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1.
Patagonia exhibits a particularly abundant record of Cretaceous dinosaurs with worldwide relevance. Although paleontological studies are relatively numerous, few include taphonomic information about these faunas. This contribution provides the first detailed sedimentological and taphonomical analyses of a dinosaur bone quarry from northern Neuquén Basin. At Arroyo Seco (Mendoza Province, Argentina), a large parautochthonous/autochthonous accumulation of articulated and disarticulated bones that represent several sauropod individuals has been discovered. The fossil remains, assigned to Mendozasaurus neguyelap González Riga, correspond to a large (18–27-m long) sauropod titanosaur collected in the strata of the Río Neuquén Subgroup (late Turoronian–late Coniacian). A taphonomic viewpoint recognizes a two-fold division into biostratinomic and fossil-diagenetic processes. Biostratinomic processes include (1) subaerial biodegradation of sauropod carcasses on well-drained floodplains, (2) partial or total skeletal disarticulation, (3) reorientation of bones by sporadic overbank flows, and (4) subaerial weathering. Fossil-diagenetic processes include (1) plastic deformation of bones, (2) initial permineralization with hematite, (3) fracturing and brittle deformation due to lithostatic pressure; (4) secondary permineralization with calcite in vascular canals and fractures, and (5) postfossilization bone weathering. This type of bone concentration, also present in Rincón de los Sauces (northern Patagonia), suggests that overbank facies tended to accumulate large titanosaur bones. This taphonomic mode, referred to as “overbank bone assemblages”, outlines the potential of crevasse splay facies as important sources of paleontological data in Cretaceous meandering fluvial systems.  相似文献   

2.
From excavation at FLK North levels 1–2 in 1960–1962, Mary Leakey reported approximately 1200 Oldowan artifacts and 3300 large mammal fossils as a hominin “living floor”. Preliminary taphonomic analysis by Bunn seemed supportive, based on the presence of some cut-marked bones, the concentration of several dozen bovid individuals, and the relative abundance of limbs and mandibles over other axial elements. Recent taphonomic analysis of Leakey's entire fossil assemblage by Domínguez-Rodrigo and Barba, however, documents a minor hominin role at the site, contrasted to the dominant role of carnivores. Felids brought prey animals; hyenas scavenged from abandoned felid meals. At different times, hominins butchered several bovids and discarded artifacts at this dynamic location. Since 2006, renewed excavations at FLK North and other sites by the Olduvai Paleoanthropology and Paleoecology Project (TOPPP) have expanded artifact and fossil samples and implemented new analytical approaches to clarify taphonomic histories of the Olduvai paleolandscape. At FLK North, > 1000 new large mammal fossils from levels 1 to 2 show minimal butchery evidence amid abundant evidence of carnivore gnawing/fracture, rodent gnawing, and sediment abrasion. To help guide future excavation and analyses, we have developed several alternative working hypotheses of site formation.  相似文献   

3.
The taphonomic and archeological implications of massive mortality patterns of guanaco (Lama guanicoe) populations due to winter stress are discussed, considering both local and regional scales. The presentation has its focus in the Lake Cardiel basin, Southern Patagonia, Argentina. Winter stress is not an unusual event in Patagonian ecosystems and it produces a seasonal and predictable output: a huge number of dead guanacos in certain loci.Five faunal assemblages formed in 2000 are discussed, three in rock shelters and two at open-air settings, considering guanaco individuals/bone preservation, age and sex structure, death positions, degree of interment, carnivore damages, trampling marks, and the association between guanaco individuals/bones and archeological material (mainly lithic artifacts). Conditions exist for the commingling of naturally formed guanaco bone assemblages and archeological materials. The taphonomic signature of this mortality pattern is analysed, and criteria developed that allow identification of the potential mixture between archeological material and modern guanaco bones. From an archeological point of view, the potential scavenging opportunities brought by these guanaco mortality events to hunter-gatherer populations are explored.In sum, the results of this study are twofold. First, it brings new data on the regional taphonomy of guanacos and its implications for the archeological record. Second, it invites consideration of these mass mortality events in the context of strategies employed by hunter-gatherer populations to cope with their environment.  相似文献   

4.
The present study analyses the taphonomic features of recent small mammal bone and teeth accumulations recovered from pellets generated by Strix chacoensis in a Natural Protected Area in the central Monte Desert of Mendoza, Argentina. In order to identify the taphonomic signature of this predator in the formation of zooarchaeological and palaeontological accumulations, taxonomic profiles, relative abundances, bone breakage patterns and digestive corrosion were evaluated. Taphonomic results locate S. chacoensis in modification category 2 (intermediate). Although the general taphonomic patterns detected in the present study were similar to a typical owl pattern, some differences from that pattern were detected. This sample showed lesser relative abundances of skeletal elements, greater loss of teeth and a larger number of incisors modified by digestion than those generated by other strigiforms. The comparison with reported archaeological accumulations suggested that the small mammal remains recovered in some sites from the Monte Desert may correspond to the predatory activity of S. chacoensis. The identification of the action of S. chacoensis in archaeological and palaeontological small mammal bone and teeth accumulations is a promising tool to evaluate the conservation status of native forests in the past.  相似文献   

5.
The present study analysed the taphonomic characteristics of small mammal bone accumulations produced by small felids in an area from the central Monte Desert (Mendoza, Argentina). In order to provide criteria to identify the role that these predators had in the formation of zooarchaeological assemblages, the anatomical representation, bone breakage patterns and degrees of digestive corrosion were evaluated. The main taphonomic results are: low average values for the relative abundance of skeletal elements; greater representation of mandibles, maxillae, isolated incisors, humeri and femora than the remaining elements; preponderance of cranial elements with high proportion of isolated teeth; elevated frequencies of proximal limb bones compared with distal parts; high degree of breakage in all skeletal elements and digestive corrosion on almost all diagnostic bones (mainly moderate and heavy). The values of the studied taphonomic variables indicate that small felids in this area made severe alterations to the bones of their prey (mainly rodents), attributable to the category of extreme modifier, while preserving enough skeletal elements to allow their taphonomic characterization. Tooth marks or grooves on bone surfaces produced by scratching and chewing were not detected. The low relative abundance of skeletal elements, the high degree of breakage and the elevated frequency of elements with digestion traces represent general taphonomic patterns that fall within those reported for other South American small carnivores.  相似文献   

6.
《Quaternary Science Reviews》2003,22(5-7):595-607
A taphonomic study was conducted of large mammal remains from the Middle Pleistocene site of Treugol’naya Cave. The site is located at an elevation of 1500 m above sea level near the town of Pregradnaya in the northwestern Caucasus region of Russia, and was excavated by Doronichev (Doronichev, V.B., 2000. Lower paleolithic occupation of the northern Caucasus. ERAUL 92, 67–77.) between 1986 and 2000. Large mammal remains were identified by Baryshnikov (Baryshnikov, G.F., 1993. Krupnye mlekopitayushchie ashelskoi stoyanki v peshchere Treugol’naya na Severnom Kavkaze. Trudy Zoologicheskogo instituta RAN 249, 3–47), and reflect predominance of red deer (Cervus elaphus), bison (Bison schoetensacki), and cave bear (Spelaearctos deningeri). Less common taxa include goat (Capra sp.), wolf (Canis mosbachensis), rhinoceros (Stephanorhinus hundsheimensis) and horse (Equus altidens). Data were collected on weathering, breakage, surficial damage, skeletal-part frequencies, and age and season of death from the these remains, which are stored at the Zoological Institute, Russian Academy of Sciences in St. Petersburg. Analysis of the data revealed little evidence for accumulation of the large mammal remains by the hominid occupants of the cave. The carnivore remains probably represent natural mortality, while some of the ungulate remains were apparently accumulated by stream action. Most of the remaining ungulate remains were probably collected by carnivores.  相似文献   

7.
The avian family of barn owls (Tytonidae, Strigiformes) is the most geographically widespread group of owls in the world and one of the most common accumulators of small mammal bone and tooth remains recovered from archaeological and palaeontological sites. The present study analyses the taphonomic characteristics of modern barn owl pellet-derived small mammal bone and tooth accumulations at three roosting sites from the central Monte Desert, one of the most arid regions of South America. In order to identify the taphonomic signature of this predator in the formation of zooarchaeological and palaeontological accumulations, taxonomic structure, relative abundance of skeletal elements, bone and teeth breakage patterns, and digestion were evaluated. Taphonomic results locate the barn owl in modification category 1 (light modifier). Although the general taphonomic results obtained in the present study were similar to a common barn owl pattern, the three samples showed variability both among them and with other samples previously reported in different regions of the world. Very light digestion was detected on incisors, whereas the proportion of digestion on incisors was close to 20%. Cricetid and murid rodents as well as marsupials were detected. Salinomys delicatus, a scarcely known and endangered cricetid species, was recovered.  相似文献   

8.
Four southeastern Wisconsin mammoth localities located within a glacial landscape had well-preserved remains found in inter-morainal depressions filled with lacustrine clays covered with peat. Numerous radiocarbon ages dated the mammoths to the late Pleistocene. The taphonomic analysis focused on determining the agency or agencies involved in site formation and the agency or agencies involved in modification to the bones. Statistical approaches to bone orientation data underscored that water transport was not a factor in the formation of the bone beds nor was water movement a disturbance factor. The bone modification profile of the overall assemblage was dominated by chemical weathering, followed by other chemical and microbiological processes. Root etching was the most common biological modification, and the most frequent modifications are involved with the microenvironments of the surface to burial substrate. Potentially cultural modification accounted for only a small portion of the cortical damage to the bones. This general profile indicated an assemblage far more influenced by the immediate environment than by passing animals or people.  相似文献   

9.
Investigations at a Late Weichselian freshwater basin in northwestern Jutland, Denmark, yielded a fairly rich assemblage of vertebrate remains, mostly bones and teeth of small mammals. The remains are primarily allochthonous and the bones have been subjected to different taphonomic pathways and agents. AMS 14C-dates on terrestrial organic remains provided ages of Middle to Late Allerød time. Identifications revealed the first fossil record in Scandinavia of Rana arvalis, Sorex minutus, Ochotona cf. pusilla, Microtus gregalis, Microtus oeconomus , and Sicista cf. betulinu. Spermophilus cf. major and Desmana moschata , previously found only once and twice respectively, were retrieved, and Sorex araneus and Arvicola terrestris were recovered for the first time beyond the Atlantic chronozone. Ecologically, the Nørre Lyngby small mammal fauna can be characterized by its very high and almost equal proportions of boreal forest and steppe elements followed by a relatively high proportion of tundra elements. The fossil species share a modern area of sympatry north of the Caspian Sea from the river Volga in the west to the southern and western slopes of the Urals. If, however, the large Allerød mammals are added, the fauna is without modern analogues. The Nørre Lyngby fauna can be seen as a last expansion of the North European glacial fauna. Provided that an absolute chronology and a differentiated sea-level curve for the area can be established, the Nsrre Lyngby fauna could become important for studies in mammalian dispersal and migration rates.  相似文献   

10.
Sediments from the Upper Cretaceous (Cenomanian-Turonian) Tropic Shale were deposited along the western margin of the Western Interior Seaway, in present-day southern Utah. Marine vertebrates from this formation include plesiosaurs, mosasaurs, bony fish, sharks, and turtles. They are concentrated in the lower portion of the Tropic Shale, mostly between Bentonites B and D. Study of the taphonomic condition of these vertebrates has contributed to an understanding of how they were preserved as well as a detailed paleoenvironment for the Tropic Shale. Physical factors played the dominant role in their preservation, with robust and durable skeletal elements, such as teeth and vertebrae, being most common within the shale. Isolated bones and teeth are also relatively common within the formation, while complete and nearly complete skeletons are more rare. Biological factors played a less dominant role, with no evidence of epifaunal or infaunal activity preserved with any of the skeletal remains. In addition, scavenging marks (both bite and gnaw marks) are relatively uncommon, typically only being found on more complete specimens. A signature of post-burial alteration can be recognized as low levels of abrasion, weathering, and compression and high levels of fracturing of vertebrate skeletal material. Slightly higher levels of abrasion and weathering occur to the west, closer to the ancient shoreline, suggesting some pre-burial alteration. The preservation of marine vertebrates in the Tropic Shale suggests a low energy marine environment with some weak bottom currents and low levels of benthic oxygen. The substrate ranged from soft and soupy to firm, with moderate sedimentation rates resulting in relatively rapid burial.  相似文献   

11.
Fossil bones and teeth are potentially important repository for geochemical proxy data and a target for radiometric dating. The concentration of many trace elements in bones and teeth increases by orders of magnitude after death and it is this diagenetic incorporation that forms the basis for several areas of geochemical study. The use of bones and teeth in this context relies on two assumptions: first, that target metal ions are incorporated rapidly after death, reflecting a known environmental signal, and second, that after early incorporation, the bone or tooth remains as an essentially closed system, resistant to later diagenetic change. A wide literature has developed exploring these assumptions, but relatively little direct evidence has been used to assess the long-term diagenetic stability of trace elements within bones and teeth. In this study, we use the Lu-Hf isotope system to show that bones and teeth of Cretaceous and Triassic age from both terrestrial and marine settings experience continued, long-term diagenetic change, most likely through gradual addition of trace elements. Modelling suggests that diagenetic addition after initial recrystallisation may account for >50% of the total REE content in the sampled bones, the extent depending on initial uptake conditions. Tooth enamel and enameloid may be more resistant to late diagenetic changes, but dentine is probably altered to the same extent as bone. These results have significant implications for the use of bones and teeth as hosts of chronological, palaeoceanographic, palaeoenvironmental and taphonomic information, particularly in Mesozoic and Palaeozoic contexts.  相似文献   

12.
Modes of faunal exploitation in NW Chubut Province (Patagonia, Argentina) during the last 3500 calibrated years BP are discussed, based on taphonomic and zooarchaeological analyses. Bone assemblages were recovered from archaeological sites of the Sub-Antarctic forests and the extra-Andean Patagonian steppe, where huemul (Hippocamelus bisulcus) and guanaco (Lama guanicoe) were, respectively, the main staple. In the steppe, lesser rhea (Pterocnemia pennata), a medium-sized flightless bird, also was integrated into the hunter-gatherer diet. Rodents, carnivores, flying birds, and fishes also were recovered. Different processes and agents were involved in the accumulation of small vertebrates bone assemblages (e.g. birds of prey, foxes, and small carnivores). In the steppe, only two taxa of small vertebrates—birds and mountain viscacha (Lagidium viscacia)—can be related with human subsistence activities. These two taxa and all large vertebrates reveal fat-oriented carcass processing.Guanaco bone assemblages associated both with ceramic and non-ceramic technologies do not show changes in carcass processing. Moreover, the similarities in bone grease exploitation suggest boiling prior to the appearance of pottery.  相似文献   

13.
Understanding resource utilization and economic diversification amongst Holocene hunter‐gatherers in southern Brazil requires in‐depth taphonomic analysis of faunal assemblages. Three Early to Late Holocene archaeological sites (Garivaldino, Pilger and Sangão) in Rio Grande do Sul State, southern Brazil, revealed large assemblages of small mammal (?1 kg) remains, composed mainly of rodents. To appreciate depositional processes of fauna in relation to human consumption, taphonomic attributes and processes were evaluated. The results indicated that the remains accumulated through different antemortem taphonomic pathways, apparently linked to the size and natural history of the species represented. The small‐ and medium‐sized (<150 g) cricetid rodents (e.g. Pseudoryzomys simplex and Sooretamys angouya) were represented by a low proportion of teeth with lightly digestive corrosion suggesting that they were derived from the predatory activity of owls, possibly Tyto alba. In contrast, large‐sized cricetids (>150 g; Kunsia tomentosus, Gyldenstolpia sp. and Holochilus sp.), and several caviomorph rodents with aggregated spatial distributions (Echimyidae (?Dicolpomys fossor, Phyllomys sp., ?Clyomys riograndensis and ?Euryzygomatomys mordax), Caviidae (Cavia sp.) and Ctenomyidae (Ctenomys sp.)) showed evidence of thermoalteration patterns and cut marks on bones, suggesting human exploitation. A postmortem depositional environment was deduced from dendritic and branched patterns of rootlet etching, apparent mostly at Garivaldino and Pilger. Polished areas and holes on bone surfaces, and impregnation of manganese showed moderate (Sangão) or low (Garivaldino and Pilger) incidence, pointing to water transport under moderate (Pilger) and low (Garivaldino and Sangão) energy over short distances. The results obtained here provide the first clear evidence of early human exploitation of small mammals in southern Brazil, suggesting a diversification of economies. In addition, as several of the recorded rodents are today regionally or biologically extinct, a preliminary discussion about the potential impact of humans on this process is provided.  相似文献   

14.
Comparisons of historical and modern assemblages of mammals can yield important insights into patterns and processes of environmental change. Here, we present the first analyses of small mammal assemblages present in northern Patagonia during the last glacial maximum (LGM). Using remains obtained from owl pellets excavated from an archeological cave site (Arroyo Corral I, levels VII–V, carbon dates of 22,400–21,530 cal yr BP), we generate estimates of the minimum number of individuals for all species detected; these estimates, in turn are used to determine relative species abundances. Comparisons of these data with similar analyses of small mammal remains obtained from a second archeological site (ACoII, levels IV–V, carbon dates of 10,010–9220 cal yr BP) as well as from modern owl pellets reveal pronounced changes in relative species abundance since the LGM. In particular, Euneomys chinchilloides and Ctenomys sociabilis – the predominant species during the LGM – declined markedly, suggesting a change from open, bare habitat punctuated by patches of wet meadows and shrubs to the more densely vegetated mosaic of ecotone habitats found in this region today. These data provide important new insights into the environmental changes that have occurred in northern Patagonia over the last 20,000 years.  相似文献   

15.
Leporids (Lagomorpha, Mammalia), especially the mountain hare (Lepus timidus) and the European rabbit (Oryctolagus cuniculus), have been among the most abundant small game in western Europe since the Pleistocene. These animals are preferred prey for more than 40 predators, which increases the chances of finding their skeletal remains at archaeological or natural sites, and in particular karstic formations alternately occupied by Palaeolithic human societies and other predators. Moreover, specific eco-ethological characteristics for both species also offer the possibility to produce a substantial quantity of their bones accumulated by natural mortality without predation, making it more difficult to identify their origin in the fossil record. Despite this fact, the taphonomic signature of accidental-natural accumulations of leporids in karsts, such as pitfalls, has never been properly characterised. In order to address these issues, we carried out a detailed taphonomic study of leporid remains in a pitfall without any evidence of human activity, namely at Coulet des Roches (Monieux, Vaucluse, south-eastern France). At this site, leporids are the most abundant species throughout the Last Glacial Maximum sedimentary sequence. The bone accumulation was analysed in order to determine the most relevant criteria to identify a natural accumulation that results from the accidental fall of individuals in the pitfall. The relative proportions of each species are consistent with their current ethology, and their mortality profiles are compatible with the structure of a living natural population. The completeness rate of the different skeletal portions for both species is relatively high, bones are less fragmented and, unlike what might be expected, very few anatomical connections were preserved. The observed anatomical representation, breakage, and bone surface modification patterns are discussed and compared with available data concerning accumulations with predation. Taken together, the identified criteria contribute valuable help in the recognition of accidental mortality in the fossil record, as well as to evaluate the proportion of leporid bones accumulated naturally and those accumulated by predation, human or not, in a deposit.  相似文献   

16.
Geomorphological evidence and historical wind records indicate that eolian processes have heavily influenced San Miguel Island environments for much of the Late Quaternary. The island is almost constantly bombarded by prevailing northwesterly winds, with peak velocities exceeding 75 km/h and wind gusts reaching over 100 km/h. These strong winds played an important role in the location, formation, and preservation of the island's more than 600 archaeological sites. Excavation and surface collection at a stratified Middle and Late Holocene archaeological site on the island's north coast suggest that wind related disturbances result in significant displacement of light fish bones, produce concentrations of shellfish and heavy mammal bones, and cause significant abrasion, etching, and polishing of bones, shells, and artifacts. These data illustrate that wind not only alters surface materials but can significantly disturb subsurface deposits to a depth of at least 20 cm. Working in concert with a variety of taphonomic processes, wind can play a fundamental role in the preservation of archaeological sites and careful scrutiny during excavation and laboratory analysis is required to delineate its effects. © 2002 Wiley Periodicals, Inc.  相似文献   

17.
The preservation process of faunal remains from the Middle Paleolithic levels of Kůlna Cave was studied within a taphonomic and palaeodietary framework. A visual examination of bones and teeth showed that they display a wide variety of colors. Chemical analyses (C, N, Fe, Mn) and a combination of analytical techniques, such as Fourier transform infrared (FT‐IR), X‐ray diffraction (XRD), and electron spin resonance (ESR) spectrometry, were conducted on 13 fossil remains. The crystallinity indexes, IR‐SF, XRD‐CI, XRD‐L200, the IR area band (Amide I), and the ESR analyses showed that the bones were not burned at a temperature higher than 200°C. This implies that the bone remains were not used as fuel by Neanderthals at Kůlna Cave. Coloring in these bones is largely due to the presence of Mn and Fe. © 2006 Wiley Periodicals, Inc.  相似文献   

18.
The world’s first archaeological traces from 2.6 million years ago (Ma) at Gona, in Ethiopia, include sharp-edged cutting tools and cut-marked animal bones, which indicate consumption of skeletal muscle by early hominin butchers. From that point, evidence of hominin meat-eating becomes increasingly more common throughout the Pleistocene archaeological record. Thus, the substantive debate about hominin meat-eating now centers on mode(s) of carcass resource acquisition. Two prominent hypotheses suggest, alternatively, (1) that early Homo hunted ungulate prey by running them to physiological failure and then dispatching them, or (2) that early Homo was relegated to passively scavenging carcass residues abandoned by carnivore predators. Various paleontologically testable predictions can be formulated for both hypotheses. Here we test four predictions concerning age-frequency distributions for bovids that contributed carcass remains to the 1.8 Ma. old FLK 22 Zinjanthropus (FLK Zinj, Olduvai Gorge, Tanzania) fauna, which zooarchaeological and taphonomic data indicate was formed predominantly by early Homo. In all but one case, the bovid mortality data from FLK Zinj violate test predictions of the endurance running-hunting and passive scavenging hypotheses. When combined with other taphonomic data, these results falsify both hypotheses, and lead to the hypothesis that early Homo operated successfully as an ambush predator.  相似文献   

19.
The Lapa do Santo archaeological site, located in the Lagoa Santa region, Brazil, represents an important hunter-gatherer occupation dated from the Early and Middle Holocene. Prior studies of archaeofauna dating to this period are few and most of them only provide basic faunal identification with limited information on taphonomic processes. The main goal of this study is to identify the archaeofauna, record the taphonomic processes, and make inferences about its natural (interpreted as the death of the animal in the rockshelter area due to natural causes or due to predation) or anthropic origins. No extinct species (including megamastofauna) were identified. Taxonomic analysis indicated that most faunal remains consist of taxa that may be of mixed origin (natural or anthropic), such as microvertebrates and carnivorous mammals. The Cervidae are the main vertebrate family found in the studied material and these are likely the result of hunting. The Cervidae Ozotoceros support the presence of a fauna that is typical of savannah environments, although the assemblage contains mostly generalist fauna that can thrive in a variety of environments. Very few remains showed clear evidence for taphonomic processes related to human interaction, such as burnt bones.  相似文献   

20.
Archaeological sites composed only of surficial lithics are widespread in arid environments. Numerical dating of such sites is challenging, however, and even establishing a relative chronology can be daunting. One potentially helpful method for assigning relative chronologies is to use lithic weathering, on the assumption that the most weathered artifacts are also the oldest. Yet, few studies have systematically assessed how local environmental processes affect weathering of surficial lithics. Using macroscopic analyses, we compared the weathering of surficial lithic assemblages from seven mid‐to‐late Holocene archaeological sites sampled from four different microenvironments in the Atacama Desert of northern Chile. Changes in polish, texture, shine, and color were used to establish significant differences in weathering between two kinds of locations: interfluves and canyon sites. Lithics from interfluve sites were moderately to highly weathered by wind and possessed a dark coating, whereas canyon lithics were mildly weathered despite greater exposure to moisture, often lacked indications of eolian abrasion, and lacked dark coatings. Our results show that lithic weathering can be used as a proxy for relative age, but only after considering local environmental factors. The power of such chronologies can be improved by combining archaeological, paleoenvironmental, geomorphological, and taphonomic data.  相似文献   

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