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1.
Hydro-acoustic surveys have been used to provide annual estimates of May recruitment and November spawner biomass of the South African sardine Sardinops sagax and anchovy Engraulis encrasicolus resources since 1984. These time-series of abundance estimates form the backbone of the assessment of these resources, and consequently the management of the South African sardine and anchovy is critically dependent on them. Upgrades to survey equipment over time have resulted in recent surveys providing more accurate estimates of abundance, yet in order to maintain comparability across the full time-series, estimates of biomass mimicking the old equipment were used for a number of years. In this paper we develop a method to revise the earlier part of the time-series to correct for receiver saturation in the older generation SIMRAD EK400 and EKS-38 echo sounders and to account for attenuation in dense sardine schools. This is applied to provide a revised time-series of biomass estimates for the South African sardine and anchovy resources with associated variance–covariance matrices. Furthermore, the time-series presented here are based on updated acoustic target strength estimates, making this the most reliable time-series currently available for both resources.  相似文献   

2.
The diet of sardine Sardinops sagax in the southern Benguela was investigated by microscopic examination of stomach contents. The relative dietary importance of prey size and prey type was assessed by calculating the carbon content of prey items. Sardine is an omnivorous clupeoid, ingesting both phytoplankton and zooplankton, with the relative importance of these two food types varying both spatially and temporally. Stomach contents were numerically dominated by small prey items, principally dinoflagellates, followed by crustacean eggs, cyclopoid copepods, calanoid copepods and diatoms. Virtually all prey items ingested by sardine were <1.2 mm maximum dimension, the particle size below which sardine only filter-feed. Despite the numerical dominance by phytoplankton, zooplankton contributed the major portion to sardine dietary carbon, small calanoid and cyclopoid copepods, anchovy eggs and crustacean eggs being the primary prey types. These results indicate that, like anchovy Engraulis capensis, sardine in the southern Benguela are primarily zoophagous, and contrast with earlier dietary studies on sardine in the region. However, the two species appear to partition their prey on the basis of size; sardine consume small zooplankton, whereas anchovy consume large zooplankton. This difference has been observed in other upwelling ecosystems where the two genera co-exist and is likely to contribute to the regime shifts observed between sardine and anchovy.  相似文献   

3.
The occurrence and distribution of eggs and larvae of anchovy, Engraulis japonicus, in Jeju Strait, Korea from May to November 2002 were investigated, and the relationships between the abundances of anchovy eggs and larvae and the oceanographic conditions, including meteorological data, were analyzed. During the sampling periods, the anchovy eggs occurred from May to October and about 95% of total egg abundance occurred in only two months from July to August, suggesting that the main spawning season of anchovies is confirmed in summer from July and August in Jeju Strait. The spawning area of anchovies gradually moved to the southern part of the survey area with time. The predominant anchovy larvae collected in this study were mostly smaller than 16.0 mm TL. Correlation analysis and multiple regression analysis showed that the anchovy egg abundance in the study area depended strongly on the wind conditions, especially less than 10 m/s, and increasing water temperature. Salinity variations also had significant effects on the abundance of anchovy larvae.  相似文献   

4.
The present paper investigates the effect of environmental variability on the spawning dynamics of sardine, Sardina pilchardus, in the Eastern Mediterranean. To do so, variability in four different reproductive traits, i.e. reproductive activity, incidence of recent spawning, prevalence of early ovarian atresia and relative batch fecundity was tested along a gradient of habitat conditions like water temperature, chlorophyll a (Chl a) concentration and an index of zooplankton production. Apart from atretic prevalence all remaining traits were significantly and positively correlated with Chl a concentration which indicates that sardine in the E. Mediterranean tends to enhance egg production under conditions of increased primary productivity. On the other hand, prevalence of atresia together with spawning incidence were positively affected by water temperature whilst the only factor that correlated significantly with the index of zooplankton production was relative fecundity. These findings were interpreted as an indication of direct energy flow from phytoplankton filter-feeding to gonadal development and egg production, implying that besides capitalized energy sardine also uses current income for supporting reproduction.  相似文献   

5.
This study attempts to explain the variability in recruitment of sardine in the northern Benguela and to develop potential models by including environmental information to predict recruitment. Two different recruitment and spawner number datasets were available: a VPA-developed dataset, for the period 1952-1987, and data from a simple age-structured model for 1992-2007. In all, four environmental indices were used: the degree of the intrusion of the warm Angola Current into northern Namibia, termed the Angola-Benguela front index; the extent of the upwelling area off central Namibia; average sea surface temperature (SST) over the northern and central Namibian shelf; and wind stress anomalies at Luderitz as an indicator of upwelling strength. Contrary to general belief, it was found that extremely high recruitment can happen at low spawner levels. This occurred in years in which a large upwelling area existed in association with the minimum southward intrusion of the Angola Current. These effects override the normal negative linear relationships with SST and the positive linear relationship with wind. However, when the area of upwelling is average or small, the effects of spawner biomass, SST and wind become important factors in the variability of recruitment. To estimate exceptional recruitment, the upwelling and front indices were included in the model. To measure medium and weak recruitment, spawner numbers and the SST and wind anomaly formed part of the model. These models can be used simultaneously to predict recruitment before annual acoustic surveys take place and thus aid management decisions.  相似文献   

6.
The Humboldt Current System, like all upwelling systems, has dramatic quantities of plankton-feeding fish, which suggested that their population dynamics may ‘drive’ or ‘control’ ecosystem dynamics. With this in mind we analysed the relationship between forage fish populations and their main prey, zooplankton populations. Our study combined a zooplankton sampling program (1961–2005) with simultaneous acoustic observations on fish from 40 pelagic surveys (1983–2005) conducted by the Peruvian Marine Research Institute (IMARPE) and landing statistics for anchoveta (Engraulis ringens) and sardine (Sardinops sagax) along the Peruvian coast from 1961 to 2005. The multi-year trend of anchoveta population abundance varied consistently with zooplankton biovolume trend, suggesting bottom-up control on anchovy at the population scale (since oceanographic conditions and phytoplankton production support the changes in zooplankton abundance). For a finer-scale analysis (km) we statistically modelled zooplankton biovolume as a function of geographical (latitude and distance from the 200-m isobath), environmental (sea surface temperature), temporal (year, month and time-of-day) and biological (acoustic anchovy and sardine biomass within 5 km of each zooplankton sample) covariates over all survey using both classification and regression trees (CART) and generalized additive models (GAM). CART showed local anchoveta density to have the strongest effect on zooplankton biovolume, with significantly reduced levels of biovolume for higher neighbourhood anchoveta biomass. Additionally, zooplankton biovolume was higher offshore than on the shelf. GAM results corroborated the CART findings, also showing a clear diel effect on zooplankton biovolume, probably due to diel migration or daytime net avoidance. Apparently, the observed multi-year population scale bottom-up control is not inconsistent with local depletion of zooplankton when anchoveta are locally abundant, since the latter effect was observed over a wide range of overall anchoveta abundance.  相似文献   

7.
Regime shifts in the Humboldt Current ecosystem   总被引：4，自引：0，他引：4  
Of the four major eastern boundary currents, the Humboldt Current (HC) stands out because it is extremely productive, dominated by anchovy dynamics and subject to frequent direct environmental perturbations of the El Niño Southern Oscillation (ENSO). The long-term dynamics of the HC ecosystem are controlled by shifts between alternating anchovy and sardine regimes that restructure the entire ecosystem from phytoplankton to the top predators. These regime shifts are caused by lasting periods of warm or cold temperature anomalies related to the approach or retreat of warm subtropical oceanic waters to the coast of Peru and Chile. Phases with mainly negative temperature anomalies parallel anchovy regimes (1950–1970; 1985 to the present) and the rather warm period from 1970 to 1985 was characterized by sardine dominance. The transition periods (turning points) from one regime to the other were 1968–1970 and 1984–1986. Like an El Nino, the warm periods drastically change trophic relationships in the entire HC ecosystem, exposing the Peruvian anchovy to a multitude of adverse conditions. Positive temperature anomalies off Peru drive the anchovy population close to the coast as the coastal upwelling cells usually offer the coolest environment, thereby substantially decreasing the extent of the areas of anchovy distribution and spawning. This enhances the effects of negative density-dependent processes such as egg and larval cannibalism and dramatically increases its catchability. Increased spatial overlap between anchovies and the warmer water preferring sardines intensifies anchovy egg mortality further as sardines feed heavily on anchovy eggs.Food sources for juvenile and adult anchovies which prey on a mixed diet of phyto- and zooplankton are drastically reduced because of decreased plankton production due to restricted upwelling in warm years, as demonstrated by lower zooplankton and phytoplankton volumes and the diminution of the fraction of large copepods, their main food source.Horse mackerel and mackerel, the main predators of anchovy, increase predation pressure on juvenile and adult anchovies due to extended invasion into the anchovy habitat in warmer years. In contrast to these periods of warm and cold temperature anomalies on the decadal scale, ENSO events do not play an important role for long-term anchovy dynamics, as the anchovy can recover even from strong ENSO events within 1–2 years. Consequently, the strong 1972–1973 ENSO event (in combination with overfishing) was not the cause of the famous crash of the Peruvian anchovy fishery in the 1970s.  相似文献   

8.
The Humboldt Current System (HCS) is dominated by two pelagic species; Peruvian anchovy or anchoveta (Engraulis ringens) and sardine (Sardinops sagax). Using data from 43 acoustic surveys conducted from 1983 through 2005 by the Peruvian Marine Institute (IMARPE), we examined the distribution of these two species relative to water masses. We tested the hypothesis that anchovy was found more frequently in upwelled cold coastal water (CCW) and mixed waters (MCW) than in other water types and that sardine was more associated with more offshore oceanic surface subtropical water (SSW). Surface temperature, salinity, latitude, season and distance to the coast data were used to define water masses. Results using generalized additive models (GAM), modelling sardine and anchovy presence–absence as a function of year, water body, bottom depth and latitude, showed that anchovy were primarily found in CCW and MCS, while sardine were more ubiquitous relative to water masses with some predilection for SSW. These results were supported by various indexes of anchovy and sardine distribution versus water mass as well as temporal and location variables.  相似文献   

9.
The recruitment rate of Peruvian anchoveta, Engraulis ringens, was studied to test the hypothesis that long-term environmental variation (regime shifts) had a significant impact on density-dependent processes governing the anchovy recruitment during the period 1963–2004. On the basis of previous defined regimes and turning points for the Humboldt Current System, we identified two groups of years for increased recruitment of anchoveta (1963–1971 and 1986–2004), and one unfavorable period (1972–1985). A common intercept and significantly different slopes were found when the recruitment rate was plotted as a function of the spawning stock biomass during those groups of years, suggesting that density-dependent effects on recruitment were affected during different climate regimes. The favorable (unfavorable) regime was characterized by higher (lower) zooplankton volumes, and with a higher frequency of colder (warmer) waters. Dome-shaped relationships between recruitment rate, spawning stock biomass and SST, were detected with a Generalized Additive Model for the favorable regime. Thus, recruitment could be explained by non-linear effects of environmental variables. Ultimately, climatic regimes are affecting the density-dependent effects on recruitment of anchoveta and the mechanisms involved may be associated with changes in the carrying capacity of the spawning habitat of anchoveta off Peru, which in turn are related with the effects of cold and warm regimes.  相似文献   

10.
Despite much public awareness surrounding the annual migration of sardine Sardinops sagax northward along the east coast of South Africa in winter each year, relatively little research effort has been expended to improve understanding of the ‘sardine run’. For this reason, a dedicated multidisciplinary survey, timed to coincide with the annual sardine run, was conducted off the East Coast in June and July of 2005. The major objective of the survey was to estimate the biomass of sardine off the East Coast during the run, and to compare this with biomass estimates collected during previous surveys conducted in this area during the late 1980s when the South African sardine population was at a considerably smaller size. We also collected data on the distribution of sardine and other small pelagic fish species and their eggs, the biological characteristics of sardine during the run, and data on the hydrography (temperature and currents) and lower trophic levels (phytoplankton and zooplankton) of the region. Results suggest that the biomass of sardine off the East Coast in winter remains relatively small and consistent, regardless of overall sardine population size. The narrow continental shelf to the east of Port Alfred, which is dominated offshore by the fast-flowing warm Agulhas Current, constrains the amount of suitable habitat for sardine and other clupeoids such as anchovy Engraulis encrasicolus, West Coast round herring Etrumeus whiteheadi and East Coast round herring Etrumeus teres, and hence precludes these species from attaining a high biomass in this region. Additionally, primary and secondary productivity levels are much lower than elsewhere on the western and eastern Agulhas Bank off the south coast of South Africa, suggesting that the sardine run is not a feeding migration. A previous hypothesis that the run is mainly a result of an expansion of the distributional range of these fish as conditions become favourable in winter due to sporadic cooling off the East Coast is also not entirely supported by results from the survey. It is suggested that a migration for the purposes of spawning off this coast when conditions become favourable is a more likely incentive for sardine to undertake this arduous journey, despite increased predation and poor feeding conditions.  相似文献   

11.
Trends in the spatial pattern of spawning of pilchard Sardinops ocellatus and anchovy Engraulis capensis during the period 1971–1985 were examined in relation to large-scale environmental anomalies outside the spawning area. Shifts in the locality of peak spawning of anchovy often coincided with that of pilchard, although sometimes anchovy were slower to respond to an environmental change. With negative sea surface temperature (SST) anomalies spawning shifted northwards, whereas with positive anomalies spawning took place both in the north and farther south. With a transition from negative SST anomalies early in the spawning season to zero or positive anomalies later in the spawning season, the impact on anchovy and pilchard spawning differed. Anchovy spawning was either reduced or absent during the negative phase of transitional seasons, thus avoiding unfavourable conditions. Pilchard, on the other hand, were spawning mainly in the north during the negative phase of the transitional seasons, but also in the south during the zero to positive phase. The shifts in spawning locality in response to environmental anomalies outside the spawning area imply that explanations for such shifts need not be sought in changes in biomass or age structure or in the fact that stocks may be genetically different, all theories previously advanced.  相似文献   

12.
Potential spawning habitat is defined as the area where environmental conditions are suitable for spawning to occur. Spawning adult data from the first quarter (January–March) of the International Bottom Trawl Survey have been used to study the inter-annual variability of the potential spawning habitat of North Sea plaice from 1980 to 2007. Generalised additive models (GAM) were used to create a model that related five environmental variables (depth, bottom temperature and salinity, seabed stress and sediment type) to presence–absence and abundance of spawning adults. Then, the habitat model was applied each year from 1970 to 2007 to predict inter-annual variability of the potential spawning habitat. Predicted responses obtained by GAM for each year were mapped using kriging. A hierarchical classification associated with a correspondence analysis was performed to cluster spawning suitable areas and to determine how they evolved across years. The potential spawning habitat was consistent with historical spawning ground locations described in the literature from eggs surveys. It was also found that the potential spawning habitat varied across years. Suitable areas were located in the southern part of the North Sea and along the eastern coast of England and Scotland in the eighties; they expanded further north from the nineties. Annual survey distributions did not show such northward expansion and remained located in the southern North Sea. This suggests that this species' actual spatial distribution remains stable against changing environmental conditions, and that the potential spawning habitat is not fully occupied. Changes in environmental conditions appear to remain within plaice environmental ranges, meaning that other factors may control the spatial distribution of plaice spawning habitat.  相似文献   

13.
Potential spawning habitat is defined as the area where environmental conditions are suitable for spawning to occur. Spawning adult data from the first quarter (January–March) of the International Bottom Trawl Survey have been used to study the inter-annual variability of the potential spawning habitat of North Sea plaice from 1980 to 2007. Generalised additive models (GAM) were used to create a model that related five environmental variables (depth, bottom temperature and salinity, seabed stress and sediment type) to presence–absence and abundance of spawning adults. Then, the habitat model was applied each year from 1970 to 2007 to predict inter-annual variability of the potential spawning habitat. Predicted responses obtained by GAM for each year were mapped using kriging. A hierarchical classification associated with a correspondence analysis was performed to cluster spawning suitable areas and to determine how they evolved across years. The potential spawning habitat was consistent with historical spawning ground locations described in the literature from eggs surveys. It was also found that the potential spawning habitat varied across years. Suitable areas were located in the southern part of the North Sea and along the eastern coast of England and Scotland in the eighties; they expanded further north from the nineties. Annual survey distributions did not show such northward expansion and remained located in the southern North Sea. This suggests that this species' actual spatial distribution remains stable against changing environmental conditions, and that the potential spawning habitat is not fully occupied. Changes in environmental conditions appear to remain within plaice environmental ranges, meaning that other factors may control the spatial distribution of plaice spawning habitat.  相似文献   

14.
The annual movement of South African sardine Sardinops sagax up the east coast of South Africa, known as the ‘sardine run’, was investigated using data from aerial surveys for the period 1988–2005 and compared with remotely sensed sea surface temperature (SST) and chlorophyll a data. Sardine sighting rates were highest within the Waterfall Bluff Bight off the Eastern Cape Coast, where conditions appeared to be most favourable. Sardine and predator sightings decreased significantly northwards of Mdoni on the KwaZulu-Natal (KZN) coast, whereas the proportion of nearshore sightings increased. The causal mechanism for this inshore concentration is suggested to be the influx of warm Agulhas Current water from the Durban Eddy that forces sardine shoreward. Cape gannet Morus capensis, common dolphin Delphinus capensis and sardine distributions were associated, and there was an association between SST and sardine and predator distributions. There was a marked increase in bottlenose dolphin Tursiops aduncus sightings upon commencement of the sardine run, with these dolphins being consid-ered to be a ‘migratory’ stock that enters KZN waters every winter.  相似文献   

15.
Climate variability and pelagic fisheries in northern Chile   总被引：5，自引：0，他引：5  
A time series analysis of long-term climate variability in northern Chile (18°21′–24°00′S) shows anomalies associated with the El Niño events and the longer warm period observed since 1976, followed by a cooling trend since mid 1980s. The succession of pelagic fisheries, anchovy (Engraulis ringens) and sardine (Sardinops sagax), occurring in this fishing zone was analyzed taking into account the landings, the CPUE abundance index, the fishing effort, and the environmental variables. The anchovy production model is a negative linear function of fishing effort and turbulence. For sardine, the production model is a negative linear function of fishing effort and a quadratic function of the sea surface temperature.An analysis of the relationship between recruitment, adult biomass and the environment shows that the annual recruitment of anchovy increases with turbulence intensity until wind speed reaches a value of 5.46 m s−1, decreasing for higher values. For sardine, the recruitment increases with turbulence intensity until 5.63 m s−1, stabilizing thereafter.It is deduced that the climatic variations associated to the El Niño events affect the abundance of coastal pelagic fishes, without forgetting the most likely effects upon its distribution and the fishing effort. However, it is the long-term variability that mainly affects the fishing activity.  相似文献   

16.
福清湾及附近海域浮游动物的数量和分布   总被引：3，自引：0，他引：3  
林君卓 《台湾海峡》2008,27(1):58-63
本文分析了2005年10月和2006年5月在福清湾及其附近海域采集的浮游动物样品,结果表明,福清湾已鉴定的浮游动物有64种,10类浮游幼体.分为4个生态类群,包括河口低盐类群,近岸暖温类群,近岸暖水类群和广布外海类群.浮游动物物种多样性指数春季高于秋季.春季浮游动物总个体数均值为(161ind/m3)高于秋季的(83ind/m3),春季生物量均值为(67.7mg/m3)低于秋季的(87.7mg/m3);浮游动物总个体数的平面分布,春季湾内西部水域最高,海坛海峡最少;秋季福清湾东部水域最高,湾内西部水域最少.浮游动物生物量和总个体数的平面分布趋势基本一致.另外还对浮游动物的分布与环境因子的关系进行了分析.  相似文献   

17.
Marine oxygen-deficient environments with high sedimentation rates and high primary productivity can provide relevant information regarding variations of ocean–climatic conditions in the past. In the Humboldt current ecosystem, which now hosts huge populations of pelagic fishes (mainly anchovy and sardine), fish scale abundance in the sedimentary record may be useful indicators of environmental change. Here we assess such a proxy record in a 42 cm-long sedimentary core collected from 80 m in Mejillones Bay (23°S, northern Chile). We also analyse fish remains in surface sediment sampled along a bathymetric transect (from 10 to 110 m water depth) in the same bay. In the core-top record, the fluctuations of sardine and anchovy scale deposition rates (SDR) agreed with those of industrial catches for these two species in northern Chile, tending to validate the SDR as a proxy of local fish biomass when bottom anoxic conditions prevail. However, apparent SDR for records prior to 1820 have probably been influenced by dissolution processes linked to the oxygenation of the bottom environment of Mejillones Bay, as suggested by other proxy records. After 1820, the fluctuations in the relative abundance of sardine and anchovy scales point to alternating warm and cold conditions during about 30 years and then a progressively cooler period. Since ca. 1870, marked fluctuations of SDR of both species are observed, probably as a consequence of the onset of a different oceanographic regime characterized by intensified upwelling, stronger subsurface oxygen deficiency, higher primary productivity, and enhanced “ENSO-like” interdecadal variability. While anchovy SDR fluctuated in periods of 25–40 years, only two peak periods of sardine SDR occurred (late 19th century and late 20th century), suggesting that sardine abundance depends on other ocean–climatic factors.  相似文献   

18.
Cape anchovy Engraulis encrasicolus spawners in the southern Benguela showed an eastward shift in their distribution on the Agulhas Bank that occurred abruptly in 1996 and has since persisted. We assessed whether this shift was environmentally mediated by examining sea surface temperature data from different regions of the Agulhas Bank, which showed that in 1996 the inner shelf of the Agulhas Bank to the east of Cape Agulhas abruptly became 0.5°C colder than in previous years and has since remained that way. In addition, signals, coherent with the 1996 shift recorded in sea surface temperatures, were also found in atmospheric surface pressure and zonal wind data for that region; interannual coastal SST variability is also shown to be correlated with zonal wind-stress forcing. As a result, increased wind-induced coastal upwelling east of Cape Agulhas is proposed as the main driver of the observed cooling in the coastal region. The synchrony between the environmental and biological signals suggests that the eastward shift in anchovy spawner distribution was environmentally mediated and arose from a change in environmental forcing that altered the relative favourability for spawning between regions to the west and east of Cape Agulhas. The results highlight how a relatively minor change in environmental conditions can lead to a drastic spatial reorganisation of the life history of one species in an ecosystem.  相似文献   

19.
Swordtip squid (Uroteuthis edulis) is one of the important economical fishing target species in the East China Sea. Uroteuthis edulis is characterized by rapid growth, extensive migration, and long spawning period and sensitive to surrounding environment. In order to assess its stock status, it is necessary to explore its spawning season, growth patterns of different populations and their relationship with the environment in advance. In this paper, based on the samples of U. edulis collected in the East China Sea from September 2017 to March 2018, we explored the relationships between daily growth of statolith microstructure and environmental variables by gradient forest method and generalized additive model. The spawning season of U. edulis was found to be nearly one year, and two dominant season groups were found: the spring group with the peak period of April and the summer group with the peak period of August. Water temperature in the depth of 25 m (Temp_25), sea surface temperature (SST) and zonal velocity were the key environmental variables for the daily growth of April-spawning group. The most suitable environmental conditions for the growth of April-spawning group were water temperature (24?27°C) and velocity (0.1?0.3 m/s). SST, Temp_25 and mixed layer depth were the key environment variables for the daily growth of August-spawning group. The most suitable environmental conditions for the growth of August-spawning group were water temperature (21?28°C) and water depth (0?50 m). Key environmental variables of different groups suggested that early growth was significantly affected by seasonal changes of water temperature, current velocity and prey abundance. This study explored the relationships between early growth and environmental variables and provided the scientific guidance for the management and conservation of U. edulis.  相似文献   

20.
于2009年4月15～23日在长江口及其邻近海域进行了32个站位的浮游动物生态调查,分析了浮游动物的种类组成、优势种、丰度和生物量的空间分布,并利用PRIMER生物统计学软件中的PCA、BIOENV和RELATE程序分析了浮游动物与环境因子的关系.共发现浮游动物成体72种,浮游幼虫9种,合计种类数为81.中华哲水蚤(C...  相似文献   

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