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1.
Discharge of saline groundwater from Eucalyptus forests on a semi-arid floodplain was directly determined by first measuring transpiration rates from the forests, and then partitioning the transpiration flux into groundwater discharge and soil water depletion. This partitioning was achieved by identifying the source of the transpired water with naturally occurring stable isotopes of water. Transpiration rates were low, being about 0.3 mm day−1 from three E. largiflorens sites and up to 2 mm day−1 from an E. camaldulensis site. There was no significant variation in transpiration across seasons, indicating that transpiration was limited by environmental factors other than evaporative demand. Despite its salinity (electrical conductivities of 11–33 dS m−1), the groundwater was used by the forests at all sites and all times, and made up 100% of transpiration in more than half of the measurements, and 40–80% in the remainder. There was some consistency in water uptake patterns. E. camaldulensis tended to take up shallow soil water and groundwater simultaneously, as did trees at one of the E. largiflorens sites. At the driest sampling time, however, groundwater was the only source of water for trees at both of these sites. Trees at the remaining two E. largiflorens sites generally relied solely on the groundwater. The tree water source results indicate that groundwater discharge fluxes were between 40 and 100% of the transpiration fluxes at these sites. These groundwater discharge fluxes were small in terms of regional groundwater balances, but would be important in the salinisation of the soils. Additionally, uptake of water from the soil profile by the trees substantially increased groundwater discharge compared with discharge from the soils had they been bare of vegetation.  相似文献   

2.
Soil water matric potentials (Ψm) and the deuterium (δ2H) composition at natural abundance levels of xylem water, soil water, river water and groundwater were used to evaluate whether trees use groundwater during the dry season in the riparian zone of the Daly River (Northern Territory, Australia). Groundwater was a significant source of water for plant transpiration, probably accounting for more than 50% of the water transpired during the dry season. Groundwater use occurred either when trees used water from the capillary fringe or when low Ψm induced by soil water uptake lifted groundwater in the vadose zone. Several water use strategies were inferred within the riparian plant community. Melaleuca argentea W. Fitzg and Barringtonia acutangula (L.) Gaertn. appeared to be obligate phreatophytes as they used groundwater almost exclusively and were associated with riverbanks and lower terraces with shallow (<5 m) water tables. Several species appeared to be facultative phreatophytes (including Cathorium umbellatum (Vahl.) Kosterm. and Acacia auriculiformis A. Cunn. ex Benth.) and tended to rely more heavily on soil water with increased elevation in the riparian zone. The levee-bound Corymbia bella K.D. Hill and L.A.S. Johnson mostly used soil water and is either a facultative phreatophyte or a non-phreatophyte. The temporal variability in groundwater utilisation by the trees is unclear because the study focused on the end of the dry season only. A decline in the regional water table as a result of groundwater pumping may affect the health of riparian zone vegetation in the Daly River because groundwater use is significant during the dry season.  相似文献   

3.
Hydrological impact of Eucalyptus plantation in India   总被引:1,自引:0,他引:1  
A brief review is given of the results of hydrological studies carried out in southern India on the effects of plantations of Eucalyptus and other fast growing exotic tree species on water resources, erosion and soil nutrients at sites of differing rainfall and soil depth in Karnataka. New results on the impacts of the plantations on raindrop-induced erosion and preliminary results from studies that are aimed at improving the water use efficiency of the plantations are also presented.

The erosion studies indicate that soil detachment resulting from net rainfall beneath Eucalyptus camaldulensis will be greater than beneath Pinus caribaea but less than that beneath the indigenous species, Tectona grandis.

The water use efficiency studies, which make use of a controlled environment facility, have confirmed that in the dry zone of southern India water availability is the principal limiting factor on growth although, when water is available, nutrient limitations become important. Removal of both water and nutrient stress results typically in a fivefold increase in volume growth for the first year of growth.  相似文献   


4.
A simple field‐based method for directly parameterizing root water uptake models is proposed. Stem psychrometers and sap flow meters are used to measure stem water potential and plant transpiration rate continuously and simultaneously. Predawn stem water potential is selected as a surrogate for root zone soil water potential to examine and parameterize the root water uptake–water stress response functions. The method is applied to two drooping sheoak (Allocasuarina verticillata) trees for a period of 80 days, covering both a dry season and a wet season. The results indicate that the S‐shape function is more appropriate than the Feddes piecewise linear function for drooping sheoak to represent the effect of soil moisture stress on its root water uptake performance. Besides, the water stress function is found to be not only a function of soil moisture but also dependent of the atmospheric demand. As a result, the water stress function is corrected for the effect of atmospheric conditions. Copyright © 2012 John Wiley & Sons, Ltd.  相似文献   

5.
Subalpine forests are hydrologically important to the function and health of mountain basins. Identifying the specific water sources and the proportions used by subalpine forests is necessary to understand potential impacts to these forests under a changing climate. The recent “Two Water Worlds” hypothesis suggests that trees can favour tightly bound soil water instead of readily available free-flowing soil water. Little is known about the specific sources of water used by subalpine trees Abies lasiocarpa (Subalpine fir) and Picea engelmannii (Engelmann spruce) in the Canadian Rocky Mountains. In this study, stable water isotope (δ18O and δ2H) samples were obtained from S. fir and Engelmann spruce trees at three points of the growing season in combination with water sources available at time of sampling (snow, vadose zone water, saturated zone water, precipitation). Using the Bayesian Mixing Model, MixSIAR, relative source water proportions were calculated. In the drought summer examined, there was a net loss of water via evapotranspiration from the system. Results highlighted the importance of tightly vadose zone, or bound soil water, to subalpine forests, providing insights of future health under sustained years of drought and net loss in summer growing seasons. This work builds upon concepts from the “Two Water Worlds” hypothesis, showing that subalpine trees can draw from different water sources depending on season and availability. In our case, water use was largely driven by a tension gradient within the soil allowing trees to utilize vadose zone water and saturated zone water at differing points of the growing season.  相似文献   

6.
Annual evaporation from groundflora, litter and soil of the jarrah forest was estimated from measurements of daily evaporation by ventilated chambers on several days over two separate 12-month periods. In the first year, when sampling ranged over 0.1 ha of forest, annual evaporation during daylight hours was estimated as 410 mm (0.32 rainfall). In the second year, sampling was more frequent, on a larger scale, and included the night hours. Annual evaporation was estimated at 360 mm (0.36 rainfall).

Similarly, in the second year, annual evaporation from two trees of the dominant middle storey species, Banksia grandis, was estimated at 7500 and 18,9001 respectively. The leaf area of these two trees was 9.6 and 22.4 m2, respectively, so that annual evaporation, when expressed as mm3 per mm2 leaf area, was similar for both trees (mean = 820 ± 30 mm). Applying that value to all Banksia trees in a hectare of forest, and using a measured estimate of leaf area index of 0.19, the estimated annual evaporation from the Banksia component was 155 mm (0.16 rainfall). For the upland part of the forest sampled, the combined annual evaporation from the lower and middle storeys accounted for about half (0.51) of the annual rainfall.

We conclude that reduced evaporation from the upper storey following clearing or thinning may be strongly counteracted by increased evaporation from the understorey due to increased availability of energy and water.  相似文献   


7.
Evaporation and infiltration were compared for tropical rainforest and pasture, near to Manaus, Brazil from October 1990 to February 1992 using measurements of soil water storage over a depth of 2 m. The soil is a clayey oxisol of low water available capacity. In both of the dry seasons studied, the maximum change in soil water storage in the forest was 154 mm and in the pasture it was 131 and 112 mm. Similar behaviour of the soil water reservoir below forest and pasture in the wet season implied that differences in evaporation and drainage were small. In the dry season, soil water storage behaviour in the upper metre of the soil was similar but there were marked differences in the second metre. The pasture took up little water from below 1.5 m but the forest appeared to utilise all of the available water in the 2 m profile in both seasons.

The water balance of the 2 m profile showed that the pasture evaporation rate was equal to that of the forest until storage had decreased 80 mm from the maximum. There was then a decline in pasture evaporation rate to 1.2 mm day−1 as the storage decreased by a further 50 mm. In contrast, the forest uptake rate remained above 3.5 mm day−1 until storage had decreased 140 mm from the maximum (within 15 mm of the extraction limit), before declining abruptly to less than 1.5 mm day−1. There was strong evidence that the forest was able to abstract water from depths greater than 3.6 m.

Spatial variability of soil water storage was significantly greater beneath the pasture than beneath the forest, particularly following rainfall events in the dry season. This was largely the result of redistribution of rainfall as local surface runoff. There was no evidence of redistribution or runoff in the forest.  相似文献   


8.
《水文科学杂志》2013,58(4):739-753
Abstract

The hydrodynamic behaviour of a sloped phreatic aquifer in the Tigray Highlands in northern Ethiopia is described. The aquifer is situated in the soils of a plateau on top of a basalt sequence and lies on steep slopes; the latter lead to hydraulic gradients that can cause high discharge fluxes. Distinct wet and dry seasons characterize the climate of the Tigray Highlands and recharge is absent during the dry season. Because of the fertile vertisols that have developed, the plateau is heavily cultivated and thus has great local economic, and hence social, importance. Water for land irrigation is almost exclusively delivered by rainfall, which is largely restricted to the period June—September. During the dry season, the water table drops dramatically and the aquifer drains nearly completely, under the strong gravity-driven, sustained discharges. This study strives to give insights into recharge and discharge mechanisms of the aquifer, in order to improve the effectiveness of the implemented water conservation measures.  相似文献   

9.
Studying the processes responsible for the distribution of water resources in a river basin over space and time is of great importance for spatial planning. In this study a multi-agent simulation approach is applied for exploring the influence of alternative reservoir operation strategies on water use distribution in the semi-arid Jaguaribe basin in case of decreasing rainfall. Water use distribution is analyzed both for one specific subbasin – our study area – and for the river basin level. Agents in this study are farmers that adapt to local variations in water availability. In this way both natural and human influences on water availability are taken into account. This study shows that a decrease in rainfall and runoff in the Jaguaribe basin leads to a transition of water use from the dry season to the wet season. The dry season water use decreases because of reduced water availability in the dry season. This mainly is the result of reduced rainfall and runoff in the wet season and the consequent increased water use for irrigation in the wet season. A decrease in rainfall and runoff also leads to a relative transition of water use from downstream to upstream at the basin scale. Strategic reservoir operation enables local water managers to offset the effect of decreasing rainfall and runoff with regards to water use at the subbasin level, at the cost of further decreasing water availability at the basin level.  相似文献   

10.
Despite the widely held assumption that trees negatively affect the local water budget in densely planted tree plantations, we still lack a clear understanding of the underlying processes by which canopy cover influences local soil water dynamics in more open, humid tropical ecosystems. In this study, we propose a new conceptual model that uses a combination of stable isotope and soil moisture measurements throughout the soil profile to assess potential mechanisms by which evaporation (of surface soil water and of canopy‐intercepted rainfall) affects the relationship between surface soil water isotopic enrichment (lc‐excess) and soil water content. Our conceptual model was derived from soil water data collected under deciduous and evergreen plants in a shade grown coffee agroforestry system in Costa Rica. Reduced soil moisture under shade trees during the “drier” season, coinciding when these trees were defoliated, was largely the result of increase soil water evaporation as indicated by the positive relationship between soil water content and lc‐excess of surface soil water. In contrast, the evergreen coffee shrubs had a higher leaf area index during the “drier” season, leading to enhanced rainfall interception and a negative relationship between lc‐excess and soil water content. During the wet season, there was no clear relationship between soil water content and between lc‐excess of surface soil water. Greater surface soil water under coffee during the dry season may, in part, explain greater preferential flow under coffee compared with under trees in conditions of low rainfall intensities. However, with increasing rainfall intensities during the wet season, there was no obvious difference in preferential flow between the two canopy covers. Results from this study indicate that our new conceptual model can be used to help disentangling the relative influence of canopy cover on local soil water isotopic composition and dynamics, yet also stresses the need for additional measurements to better resolve the underlying processes by which canopy structure influences local water dynamics.  相似文献   

11.
Although urban wet weather discharges may have elevated concentrations of fecal indicator bacteria impacting water quality at swimming beaches, not all of these bacteria may arise from human sources. In this study, the contribution of non-human fecal indicator bacteria was quantified by sampling coastal reference beaches in southern California. Samples were collected at beaches near stormwater discharges from undeveloped watersheds and analyzed for total coliform, Escherichia coli, and enterococci. Surfzone samples exceeded water quality thresholds >10 times more frequently during wet weather than dry weather. Exceedences were greatest <24 h following rainfall, then steadily declined on successive days. Early season storms exceeded thresholds more frequently, and by greater magnitude, compared to late season storms. Large storms exceeded thresholds more frequently than smaller-sized storms, partly due to the breaching of sand berms. When discharges did reach the surf zone, bacterial concentrations in the wave wash were correlated with watershed bacterial flux.  相似文献   

12.
In-depth studies of water and sediment fluxes from rivers into the sea are very important for understanding the interactions between land and sea. This paper is concerned with identifying the changes in the time series of water and sediment fluxes from Feiyun River in Zhejiang Province, China. Inter- and intraannual variability in the water discharge and sediment load of the Feiyun River into the sea are analyzed using the observed data of runoff (1956–2008) and sediment (1957–2008) at Xuekou Station, which is in the main channel. The results show that there is a good peak–valley correlation between the water discharge and sediment load, and there are obvious seasonal variations, with a 65.7% water discharge and 89.2% sediment load during the flooding periods. Water discharge is mainly controlled by natural rainfall, but the construction of the upstream reservoirs in 1997 increased the discharge amount in the dry season and decreased the amount in the flooding season. Sediment loads were reduced after a huge flood in 1990 and construction of upstream reservoirs, while the latter also decreased the sediment load during the typhoon flooding period. Furthermore, the correlation between water discharge and sediment load is also affected by the flood and reservoir construction. There are some differences in the regression equations of sediment load and water discharge for 1957–1989, 1991–1996, and 1997–2008.  相似文献   

13.
Water table response to rainfall was investigated at six sites in the Upper, Middle and Lower Chalk of southern England. Daily time series of rainfall and borehole water level were cross-correlated to investigate seasonal variations in groundwater-level response times, based on periods of 3-month duration. The time lags (in days) yielding significant correlations were compared with the average unsaturated zone thickness during each 3-month period. In general, for cases when the unsaturated zone was greater than 18 m thick, the time lag for a significant water-level response increased rapidly once the depth to the water table exceeded a critical value, which varied from site to site. For shallower water tables, a linear relationship between the depth to the water table and the water-level response time was evident. The observed variations in response time can only be partially accounted for using a diffusive model for propagation through the unsaturated matrix, suggesting that some fissure flow was occurring. The majority of rapid responses were observed during the winter/spring recharge period, when the unsaturated zone is thinnest and the unsaturated zone moisture content is highest, and were more likely to occur when the rainfall intensity exceeded 5 mm/day. At some sites, a very rapid response within 24 h of rainfall was observed in addition to the longer term responses even when the unsaturated zone was up to 64 m thick. This response was generally associated with the autumn period. The results of the cross-correlation analysis provide statistical support for the presence of fissure flow and for the contribution of multiple pathways through the unsaturated zone to groundwater recharge.  相似文献   

14.
Recent research has shown that planting deep-rooted trees, such as poplar, can take up and degrade important ground water pollutants such as trichloroethylene (TCE) as they transpire water from the capillary fringe of shallow contaminated aquifers. The effect of hydrogeologic factors on the minimum plantation area needed to prevent downgradient migration of contaminated ground water is not well known. Accordingly, the objective of this research was to identify the hydrogeologic parameters that control phytoremediation effectiveness. We used a numerical ground water flow model to evaluate the effect that natural variations in hydrogeologic parameters and growing season duration have on the minimum plantation area required for capture. We found that the plantation area that was needed to completely capture a ground water contamination plume was directly proportional to aquifer horizontal hydraulic conductivity, saturated thickness, and ground water gradient. The plantation area needed for capture increased nonlinearly with increasing plume width, aquifer anisotropy, and decreasing growing season duration. The plantation area needed for capture was generally insensitive to aquifer-specific yield and storativity. Steady-state simulations can be used to predict the plantation area needed for capture in many applications. A particularly important finding of this work is that evapotranspiration fluxes through plantations appropriately sized to contain the plume substantially exceeded the ground water flux through the plume itself.  相似文献   

15.
D.F. Scott 《Journal of Hydrology》1993,150(2-4):409-432
Streamflow and its storm-flow elements in four catchments were analyzed by the paired catchment method for a response to fire. Prior to burning two of the catchments were vegetated with over-mature fynbos (the indigenous scrub vegetation of the southwestern Cape, South Africa), one was afforested with Pinus radiata and the fourth with Eucalyptus fastigata. One of the fynbos catchments was burned in a prescribed fire in the late dry season. The other catchments burned in wildfires.

Neither of the fynbos catchments showed a change in storm-flow. Annual total flow increases of around 16% were in agreement with model predictions, being related to the reductions in transpiration and interception. The manner of streamflow generation appeared to have remained unaltered despite the presence of some water repellency in the soils and consequent overland flow on some steep midslope sites.

The two timber plantation catchments experienced large and significant increases in storm-flows and soil losses, while total flow increased by 12% in the pine catchment and decreased marginally in the eucalypt catchment. The pattern of the storm-flow increases was similar in both cases. After fire, storm hydrographs were higher and steeper though their duration was little changed. The respective first year increases in the pine and eucalypt catchments were 290% and 1110% for peak discharge, 201% and 92% for quick-flow volume, and 242% and 319% for storm response ratio. These fire effects are considered to be due to changes in storm-flow generation consistent with an increased delivery of overland flow (surface runoff) to the stream channel. This was caused, in part, by reduced infiltration resulting from water repellency in the soils of the burned catchments. Overall the hydrological effects of fire are related to numerous interactive factors, including the degree of soil heating, the vegetation type and soil properties.  相似文献   


16.
In sparsely cropped farming systems in semi-arid tropics, rainfall partitioning can be complex due to various interactions between vertical and horizontal water flows, both in the atmosphere and in the soil. Despite this, quantifying the seasonal rainfall partitioning is essential, in order to identify options for increased yields. Results are presented on water flow components, based on field measurements and water balance modelling, for three years (1994–96) in a farmer's field cultivated with pearl millet [Pennisetum glaucum (L.) Br.] in the Sahel (Niger). Water balance modelling was carried out for three common infiltration categories: runoff producing surfaces, surfaces receiving inflow of runon water from upstream zones, and a reference surface with zero runoff and runon. Runoff was calculated to 25%–30% of annual rainfall (which ranged from 488 to 596 mm), from crust observations, rainfall, soil wetness data, and infiltration estimates. Inflow of runon was estimated from field observations to 8%–18% of annual rainfall. The parameters in the functions for soil surface and canopy resistances were calibrated with field measurements of soil evaporation, stomatal conductance and leaf area. The model estimates of soil water contents, which were validated against neutron probe measurements, showed a reasonable agreement with observed data, with a root mean square error (RMSE) of approximately 0.02 m3 m−3 for 0–160 cm soil depth. Estimated productive water flow as plant transpiration was low, amounting to 4%–9% of the available water for the non-fertilised crop and 7%–24% for the fertilised crop. Soil evaporation accounted for 31%–50% of the available water, and showed a low variation for the observed range of leaf area (LAI <1 m2 m−2). Deep percolation was high, amounting to 200–330 mm for the non-crusted surfaces, which exceeded soil evaporation losses, for 1994–95 with relatively high annual rainfall (517–596 mm). Even a year with lower rainfall (488 mm) and a distinct dry spell during flowering (1996), resulted in an estimated deep percolation of 160 mm for the non-fertilised crop. The crop did not benefit from the additional inflow of runon water, which was partitioned between soil water storage and deep percolation. The only exception to this was the fertilised crop in 1996, where runon somewhat compensated for the limited rainfall and the higher water demand as a result of a larger leaf area than the non-fertilised crop. The effects of rainfall erraticness, resulting in episodic droughts, explain why a crop that uses such a small proportion of the available water, in an environment with substantial deep percolation, still suffers from water scarcity. Application of small levels of phosphorus and nitrogen roughly doubled yields, from 380 to 620 kg ha−1, and plant transpiration, from 33 to 78 mm. Evapotranspirational water use efficiency (WUEET) was low, 6500–8300 m3 ton−1 grain for non-fertilised crop, which is an effect of the low on-farm yields and high non-productive water losses. The estimated seasonal rainfall partitioning indicates the possibility of quantifying vertical water flows in on-farm environments in the Sahel, despite the presence of surface overland flow.  相似文献   

17.
Water balances have been constructed for three catchments using monthly data on rainfall (P), evapotranspiration (ET), stream discharge (Q) and groundwater levels. Length of record on the three catchments is 12, 8 and 6 years. Monthly, seasonal, and annual residuals (R) of the surface water balance equation R = PQET are used to infer changes in groundwater storage and are plotted against observed changes in groundwater storage (WTD). Linear regression analysis between R and WTD is used to examine the nature of the catchments' storage characteristics, the watertightness of the catchments, and the possibility that systematic measurement errors accumulate as the balance period lengthens.  相似文献   

18.
This study aimed to improve the understanding of hydrological processes in a humid (sub)tropical area in Africa with Inselberg topography. Additionally, the study intended to develop an approach for selective discharge data acquisition to determine water availability for smallholder irrigation in similar data-scarce catchments. During the December 2012–August 2013 field campaign meteorological and river stage data were collected at the Messica catchment in Central Mozambique. The 220 km2 catchment has an estimated 1000 ha of irrigated land, developed by smallholder farmers. Baseflow in the perennial tributary streams on the slopes of a meta-sedimentary Inselberg is the source of irrigation water. The baseflow recession curve of one of these tributaries is analysed and the water balance of an average year was determined. Precipitation, potential evapotranspiration, actual evapotranspiration and discharge were estimated to be 1224, 1462, 949 and 266 mm/year respectively. Differential gauging showed that the perennial tributaries gain water; the groundwater contribution increased with approximately 50% over two and a half month relative to the downstream discharge from March to May. In the downstream parts the groundwater contribution per metre stream length is between 30% and 100% higher compared to the upstream parts for two of the tributaries. Nevertheless, due to natural streambed infiltration and irrigation canals, discharge varies over the length of these tributaries. A rainfall–runoff model (HBV) was calibrated using the field data to examine the relation between precipitation characteristics and discharge at the start of the dry season. For precipitation scenarios with low and high intensity precipitation, discharges from June onwards were approximately similar in size according to the calibrated model. This suggest that discharge at the start of the dry season is mainly determined by total precipitation and the timing of precipitation (i.e. early or late in the wet season), not by individual rainfall events or rainfall intensity. It is concluded that the use of selective discharge measurements and low frequency precipitation measurements can effectively be used for water availability assessments in Inselberg catchments. Further research should be conducted to verify the validity of the used techniques in other humid sub-tropical Inselberg areas.  相似文献   

19.
ABSTRACT

The suspended sediment rating curves for six stations on four rivers in western and northern Greece are investigated. For each station the suspended sediment load is a power function of the water discharge, which may be distinguished according to wet and dry seasons; the latter yields higher sediment loads for a given discharge than the former. This is due to the higher erosivity of dry season rainfall compared to wet season rainfall producing the same runoff. All rating curve exponents b lie in the range 2.5–3.5 for the wet and 2.0–3.0 for the dry season and are related to the constants a of the curves by empirical equations. The variation in a and b is explained in terms of the annual precipitation and area of the basin, the hypsometric fall, the main channel length, and the average bedslope of the river from the basin divide to the station, through empirical relationships, which also permit the prediction of rating curves for ungauged basins.  相似文献   

20.
Hydraulic redistribution defined as the translocation of soil moisture by plant root systems in response to water potential gradients is a phenomenon widely documented in different climate, vegetation, and soil conditions. Past research has largely focused on hydraulic redistribution in deep tree roots with access to groundwater and/or winter rainfall, while the case of relatively shallow (i.e., ≈1–2 m deep) tree roots has remained poorly investigated. In fact, it is not clear how hydraulic redistribution in shallow root zones is affected by climate, vegetation, and soil properties. In this study, we developed a model to investigate the climate, vegetation, and soil controls on the net direction and magnitude of hydraulic redistribution in shallow tree root systems at the growing season to yearly timescale. We used the model to evaluate the effect of hydraulic redistribution on the water stress of trees and grasses. We found that hydraulic lift increases with decreasing rainfall frequency, depth of the rooting zone, root density in the deep soil and tree leaf area index; at the same time for a given rainfall frequency, hydraulic lift increases with increasing average rainstorm depth and soil hydraulic conductivity. We propose that water drainage into deeper soil layers can lead to the emergence of vertical water potential gradients sufficient to explain the occurrence of hydraulic lift in shallow tree roots without invoking the presence of a shallow water table or winter precipitation. We also found that hydraulic descent reduces the water stress of trees and hydraulic lift reduces the water stress of grass with important implications on tree–grass interactions.  相似文献   

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