom events, we discuss the primary factors affecting fluorescence signals and suggest possible methods for mitigating these issues.  相似文献   

16.
The present paper describes the chemical and biological characteristics of an upwelling filament off southern Iberia in October 2004, based on a total of 42 CTD casts, along with ADCP velocity measurements distributed by an almost regular grid of 15 km mean spacing. Stations were sampled from the surface to a maximum depth of 400 dbar, for nutrients (nitrate, phosphate and silicate) and chlorophyll a determination. The effect of cross-shelf exchange of nutrients and chlorophyll a between the coastal and oceanic waters was also investigated. Data revealed that, during the cruise conducted under relaxed winds, a relatively small filament was formed down to 75 m. However, an amount of 180 t of chlorophyll a was estimated there accompanied by low nutrient concentrations. The observed water properties reveal that SW Iberia is similar to NW Iberia, showing the continuity of the Canary Upwelling System along the Iberian Peninsula, one of poorest upwelling systems of the oceanic eastern boundaries. Nevertheless, the amounts of chlorophyll and nutrients transported through the filament are significant, revealing that those play a key role in the seaward export of matter with an important impact on the regional oceanography of this region. Considering the periods of strong upwelling events and the extent of their duration along the year, the amounts of exported matter must be hugely increased and responsible for the high productivity of these waters.  相似文献   

17.
Hypoxic conditions (dissolved oxygen (DO)<2 mg l−1) have been documented in the nearshore coastal waters of Long Bay, South Carolina, United States of America, during summer months over the past several years. Hypoxia was documented in August 2009 in the nearshore (<500 m offshore) for ten consecutive days and four days in September 2009 corresponding with spring tides. This study measured radon activities of shallow beachface groundwater and nearshore bottom waters to estimate mixing rates and submarine groundwater discharge (SGD) in the nearshore waters of central Long Bay. Statistical analyses demonstrate significant correlations between high bottom water radon activities, low DO, and cooler bottom water temperatures during hypoxic conditions. Elevated radon activities during hypoxia were significantly influenced by upwelling favorable conditions which severely limited cross-shelf mixing. Model results indicate mixing of nearshore and offshore waters was limited by up to 93% (range: 43-100%) relative to non-hypoxic conditions. Data suggests previously overlooked natural phenomena including limited cross-shelf mixing and SGD can significantly influence nearshore water quality.  相似文献   

18.
From 15 to 28 August in 2007, a Chaetoceros socialis bloom was detected in the Pearl River Estuary water with chlorophyll a concentration (Chl a) up to 30 mg m−3 and cell density up to 106 cells L−1. Time series of bio-optical measurements was obtained at a single site (114.29°E, 22.06°N) with the mooring of marine optical buoy. Light absorption properties of seawater experienced large variability throughout the algal bloom. Absorption by colored dissolved organic matter (CDOM) was one of the dominant optical components of the light absorption (30–70%) especially for pre- and post-bloom waters, and it tended to decrease with Chl a during the algal bloom. Absorption by phytoplankton was another dominant optical component (18–50%) and increased rapidly with Chl a. Phytoplankton and accompanying material played dominant roles in light absorption as indicated by the relationship between absorption coefficient and Chl a. At high pigment concentrations, water samples showed significantly lower specific phytoplankton absorption, compared with pre- and post-bloom conditions, with the specific phytoplankton concentration at 443 nm varied between 0.011 and 0.022 m2 mg−1 and that at 676 nm between 0.007 and 0.018 m2 mg−1; small values of blue-to-red ratio of phytoplankton were also observed. These lower values were associated with variations in phytoplankton size structure. Spectral variability of phytoplankton absorption and total absorption (not including the fixed background absorption by pure water itself) could be expressed as simple linear functions linking absorption at one wavelength to the absorption at the other wavelengths, with the slope of the relationship changing with wavelength. The absorption coefficients by non-algal particles and CDOM follow the general exponential functions with remarkably limited variability in the exponent with means of 0.0105 and 0.0166 nm−1, respectively. These spectral dependencies of absorption coefficients provide useful information for retrieving inherent optical properties from reflectance data in a remote-sensing context.  相似文献   

19.
We use the hydrographic data obtained during the joint survey of the Yellow Sea by the First Institute of Oceanography, China and the Korea Ocean Research and Development Institute, Korea, to quantify the spatial structures and temporal evolution of the southern Yellow Sea Cold Water Mass (YSCWM). It is indicated that the southern YSCWM is a water mass that develops in summer and decays in fall. In winter, due to the intrusion of the Yellow Sea Warm Current (YSWC), the central area (approximately between 34°N and 35°N, 122°E and 124°E) of the Yellow Sea is mainly occupied by relatively high temperature water (T>10 °C). By contrast, from early summer to fall, under the seasonal thermocline, the central area of Yellow Sea is occupied by cold water (T<10 °C). In summer, the southern YSCWM has two cold cores. One is formed locally southeast of Shandong Peninsula, and the other one has a tongue-like feature occupying the area approximately between 34°N and 37°N, 123°E and 126°E. The bottom layer temperature anomalies from February to July in the cold tongue region, along with the trajectories of the bottom floaters, suggest that the cold water mass in the northeast region has a displacement from the north to the central area of the Yellow Sea during the summer.  相似文献   

20.
We examined the occurrence of seasonal hypoxia (O2<2 mg l−1) in the bottom waters of four river-dominated ocean margins (off the Changjiang, Mississippi, Pearl and Rhône Rivers) and compared the processes leading to the depletion of oxygen. Consumption of oxygen in bottom waters is linked to biological oxygen demand fueled by organic matter from primary production in the nutrient-rich river plume and perhaps terrigenous inputs. Hypoxia occurs when this consumption exceeds replenishment by diffusion, turbulent mixing or lateral advection of oxygenated water. The margins off the Mississippi and Changjiang are affected the most by summer hypoxia, while the margins off the Rhône and the Pearl rivers systems are less affected, although nutrient concentrations in the river water are very similar in the four systems. Spring and summer primary production is high overall for the shelves adjacent to the Mississippi, Changjiang and Pearl (1–10 g C m−2 d−1), and lower off the Rhône River (<1 g C m−2 d−1), which could be one of the reasons of the absence of hypoxia on the Rhône shelf. The residence time of the bottom water is also related to the occurrence of hypoxia, with the Mississippi margin showing a long residence time and frequent occurrences of hypoxia during summer over very large spatial scales, whereas the East China Sea (ECS)/Changjiang displays hypoxia less regularly due to a shorter residence time of the bottom water. Physical stratification plays an important role with both the Changjiang and Mississippi shelf showing strong thermohaline stratification during summer over extended periods of time, whereas summer stratification is less prominent for the Pearl and Rhône partly due to the wind effect on mixing. The shape of the shelf is the last important factor since hypoxia occurs at intermediate depths (between 5 and 50 m) on broad shelves (Gulf of Mexico and ECS). Shallow estuaries with low residence time such as the Pearl River estuary during the summer wet season when mixing and flushing are dominant features, or deeper shelves, such as the Gulf of Lion off the Rhône show little or no hypoxia.  相似文献   

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