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1.
Steppe‐tundra is considered to have been a dominant ecosystem across northern Eurasia during the Last Glacial Maximum. As the fossil record is insufficient for understanding the ecology of this vanished ecosystem, modern analogues have been sought, especially in Beringia. However, Beringian ecosystems are probably not the best analogues for more southern variants of the full‐glacial steppe‐tundra because they lack many plant and animal species of temperate steppes found in the full‐glacial fossil record from various areas of Europe and Siberia. We present new data on flora, land snails and mammals and characterize the ecology of a close modern analogue of the full‐glacial steppe‐tundra ecosystem in the southeastern Russian Altai Mountains, southern Siberia. The Altaian steppe‐tundra is a landscape mosaic of different habitat types including steppe, mesic and wet grasslands, shrubby tundra, riparian scrub, and patches of open woodland at moister sites. Habitat distribution, species diversity, primary productivity and nutrient content in plant biomass reflect precipitation patterns across a broader area and the topography‐dependent distribution of soil moisture across smaller landscape sections. Plant and snail species considered as glacial relicts occur in most habitats of the Altaian steppe‐tundra, but snails avoid the driest types of steppe. A diverse community of mammals, including many species typical of the full‐glacial ecosystems, also occurs there. Insights from the Altaian steppe‐tundra suggest that the full‐glacial steppe‐tundra was a heterogeneous mosaic of different habitats depending on landscape‐scale moisture gradients. Primary productivity of this habitat mosaic combined with shallow snow cover that facilitated winter grazing was sufficient to sustain rich communities of large herbivores.  相似文献   

2.
During the Quaternary glacial episodes, when sea level was considerably lower, Asia and North America were linked by large extensions of circumarctic land (Beringia), which remained unglaciated. This land mass served not only as a biogeographical bridge for plants, animals, and humans, but also supported a biome very different from present tundra or boreal coniferous forests, which was dominated by steppes and a rich mammalian megafauna. Carbon stable isotope ratios of Beringian late Pleistocene mummified remains of bison, equids, mammoth, caribou, musk-ox, moose, woolly rhino, and other undetermined species, found preserved in permafrost, indicate that these megaherbivores fed exclusively on C3 plants, and that C4 grasses were not differentially ingested by bison, as previously suggested. Paleoclimatic constraints probably prevented the formation of a warm-season (C4) guild during the later part of the growing season in the steppes of Beringia during the last glaciation.  相似文献   

3.
Chukotka is a key region for understanding both Quaternary environmental history and transcontinental migrations of flora and fauna during the Pleistocene as it lies at the far eastern edge of Asia bordering the Bering Sea. The now submerged land bridge is the least understood region of Beringia yet the most critical to understanding migrations between the Old and New Worlds. The insect fauna of the Main River Ledovy Obryv (Ice Bluff) section, which is late Pleistocene in age (MIS 3-2), is markedly different from coeval faunas of areas further to the west, as it is characterized by very few thermophilous steppe elements. From the fauna we reconstruct a steppe-tundra environment and relatively cold conditions; the reconstructed environment was moister than that of typical steppe-tundra described from further west. The data from this locality, if typical of the Chukotka Peninsula as a whole, may indicate that a barrier associated with the environments of the land bridge restricted trans-Beringian migrations, particularly the more thermophilous and xeric-adapted elements of the Beringian biota, supporting the hypothesis of a cool but moist land-bridge filter inferred from evidence from several other studies.  相似文献   

4.
Fossil arctic ground squirrel (Spermophilus parryii) middens were recovered from ice-rich loess sediments in association with Sheep Creek-Klondike and Dominion Creek tephras (ca 80 ka) exposed in west-central Yukon. These middens provide plant and insect macrofossil evidence for a steppe-tundra ecosystem during the Early Wisconsinan (MIS 4) glacial interval. Midden plant and insect macrofossil data are compared with those previously published for Late Wisconsinan middens dating to ～25–2914C ka BP (MIS 3/2) from the region. Although multivariate statistical comparisons suggest differences between the relative abundances of plant macrofossils, the co-occurrence of steppe-tundra plants and insects (e.g., Elymus trachycaulus, Kobresia myosuroides, Artemisia frigida, Phlox hoodii, Connatichela artemisiae) provides evidence for successive reestablishment of the zonal steppe-tundra habitats during cold stages of the Late Pleistocene. Arctic ground squirrels were well adapted to the cold, arid climates, steppe-tundra vegetation and well-drained loessal soils that characterize cold stages of Late Pleistocene Beringia. These glacial conditions enabled arctic ground squirrel populations to expand their range to the interior regions of Alaska and Yukon, including the Klondike, where they are absent today. Arctic ground squirrels have endured numerous Quaternary climate oscillations by retracting populations to disjunct “interglacial refugia” during warm interglacial periods (e.g., south-facing steppe slopes, well-drained arctic and alpine tundra areas) and expanding their distribution across the mammoth-steppe biome during cold, arid glacial intervals.  相似文献   

5.
Botanical analyses of fossil and modern arctic ground squirrel (Urocitellus parryii) caches and nests have been used to reconstruct the past vegetation from some parts of Beringia, but such archives are understudied in Alaska. Five modern and four fossil samples from arctic ground squirrel caches and nests provide information on late Pleistocene vegetation in Eastern Beringia. Modern arctic ground squirrel caches from Alaska's arctic tundra were dominated by willow and grass leaves and grass seeds and bearberries, which were widespread in the local vegetation as confirmed by vegetation surveys. Late Pleistocene caches from Interior Alaska were primarily composed of steppe and dry tundra graminoid and herb seeds. Graminoid cuticle analysis of fossil leaves identified Calamagrostis canadensis, Koeleria sp. and Carex albonigra as being common in the fossil samples. Stable carbon isotopes analysis of these graminoid specimens indicated that plants using the C3 photosynthetic pathways were present and functioning with medium to high water-use efficiency. Fossil plant taxa and environments from ground squirrel caches in Alaska are similar to other macrofossil assemblages from the Yukon Territory, which supports the existence of a widespread mammoth steppe ecosystem type in Eastern Beringia that persisted throughout much of the late Pleistocene.  相似文献   

6.
Plant and insect macrofossil assemblages dating to the full-glacial (late Wisconsinan) are rare from eastern Beringia. Here we present an assemblage of fossil pollen, insect and plant macrofossils recovered from alluvium at the Bluefish Exposure, northern Yukon Territory. Nine AMS radiocarbon ages place these data between ca. 18,880–16,440 14C yr BP (22,313–19,597 cal. yr BP). These data indicate that xeric steppe, rich in bunchgrasses Poa and Elymus, Artemisia frigida and diverse forbs was interspersed within a mosaic of local vegetation types, including mid-rich fens, mesic graminoid meadows, steppe-tundra and herb-tundra. Macrofossils and minor pollen of tundra forbs suggest steppe-tundra plant associations within midslope elevations and discontinuous herb-tundra on high elevation uplands on exposed bedrock ridges. The composition and distribution of local vegetation was dependent on available moisture, drainage, aspect and elevation. Compositional and physiognomic similarities can be made with extrazonal steppe-dominated dry slopes and high elevation steppe-tundra ecotones in central Alaska and Yukon Territory. Our paleoecological data reflect environments inhabited by the diverse late Pleistocene Bluefish Caves fauna, including woolly mammoth, horse, steppe bison, and saiga antelope.  相似文献   

7.
《Quaternary Science Reviews》2007,26(7-8):979-1003
This paper presents paleoecological analyses of 48 fossil arctic ground squirrel (Spermophilus parryii) middens (nests and caches) recovered from ice-rich loess sediments in the Klondike region of west-central Yukon Territory. AMS radiocarbon dates and stratigraphic association of middens with Dawson tephra (∼25 300 14C yr BP), indicate these paleoecological data reflect the onset of glacial conditions of early Marine Isotope Stage (MIS) 2 and terminal MIS 3 (∼24 000–29 450 14C yr BP). Plant macrofossils include at least 60 plant taxa, including diverse graminoids (Poa, Elymus trachycaulus, Kobresia myosuroides), steppe forbs (Penstemon gormanii, Anemone patens var. multifida, Plantago cf. canescens), tundra forbs (Draba spp., Bistorta vivipara), dwarf shrubs (Salix cf. arctica, S. cf. polaris), sage (Artemisia frigida) and rare trees (Picea mariana). Many of these taxa identified in the middens represent the first recorded fossils for these plants in Eastern Beringia and add to our knowledge of the floristic composition of Pleistocene vegetation and biogeography in this region. Fossil beetles include typical members of the Eastern Beringian steppe–tundra fauna (Lepidophorus lineaticollis and Connatichela artemisiae) and others suggesting predominantly dry, open habitats. Cache forage selection is suggested by some plant taxa which were particularly frequent and abundant in the middens (Bistorta vivipara, Kobresia myosuroides, Ranunculus spp., Potentilla, Erysimum cf. cheiranthoides, Poa, Carex and Draba). Factors such as proximity of vegetation to burrows and abundance of fruits and seeds per plant were probably important in cache selection. Glacial conditions enabled arctic ground squirrels to form widespread and dense populations in regions such as the Klondike in which they are rare or absent at present. This fossil midden record supports previous hypotheses that suggest arctic ground squirrels evolved in and are well-adapted to the open, steppe–tundra vegetation, loessal soils and glacial climates of the mammoth-steppe biome.  相似文献   

8.
Herbaceous communities in forest ecosystems on the southern part of the Russian Plain appeared in the Middle Miocene (??10 Ma BP). In the Late Miocene (??7 Ma BP), feather-grass steppe associations appeared among them. In the time span of 2.7 to 2.1 Ma BP (i.e., in the Early Quaternary, according to the current chronostratigraphic scale), the steppe zone arose on the southern Russian Plain in the Don-Azov Region. Seven stages of this zone development here have been distinguished throughout the Quaternary. The first one (Eopleistocene-Early Pleistocene) was characterized by savanna-like subtropic ecosystems. Then, in the Middle Pleistocene, the temperate zone ecosystems (tallgrass prairie-like steppes) developed here and were followed by steppe ecosystems close to the modern ones in Central Europe. The ecosystems of rich-species forb steppes developed in the Late Pleistocene. Finally, in the optimum of the modern interglacial (Holocene), steppes became similar to the modern ones here, but with a slightly higher precipitation. The general trend is characterized by reduction in heat and water provision and increase in aridization progressing from earlier to later stages.  相似文献   

9.
Cryolithological, ground ice and fossil bioindicator (pollen, diatoms, plant macrofossils, rhizopods, insects, mammal bones) records from Bol'shoy Lyakhovsky Island permafrost sequences (73°20′N, 141°30′E) document the environmental history in the region for the past c. 115 kyr. Vegetation similar to modern subarctic tundra communities prevailed during the Eemian/Early Weichselian transition with a climate warmer than the present. Sparse tundra‐like vegetation and harsher climate conditions were predominant during the Early Weichselian. The Middle Weichselian deposits contain peat and peaty soil horizons with bioindicators documenting climate amelioration. Although dwarf willows grew in more protected places, tundra and steppe vegetation prevailed. Climate conditions became colder and drier c. 30 kyr BP. No sediments dated between c. 28.5 and 12.05 14C kyr BP were found, which may reflect active erosion during that time. Herb and shrubby vegetation were predominant 11.6–11.3 14C kyr BP. Summer temperatures were c. 4 °C higher than today. Typical arctic environments prevailed around 10.5 14C kyr BP. Shrub alder and dwarf birch tundra were predominant between c. 9 and 7.6 kyr BP. Reconstructed summer temperatures were at least 4 °C higher than present. However, insect remains reflect that steppe‐like habitats existed until c. 8 kyr BP. After 7.6 kyr BP, shrubs gradually disappeared and the vegetation cover became similar to that of modern tundra. Pollen and beetles indicate a severe arctic environment c. 3.7 kyr BP. However, Betula nana, absent on the island today, was still present. Together with our previous study on Bol'shoy Lyakhovsky Island covering the period between about 200 and 115 kyr, a comprehensive terrestrial palaeoenvironmental data set from this area in western Beringia is now available for the past two glacial–interglacial cycles.  相似文献   

10.
Two conflicting stratigraphic schemes describe the Siberian Karginskii interstade (Oxygen Isotope Stage 3 equivalent) as having: 1) relatively stable climate with environments more similar to the full glaciation; or 2) variable climate with landscapes that more closely approximate contemporary ones. New data from continuous lake cores and a nearly continuous section from western Beringia (WB) suggest that both schemes are valid. Herb-dominated communities, possibly with isolated populations of Larix, characterized northern WB with only a slight shift from relatively warm to cool summers during the mid-interstade. In contrast, herb and shrub tundra, steppe, forest-tundra, and modern Larix forest occurred at various times in areas of southern WB, suggesting greater climatic instability. A thermal optimum is evident in the south during the mid-interstade, with modern vegetation in southeastern WB and Larix forest-tundra in the southwest. Variations in Pinus pumila pollen indicate summer warm/winter dry and summer warm/winter wet conditions in southeastern WB. These fluctuations contrast to other areas of WB, where summers and probably winters were consistently arid. Although the interstade presents a unique interval within the Late Pleistocene, paleodata and paleoclimatic models suggest that changes in marine conditions, including sea level, were likely key drivers in the regional climate history.  相似文献   

11.
Although much emphasis has been placed on the effects of Pleistocene ice sheet as the determinant of Pleistocene climate in Europe, only through the combined evaluation of all useful climatic indicators can Pleistocene climatic zones be differentiated. The relationships between a paleoclimatically determined snowline and polar treelines and extent of loess deposition, determined stratigraphically, botanically, and morphologically, also indicate climatic conditions of Pleistocene Europe. Five great climatic-morphologic and plant-geographic zones, namely, forest-rubble tundra, forest tundra, loess tundra, loess steppe, and loess-forest steppe, may be distinguished. Considering these factors and their characteristics and spatial and temporal extents, it is possible to chronologically present the differentiation of the Würm glacial period independent of the retardation effect of ice sheets. --G. E. Denegar.  相似文献   

12.
Wisconsinan full-glacial silts filling a swale exposed in Conklin Quarry, Johnson Co., Iowa, contain a large and diverse biota that includes pollen, bryophytes, vascular-plant macrofossils, small mammals, molluscs, and insects. Radiocarbon dates on wood from the top, middle and bottom of the swale fill respectively were 16710 ± 270, 17 170 ± 205, and 18090 ± 190 yr BP. The pollen diagram is dominated by Picea (spruce), Pinus (pine), and Cyperaceae (sedge), and it records low pollen accumulation rates. Plant macrofossils include a number of tundra species along with Picea and Larix (larch) needles and small pieces of wood. The insect fauna contains many species now confined to the forest-tundra transition zone of northwestern Yukon and Alaska. Small mammals include the tundra indicators Dicrostonyx (collared lemming), and probably Microtus miurus (singing vole) together with boreal forest taxa. The molluscs include extinct and relict species and show the widest range in present geographic distribution, but Rocky Mountain and especially northern elements predominate in the swale fill. All these lines of evidence lead to consistent palaeoclimato-logical interpretation and palaeoecological reconstruction. The dominant habitats represented by the biota and sedimentary environment collectively included open calcareous silty to sandy or gravelly upland sites, minerotrophic fens (wetlands), pond- or stream-side clayey to sandy shores, and shallow (possibly ephemeral), cold, clear-water ponds. Mean July temperatures were probably 11° to 13°C cooler than at present. The biota indicates that a Picea-Larix krummholz with extensive tundra openings was present in southeastern Iowa between 18090 and 16710 yr BP.  相似文献   

13.
Our understanding of the timing of human arrival to the Americas remains fragmented, despite decades of active research and debate. Genetic research has recently led to the ‘Beringian standstill hypothesis’ (BSH), which suggests an isolated group of humans lived somewhere in Beringia for millennia during the Last Glacial, before a subgroup migrated southward into the American continents about 14 ka. Recently published organic geochemical data suggest human presence around Lake E5 on the Alaskan North Slope during the Last Glacial; however, these biomarker proxies, namely faecal sterols and polycyclic aromatic hydrocarbons (PAHs), are relatively novel and require replication to bolster their support of the BSH. We present new analyses of these biomarkers in the sediment archive of Burial Lake (latitude 68°26′N, longitude 159°10′W m a.s.l.) in northwestern Alaska. Our analyses corroborate that humans were present in Beringia during the Last Glacial and that they likely promoted fire activity. Our data also suggest that humans coexisted with Ice Age megafauna for millennia prior to their eventual extinction at the end of the Last Glacial. Lastly, we identify fire as an overlooked ecological component of the mammoth steppe ecosystem.  相似文献   

14.
Thousands of Late Pleistocene remains are found in sites throughout Beringia. These specimens comprise an Ice Age genetic museum, and the DNA contained within them provide a means to observe evolutionary processes within populations over geologically significant time scales. Phylogenetic analyses can identify the taxonomic positions of extinct species and provide estimates of speciation dates. Geographic and temporal divisions apparent in the genetic data can be related to ecological change, human impacts, and possible landscape mosaics in Beringia. The application of ancient DNA techniques to traditional paleontological studies provides a new perspective to long-standing questions regarding the paleoenvironment and diversity of Late Pleistocene Beringia.  相似文献   

15.
Field investigations of caves along Alaska's Porcupine River document three major mechanisms which modify bone in patterns similar to alterations produced by man: (1) carnivore fracture; (2) rodent gnawing; and (3) rock fall and rubble scarring. A late Wisconsin faunal assemblage composed of Equus sp., Rangifer tarandus, Ovis dalli, Bison sp., proboscidean, numerous small mammal species, birds, and fish is well documented. This faunal assemblage suggests a mosaic environment of grassland-tundra-forest in the immediate vicinity of these caves and implies that the late Wisconsin environment in north-central Alaska may have been characterized by a number of microenvironments and colder, dryer, steppe conditions. Taphonomic data which have historically been interpreted to support human occupation of eastern Beringia during the Pleistocene are critically examined and the context of these discoveries (not the specimens themselves) provides the test essential to document the antiquity of man in North America prior to 12,000 yr ago.  相似文献   

16.
Chlachula, J. & Serikov, Y. B. 2010: Last glacial ecology and geoarchaeology of the Central Trans‐Ural area: the Sosva River Upper Palaeolithic Complex, western Siberia. Boreas, 10.1111/j.1502‐3885.2010.00166.x. ISSN 0300‐9483. Quaternary and geoarchaeology studies from the eastern limits of the Ural Mountains provide multiple lines of evidence of the Palaeolithic peopling of this geographically marginal and still poorly explored territory of western Siberia following the mid‐last glacial (MIS 3) warming. A complex of investigated open‐air localities in the Sosva River basin (the north‐central Trans‐Ural area) at the periphery of the western Siberian Plain, distinguished by very high concentrations of Pleistocene megafaunal remains previously regarded as ‘mammoth cemeteries’, indicate, in conjunction with the associated diagnostic ivory/bone and stone industry, open occupation sites during the Last Glacial (MIS 2). Fossil faunal remains, dominated by mammoth (98%) together with bird and fish species, indicate various methods of exploitation of the Late Pleistocene natural resources and successful behavioural adaptation to the last glacial sub‐polar tundra‐steppe environment. The taphonomy and composition of the well‐preserved skeletal remains from the main occupation sites suggest both active hunting and anthropogenic ‘scavenging’ practices. The contextual geology and the cultural and biotic multi‐proxy records from the Trans‐Ural Upper Palaeolithic Complex provide new insights into the timing and palaeoecological conditions of the Pleistocene human occupation of north‐central Asia.  相似文献   

17.
Unglaciated parts of the Yukon constitute one of the most important areas in North America for yielding Pleistocene vertebrate fossils. Nearly 30 vertebrate faunal localities are reviewed spanning a period of about 1.6 Ma (million years ago) to the close of the Pleistocene some 10 000 BP (radiocarbon years before present, taken as 1950). The vertebrate fossils represent at least 8 species of fishes, 1 amphibian, 41 species of birds and 83 species of mammals. Dominant among the large mammals are: steppe bison (Bison priscus), horse (Equus sp.), woolly mammoth (Mammuthus primigenius), and caribou (Rangifer tarandus) – signature species of the Mammoth Steppe fauna (Fig. 1), which was widespread from the British Isles, through northern Europe, and Siberia to Alaska, Yukon and adjacent Northwest Territories. The Yukon faunas extend from Herschel Island in the north to Revenue Creek in the south and from the Alaskan border in the west to Ketza River in the east. The Yukon holds evidence of the earliest-known people in North America. Artifacts made from bison, mammoth and caribou bones from Bluefish Caves, Old Crow Basin and Dawson City areas show that people had a substantial knowledge of making and using bone tools at least by 25 000 BP, and possibly as early as 40 000 BP. A suggested chronological sequence of Yukon Pleistocene vertebrates (Table 1) facilitates comparison of selected faunas and indicates the known duration of various taxa.  相似文献   

18.
The malacological material of the mound bodies (kurgans) of the Great Hungarian Plain indicates a mixed vegetation of dry and humid environments, developed on a mosaic of alkaline and chernozem soils in the period of the construction of the kurgan. The malacofauna that evolved in the upper soil horizon of the mound indicates the extremely dry environmental conditions of steppes, charaterized by the dominance of thermoxerophilous species. Dominant species of this kurgan are Chondrula tridens, Helicopsis striata, Granaria frumentum and Cepaea vindobonensis. The species composition shows that there are differences in the malacofauna of the Danube-Tisa Interfluves region compared to that of the lowlands east of River Tisza, indicated by the higher dominance of Granaria frumentum and Helicopsis striata in the former region. Following the construction of the kurgans an island-like, dry habitat developed on their surface, covered by black soil and populated by a steppe fauna, the composition of which is comparable mostly with the mollusc fauna of loess steppes and forest steppe, irrespectively of the floodplain or wind-blown sand character of the original surface.  相似文献   

19.
The comparative analysis of palynomorphs and plant megafossils (fruits, seeds, twigs, leaves) in the Upper Pleistocene host sediments and materials filling in fossil burrows of gophers, their coprolites included, at the Duvannyi Yar, Stanchikovskii Yar and Zelenyi Mys sites of the Kolyma Lowland is carried out. Genera Salix, Lychnis, Silene, Draba, Potentilla, Larix, and families Poaceae, Polygonaceae, Cyperaceae, Compositae, and Leguminosae are determined among palynological remains and megafossils. Factors responsible for qualitative and quantitative differences in taxonomic compositions of palynological and megafossil assemblages are biological peculiarities of plants, different character of fossilization of palynomorphs and large plant remains, geographic conditions, different genesis of assemblages (allochthonous for microfossils and autochthonous for megafossils), and inadequately known morphology of certain spore and pollen taxa. The comprehensive paleobotanical analysis leads to the conclusion that the study region was occupied in the Late Pleistocene by plant communities of humid to somewhat dryer tundra with separate areas of pioneering and steppe vegetation.  相似文献   

20.
Plant macrofossils from the “Mamontovy Khayata” permafrost sequence (71°60′N, 129°25′E) on the Bykovsky Peninsula reflect climate and plant biodiversity in west Beringia during the last cold stage. 70 AMS and 20 conventional 14C dates suggest sediment accumulation between about 60,000 and 7500 14C yr B.P. The plant remains prove that during the last cold-stage arctic species (Minuartia arctica, Draba spp., Kobresia myosuroides) coexisted with aquatic (Potamogeton vaginatus, Callitriche hermaphroditica), littoral (Ranunculus reptans, Rumex maritimus), meadow (Hordeum brevisubulatum, Puccinellia tenuiflora) and steppe taxa (Alyssum obovatum, Silene repens, Koeleria cristata, Linum perenne). The reconstructed vegetation composition is similar to modern vegetation mosaics in central and northeast Yakutian relict steppe areas. Thus, productive meadow and steppe communities played an important role in the Siberian Arctic vegetation during the late Pleistocene and could have served as food resource for large populations of herbivores. The floristic composition reflects an extremely continental, arid climate with winters colder and summers distinctly warmer than at present. Holocene macrofossil assemblages indicate a successive paludification possibly connected with marine transgression, increased oceanic influence and atmospheric humidity. Although some steppe taxa were still present in the early Holocene, they disappeared completely before 2900 14C yr B.P.  相似文献   

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