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1.
Reindeer (Rangifer tarandus) populations reached the Iberian Peninsula during the end of the Middle Pleistocene and there are numerous examples of this species from Late Pleistocene Mousterian and Upper Palaeolithic sites. In this paper, new evidence of reindeer in the east Cantabrian region is presented that further supports the timing of this species’ first appearance, and previous assessments are updated. To date, the presence of this species has been identified at 55 sites in the Iberian Peninsula, nearly as many as those of mammoth (Mammuthus primigenius) and woolly rhinoceros (Coelodonta antiquitatis) combined. Most of the sites with presence of reindeer (50) are located in the Cantabrian region with a clear increase in the density of sites and remains towards the Pyrenees. The remaining five sites with evidence of reindeer are located on the other side of the Pyrenees in the NW corner of Catalonia. In contrast, archaeological evidence of reindeer in the form of art (both parietal and portable) is more scarce and scattered. Evidence for the representation of these animals has been found outside the northern fringe of the Iberian Peninsula, which could reflect either long‐distance cultural communication or the movement of human groups.  相似文献   

2.
Kuzmin, Y. V. 2009: Extinction of the woolly mammoth (Mammuthus primigenius) and woolly rhinoceros (Coelodonta antiquitatis) in Eurasia: Review of chronological and environmental issues. Boreas, 10.1111/j.1502‐3885.2009.00122.x. ISSN 0300‐9483. The current evidence for date and environmental preferences of the extinction of two middle–late Pleistocene megafaunal species, the woolly mammoth (Mammuthus primigenius Blum.) and woolly rhinoceros (Coelodonta antiquitatis Blum.), is presented in this review. It is suggested that extinction of these large herbivores in Eurasia was closely related to landscape changes near the Pleistocene–Holocene boundary (c. 12 000–9000 uncalibrated radiocarbon years ago, yr BP), mainly involving the widespread forest formations in the temperate and arctic regions of northern Eurasia and the loss of grasslands crucial to the existence of woolly mammoth and rhinoceros. However, some woolly mammoth populations survived well into the Holocene (up to c. 3700 yr BP), showing that the process of final extinction was fairly complex, with delays in some regions of up to several millennia. The possible role of Palaeolithic humans in the extinction of Late Pleistocene megafauna is also considered.  相似文献   

3.
The La Parte (Asturias) northern Spain site contains a cold-adapted mammal faunal assemblage that corresponds to a level radiometrically dated to a minimum age of 150 ka. It represents the most ancient site with cold resistant fauna in the Iberian Peninsula.Among the species recorded in La Parte, Coelodonta antiquitatis and Rangifer tarandus represent the typical cold-adapted large mammal association.The presence of the woolly rhino at La Parte provides relevant environmental information about the Mammoth Steppe. This species was not as abundant in the Middle Pleistocene as during the Late Pleistocene, when its cold-adapted features can be studied from an evolutionary perspective. The remains attributed to R. tarandus suggest an open steppe ecosystem.The presence of Marmota sp. is also confirmed in La Parte. A first interpretation would suggest an alpine or periglacial ecosystem, but based on recent results which do not support the correlation between phylogeny and climatic tolerance for extant species of Marmota (López and Cuenca, 2002. Palaeogeography, Palaeoclimatology, Palaeoecology 186, 311), we propose the presence of this rodent suggests an open landscape with cold conditions.The rest of the taxa included in the La Parte assemblage (Crocuta crocuta, Panthera leo, Cervus elaphus, Megaloceros cf. giganteus, Bison priscus and Equus caballus) are usually associated with typical cold-adapted faunas, but they are also found in woody temperate zones so they do not characterize by themselves a cold period. The faunal association from La Parte suggests a combination of steppic and open woodland ecological conditions.  相似文献   

4.
从采自辽宁省海城县小孤山晚更新世披毛犀化石样品中成功地获得了1 080 bp细胞色素b基因序列,调用Genbank中已发表的披毛犀序列及5种现生犀牛的同源序列,以马作为外类群,采用邻接法(NJ法)和最大似然法(ME法)构建系统发育树,结果均显示披毛犀与现生苏门答腊犀亲缘关系最近,且辽宁省海城县小孤山样品处于披毛犀分支的根部;对披毛犀样品进行基因分异度分析结果表明,小孤山的披毛犀样品具有较高的基因分异度,为探讨披毛犀的起源、迁徙及演化等问题提供了重要的分子依据.  相似文献   

5.
The so-called Hundsheim rhinoceros, Stephanorhinus hundsheimensis, was a very common faunal element of the Early to early Middle Pleistocene period in the western Palaearctic. In this study, individuals from two different central European populations of the Hundsheim rhinoceros were analysed in order to determine whether their local dietary signals could reflect differing food availability between the two populations, and whether such information could provide a better understanding of the ecological role of S. hundsheimensis within corresponding faunal assemblages, and of its principal subsistence strategy in the western Palaearctic. The mesowear traits observed in the studied S. hundsheimensis populations have been interpreted as representing biome-specific signals, indicating grassland vegetation at the site of Süßenborn, and dense to open forests at Voigtstedt (both localities in Germany). The analyses performed on the fossil rhino material demonstrate the most pronounced dietary variability ever established for a single herbivorous ungulate species by mesowear studies. This variability ranges from an attrition dominated grazing regime, to a one of predominantly browsing, and characterises S. hundsheimensis as the most ecologically tolerant rhinoceros of the Palaearctic Plio-Pleistocene. Although such dietary flexibility proved an effective enough subsistence strategy over a period of 600–900 ka (1.4/1.2–0.6/0.5 Myr) in the western Palaearctic, the situation changed dramatically after 0.6 Myr BP, when the new species of rhinoceroses, Stephanorhinus hemitoechus and Stephanorhinus kirchbergensis, appeared and started to compete for both the grass and the browse. For the generalist S. hundsheimensis, this bilateral interference was detrimental to its success in all of its habitats. The successful competition of specialised forms of rhinoceroses, which might have originated as a result of the development of 100 ka periodicity in the global climatic record, is proposed as the main reason for the extinction of S. hundsheimensis during the early Middle Pleistocene.  相似文献   

6.
New TIMS dates from British sites underpin and extend the growing model of mammalian faunal history during the Late Pleistocene. The sequence of British faunas during the three successive warm phases of MIS 5 presents a challenging model in which the mammal faunas of the first two temperate phases, 5e and 5c is dominated by species of a classically interglacial character, while that of the final phase, sub‐stage 5a, is dominated by bison and reindeer. This we see as the ‘cold fauna’ appearing in Britain during the cool isotope sub‐stage 5b and subsequently isolated by the rising sea level of isotope sub‐stage 5a. All these three faunas are true island faunas. Copyright © 2007 John Wiley & Sons, Ltd.  相似文献   

7.
Major changes in community structure and depositional relief of high-latitude coral communities in the southern Persian Gulf between marine isotope stage (MIS) 7 and the present day suggest that the area has become increasingly restricted. Corals and bivalves from outcrops on Kish Island, Iran, were identified in order to interpret the Late Pleistocene palaeoenvironmental setting. U/Th disequilibrium dating was used to constrain the ages of the stratigraphic units. During MIS 7, two coral-bearing sequences were deposited on what is now Kish Island. The lower sequence is dated as MIS 7.5 and changes laterally from an assemblage dominated by Cyphastrea sp. and Platygyra daedalea in the west to one characterized by branching Montipora in the east. By contrast, the upper sequence, dated as MIS 7.1, transitions from an assemblage dominated by platy Montipora in the west to a diverse assemblage of Platygyra and other faviids in the east. The assemblages of both sequences are within a marl matrix and bounded by thin lithified mollusc-rich layers. Corals and bivalves indicate that the sequences were deposited on gentle slopes in sheltered environments less than 20 m deep. The MIS 7 deposits may be classified as coral carpets or biostromes that developed a low-relief framework. During MIS 5, coral communities were no longer framework building and are now limited to an Acropora-rich layer of coral rubble that covers large parts of the island, and two small incipient reefs with sparse faviids. Similarities between the MIS 5 and modern nearshore coral communities suggest that the environmental conditions during MIS 5 were comparable to those of today. The late Pleistocene coral carpets and non-framework coral communities of the southern Persian Gulf may serve as models for coral biostromes in the fossil record, which formed under restricted environmental conditions such as elevated terrigenous input, high turbidity, and strong seasonal changes in temperature and/or salinity.  相似文献   

8.
A nearly complete frozen mummy of a woolly rhinoceros (Coelodonta antiquitatis Blum., 1799) was discovered in a gold mine on the lower reaches of the Kolyma River, north–eastern Siberia. This is the first find of the whole body of woolly rhino in permafrost. A large part of the mummified body was preserved, including the left part of the body, covered by skin, including skin of the head and ear, fore and hind legs. The skull with 2 horns and the lower jaw were also preserved. Most of the internal organs were lost, except the intestines, stomach, and their contents. A rib fragment from this individual was dated by AMS-radiocarbon method to 39,140 ± 390 years BP (OxA-18755). Spore and pollen analyses of the stomach contents indicate that grasses and sagebrushes formed the main part of the diet of C. antiquitatis in this region of Arctic Siberia.  相似文献   

9.
The investigation of a limestone cave on the Ryonggok-ri of Sangwon County, North Hwanghae Province, DPRK, has yielded many faunal remains including Late Pleistocene deer, brown bears and horses. Uranium series dating of fossil teeth yielded dates of ~72ka at the base of Layer 2 – the bone-bearing unit – and ~44ka at the top. The rich and diverse mammal remains of Ryonggok Cave No. 1 are indicative of warm and humid temperate conditions during the Late Pleistocene, and reflect the presence of both dense forests and open grassland. In addition, pollen and spores from the site include two families and 24 genera of trees, six families and a genus of grasses and herbs, and two families and six genera of ferns, corroborating the environments inferred from the faunal remains.  相似文献   

10.
Fossil arctic ground squirrel (Spermophilus parryii) middens were recovered from ice-rich loess sediments in association with Sheep Creek-Klondike and Dominion Creek tephras (ca 80 ka) exposed in west-central Yukon. These middens provide plant and insect macrofossil evidence for a steppe-tundra ecosystem during the Early Wisconsinan (MIS 4) glacial interval. Midden plant and insect macrofossil data are compared with those previously published for Late Wisconsinan middens dating to ~25–2914C ka BP (MIS 3/2) from the region. Although multivariate statistical comparisons suggest differences between the relative abundances of plant macrofossils, the co-occurrence of steppe-tundra plants and insects (e.g., Elymus trachycaulus, Kobresia myosuroides, Artemisia frigida, Phlox hoodii, Connatichela artemisiae) provides evidence for successive reestablishment of the zonal steppe-tundra habitats during cold stages of the Late Pleistocene. Arctic ground squirrels were well adapted to the cold, arid climates, steppe-tundra vegetation and well-drained loessal soils that characterize cold stages of Late Pleistocene Beringia. These glacial conditions enabled arctic ground squirrel populations to expand their range to the interior regions of Alaska and Yukon, including the Klondike, where they are absent today. Arctic ground squirrels have endured numerous Quaternary climate oscillations by retracting populations to disjunct “interglacial refugia” during warm interglacial periods (e.g., south-facing steppe slopes, well-drained arctic and alpine tundra areas) and expanding their distribution across the mammoth-steppe biome during cold, arid glacial intervals.  相似文献   

11.
The Balderton Terrace marks a former course of the River Trent between Newark and the Lincoln Gap. The principal deposit, the Balderton Sand and Gravel, is interpreted as a braided river sediment. Ice wedge casts truncated by intraformational erosion surfaces at many levels indicate syndepositional permafrost. Remnant cover deposits overlying the Balderton Sand and Gravel include the partly aeolian Whisby Sand. Locally, both the upper part of the Balderton Sand and Gravel and the cover deposits exhibit features indicative of temperate climate pedogenesis. All these deposits are affected by subsequent cryoturbation. On the basis of these features and the geomorphological and topographical relationship to other terrace deposits of the area, the Balderton Sand and Gravel and Whisby Sand are regarded as post-Hoxnian and pre-lpswichian, i.e. Wolstonian. Electron spin resonance age determinations for fossil elephant teeth and amino acid analyses on molluscs from the Balderton Sand and Gravel suggest correlation with Oxygen Isotope Stage 6. The Balderton Sand and Gravel has yielded a cold-climate mammalian fauna dominated by woolly mammoth and woolly rhinoceros, though rarer species suggest periods of milder climate. Silts from channels near the base of the deposit have produced pollen, mollusc, ostracod and beetle assemblages also indicating a cold climate.  相似文献   

12.
Woolly mammoths were large, herbivorous, cold-adapted mammals of the Late Pleistocene. The diet and habitat requirements of the species set certain constraints on the palaeoenvironments it could occupy. The relationship between the mammoth’s shifting range and changing environments can be explored using independent data on ice sheet configuration, temperature, and vegetation, provided the locality and age of the fossil remains can be validated. Here we present a comprehensive record of occurrence of the woolly mammoth in the circum-Baltic region of northern Europe during the last glaciation, based on a compilation of radiocarbon-dated remains. The record shows that the mammoth was widespread in northern and north-eastern Europe during Marine Isotope Stage 3 (MIS 3), at 50,000–30,000 calibrated years ago (50–30 ka). The presence of the species up to 65°N latitude supports the restriction of the Scandinavian Ice Sheet (SIS) during MIS 3. The widest distribution range round 30 ka was followed by a decline that led to the disappearance of mammoths from the area during the maximum extent of the SIS, from 22 to 18 ka. The woolly mammoth re-colonized the Baltic region and southern Scandinavia after the onset of the late-glacial deglaciation at 17 ka. The late-glacial record suggests a markedly fluctuating population changing its range in tune with the rapid environmental changes. The last appearance of mammoth in our study region was in Estonia during the Younger Dryas (Greenland Stadial 1; GS1) at about 12 ka. The two major periods of occurrence during MIS 3 and the late-glacial stadial suggest that mammoth had a wide tolerance of open to semi-open tundra and steppe-tundra habitats with intermediately cold climate, whereas the 22–18 ka disappearance suggests a major southward and/or eastward retreat in response to extremely cold, glacial conditions near the SIS margin. The final regional extinction correlates with the re-forestation during the rapid warming at the Younger Dryas–Holocene boundary.  相似文献   

13.
The late Middle Pleistocene sites in the Ebbsfleet Valley, Kent, UK, have yielded archaeological assemblages critical to understanding the early Middle Palaeolithic of northwestern Europe. Despite a long history of research, the nature and context of these assemblages are still poorly understood. This paper clarifies the stratigraphic, environmental and archaeological records at Ebbsfleet. These reflect a cold–warm–cold sequence of climatic events, preserved within part of the Taplow/Mucking Formation of the Thames (Marine Isotope Stage (MIS) 8/7/6). Levallois artefacts are shown to be restricted to the lower part of the Ebbsfleet Channel sequence (Phases I and II) and are assigned to late MIS 8/early MIS 7. This material is associated with fauna indicative of open environments during both cool and temperate conditions. Handaxe assemblages are recorded from higher up the sequence (Phases III–V), but have been redeposited from higher terrace units nearby. No primary context archaeology is apparent during these later phases of aggradation. This may indicate that humans abandoned the site once available raw material became inaccessible, and may also reflect a decline in human presence in Britain during the latter part of MIS 7. Copyright © 2010 John Wiley & Sons, Ltd.  相似文献   

14.
渤海湾西岸晚第四纪海相地层划分及地质意义   总被引:5,自引:0,他引:5  
BT113孔和BT114孔是位于渤海湾西岸地区的2个80 m全取心钻孔。通过对钻孔岩心的岩性、沉积构造及生物组合等特征分析,结合AMS14C和OSL(光释光)测年结果,揭示中更新世晚期以来本地区主要发育3个海相地层:自下而上为中更新统上部的第三海相地层(简称“海三”)、上更新统的第二海相地层(简称“海二”)和全新统的第一海相地层(简称“海一”),指示中更新世晚期以来研究区发生的3次主要海面波动。与全球海面变化曲线对比,海三,海二和海一分别发育于MIS(深海氧同位素阶段)7、MIS5和MIS1的高海面时期,说明渤海湾西岸中更新晚期以来地层记录的海面变化趋势与全球变化一致。与前人的传统认识不同,认为受全球海面及渤海海峡地貌共同影响,MIS3时期研究区并未发生大范围海侵。通过多个钻孔的对比分析,3个海相地层在渤海湾西岸地区的分布较广,具有良好的可对比性,不同钻孔之间同一海相地层的高程和厚度的较大差异指示本地区在晚更新世期间发生较强的差异性沉降活动。  相似文献   

15.
We present the results of a palaeogenetic analysis of two Late Pleistocene camelids originating in southern Chile. Our analysis of two mitochondrial DNA fragments (control region and cytochrome b gene) reveals that these specimens do not belong to an extinct taxon, but rather to extant vicuña (Vicugna vicugna), whose modern distribution is restricted to the extreme elevations of the Andes, more than 3500 km to the north of where these specimens originated. Our results also suggest fossil specimens from Patagonia that are currently assigned to the extinct taxon Lama gracilis, may actually belong to V. vicugna, implying a continuous distribution of the latter from the southern tip of South America to the Andes during the Final Pleistocene. The haplotypes of both specimens are not present in modern populations, suggesting a loss of genetic diversity concomitant with the contraction of the vicuña geographical distribution during the Final Pleistocene or early Holocene.  相似文献   

16.
Studies of invertebrates from steppe patches in the tundra and taiga zones of Beringia provide additional evidence that these areas could be relict steppes. A number of insect species common to both modern relict steppes and fossil Beringian insect faunal assemblages have been found. These provide important information on the moisture and temperature preferences of some of the surviving members of Pleistocene steppe-tundra insect communities. The most significant species of West Beringian insects are weevils in the genus Stephanocleonus (Coleoptera, Curculionidae), indicators of thermophytic steppe, and the pill beetle Morychus viridis (Coleoptera, Byrrhidae), the indicator of hemicryophytic steppe. The East Beringian invertebrate population of relict steppe is substantially different. Fossil evidence suggests that biotic exchange between the two parts of Beringia was limited during the Pleistocene; populations of steppe insects did not move across the Bering Land Bridge (BLB), while tundra species had more flexibility. The tundra environment reconstructed for the Pleistocene BLB should have facilitated amphi-beringian distributions for most tundra invertebrate species, but apparently only a few species achieved this.  相似文献   

17.
A pollen record from a sediment core drilled underneath the Acheulian site of Fontana Ranuccio provides new data on the vegetation characters in central Italy during the late Early Pleistocene, in a forest phase possibly corresponding to a substage of Marine Isotope Stage 21. A number of tree taxa nowadays extinct in Europe (Cathaya, Tsuga, Taxodium type, Carya and Pterocarya) are present at Fontana Ranuccio in a temperate phase characterised by dense forest conditions, dominated by Alnus and Pterocarya. The comparison of the Fontana Ranuccio record with other dated sites distributed along the Italian peninsula indicates that the floristic and vegetational characters of the Mid Pleistocene revolution present a rather distinct succession of biostratigraphic events. Compared to other Early Pleistocene pollen records from southern Europe, the Italian Peninsula shows greater floristic similarities with the Eastern Mediterranean regions than with the Iberian Peninsula. Copyright © 2011 John Wiley & Sons, Ltd.  相似文献   

18.
In this paper, the currently accepted correlation of the Early Pleistocene Ludhamian stage of England with the Tiglian‐A sub‐stage of the Netherlands is challenged. Recent investigations of Early Pleistocene marine North Sea deposits from a borehole near Noordwijk (the Netherlands) yielded evidence from molluscs, dinoflagellate cysts and sporomorphs for an alternation of warm‐temperate and arctic intervals within the Praetiglian and Tiglian stages. Marine equivalents of the terrestrial‐based pollen sub‐stages Tiglian A and B have been recognised in the upper part of the sequence. A Praetiglian age can be assigned to the lower part of the sequence on the basis of mollusc analysis. Within the Praetiglian, an alternation of warm and cold phases has been recognised from both the dinoflagellate cyst and molluscan records. Three cold phases within the Praetiglian are tentatively correlated with marine isotope stages (MIS) 96–100. The molluscan assemblages provide evidence for climate forcing of the sea level: highest sea levels are reached in the warm‐temperate intervals. Within the Praetiglian, an interval with an acme zone of the dinoflagellate cyst Impagidinium multiplexum, is correlated with the Ludhamian and tentatively linked to MIS 97 and/or MIS 96. The cold molluscan assemblages from the Noordwijk borehole include an acme zone of Megayoldia thraciaeformis, the first and only occurrence of this North Pacific bivalve in the North Sea Basin. Copyright © 2006 John Wiley & Sons, Ltd.  相似文献   

19.
Although a very high invertebrate faunal diversity is known from the outcrops of the Ariyalur group in the Cauvery Basin, southern India, little is known about its vertebrate fauna. Recent fieldwork in the badland exposures of the Karai Formation (Upper Cenomanian-Lower Turonian) near Garudamangalam in the basin has yielded two teeth belonging to the Late Cretaceous shark Ptychodus decurrens (Ptychodontidae). The fossil record of Ptychodus decurrens from the southern continents is very poor, being known from a single Late/Middle Albian occurrence in Australia. This finding documents the first record of fossil P. decurrens in India and second from a Gondwanan landmass, and provides the first evidence of a cosmopolitan, Pangaean, distribution of the species during the Albian-Turonian and additional insights into the palaeoecology of the Cauvery Basin during the deposition of the Karai Formation.  相似文献   

20.
Over the last 50 years the studies on terrestrial mammals of the Italian peninsula have provided a large volume of data and a more detailed knowledge of faunal events during the Late Pliocene and Quaternary. Moreover geological, sedimentological, palynological and magnetostratigraphical investigations on the Pliocene–Pleistocene continental sedimentary basins have yielded the possibility of a detailed calibration of the faunal successions. Thus, palaeontologists have been able to reconstruct faunal sequences and to propose biochronological scales based on large and small mammals, respectively. In the present contribution an integration of the two biochronological scales is proposed, and the successions of bioevents are carefully compared. This integrated approach allows the constraint of the sequence of large- and small-mammal events in a more reliable way, and therefore it results in a more detailed and consistent chronological use of mammalian assemblages. Particular attention is paid to the faunal changes that correspond to the Middle–Late Pliocene (2.6 ma), Pliocene–Pleistocene (1.8 ma) and Early–Middle Pleistocene (Gauss–Matuyama transition) chronostratigraphical boundaries.  相似文献   

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