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1.
The Lower Cretaceous (Albian) Sasayama Group in the Hyogo Prefecture of southwestern Japan has yielded various vertebrate fossils, including skeletal remains of dinosaurs, anurans, lizards, and mammals, and recently eggshell fragments. Here we report on numerous fossil eggshells from the bone-bearing Kamitaki locality in Tamba City, which represents a diverse dinosaur eggshell assemblage. Of the more than 90 eggshell fragments recovered, five different types were identified, including eggshells that likely belong to a variety of theropods (Nipponoolithus ramosus oogen. et oosp. nov., Elongatoolithus sp., Prismatoolithus sp., and Prismatoolithidae indet.) and at least one ornithopod (Spheroolithus sp.). All eggshells are relatively thin, and a new derived estimation method correlating egg mass with eggshell thickness indicates that they are among the smallest (28–135 g) theropod eggs known, likely laid by small bodied forms. The eggshell assemblage from this locality suggests that a diverse small dinosaur fauna, consisting primarily of theropods, nested in the region, a diversity yet to be evidenced from skeletal remains in Japan.  相似文献   

2.
汝阳盆地是豫西地区的一个中—新生代断陷盆地,自2006年以来发现了一个以巨型蜥脚类恐龙为代表的恐龙动物群。本文描述了近期发现于汝阳盆地下白垩统郝岭组中的若干恐龙蛋蛋壳碎片。基于以下对蛋壳宏观形态和显微特征的分析,这些蛋壳碎片可以归入长形蛋科(Elongatoolithidae)的长形蛋属(Elongatoolithus):蛋壳外表面具分散瘤点状或线性脊状的纹饰;蛋壳自内而外分为锥体层和柱状层,二层之间的界线不清晰;锥体层中锥体排列紧密;柱状层中具波浪状的平行生长纹。这一发现代表了长形蛋属在早白垩世地层中的首次记录,进一步拓展了该蛋属的地史和古地理分布。由以往已知的化石关联推测,这些蛋壳可能与该层位中的窃蛋龙类如刘店洛阳龙(Luoyanggia liudianensis)等具有相关性,表明汝阳盆地在早白垩世晚期具有适宜该类恐龙生存与繁殖的环境。  相似文献   

3.
The North American fossil record of dinosaur eggshells for the Cretaceous is primarily restricted to formations of the middle (Albian–Cenomanian) and uppermost (Campanian–Maastrichtian) stages, with a large gap in the record for intermediate stages. Here we describe a dinosaur eggshell assemblage from a formation that represents an intermediate and poorly fossiliferous stage of the Upper Cretaceous, the Santonian Milk River Formation of southern Alberta, Canada. The Milk River eggshell assemblage contains five eggshell taxa: Continuoolithus, Porituberoolithus, Prismatoolithus, Spheroolithus, and Triprismatoolithus. These ootaxa are most similar to those reported from younger Campanian–Maastrichtian formations of the northern Western Interior than they are to ootaxa reported from older middle Cretaceous formations (i.e., predominantly Macroelongatoolithus). Characteristics of the Milk River ootaxa indicate that they are ascribable to at least one ornithopod and four small theropod species. The taxonomic affinity of the eggshell assemblage is consistent with the dinosaur fauna known based on isolated teeth and fragmentary skeletal remains from the formation, although most ornithischians and large theropods are not represented by eggshell. Relative to the Milk River Formation eggshell, similar oospecies occurring in younger Cretaceous deposits tend to be somewhat thicker, which may reflect an increase in body size of various dinosaur lineages during the Late Cretaceous.  相似文献   

4.
巨齿兰州龙(新属、新种)是发现于中国甘肃省兰州盆地早白垩世地层中的一新鸟脚类恐龙,其牙齿是世界上已知植食性恐龙中最大的。兰州龙下颌长1m,每侧有14个齿槽,而单个牙齿宽约4cm,这是已知恐龙中的首次报道。分支系统学分析发现巨齿兰州龙与非洲早白垩世Lurdusaurus arenatus关系密切。它们代表了鸟脚类恐龙进化中四足行走的笨重的一新支。兰州龙的发现也表明欧亚大陆与非洲在早白垩世具有较密切的联系。  相似文献   

5.
《China Geology》2018,1(1):28-35
An alvarezsaurid dinosaur skeleton was discovered from the Late Cretaceous Qiupa Formation of Luanchuan, Henan Province of central China. It represents a new alvarezsaurid dinosaur Qiupanykus zhangi gen. et sp. nov. A phylogenetic analysis recovers Qiupanykus nested within the unresolved clade, which includes Asian and north American taxa. The skeleton of the new specimen is preserved in association with eggshells. The eggshell morphologies show that these eggs belong to oviraptorid eggs, skeletal remains of which were discovered from the same area. The alvarezsaurid skeleton associated with eggshell fragments may indicate that these eggs were broken by the strong thumb-claws of the former and that alvarezsaurid dinosaurs may be egg-eaters.  相似文献   

6.
Basal iguanodontians (“iguanodontids”) were a successful group of ornithopod dinosaurs that attained a near global distribution by the late Early Cretaceous. Despite their body fossils being known in abundance from the uppermost Jurassic and Lower Cretaceous of the western United States, their remains have never been found in western Canada. With its extensive, terrestrial sedimentary record for the whole of the Early Cretaceous, the expectation is that these dinosaurs would have been present in western Canada. This paper reports the finding of a fossil footprint from the Gladstone Formation in southwestern Alberta that is interpreted to have been made by an iguanodontid. This identification is based on the late Barremian age of the hosting rock, gross footprint details that match the skeletal foot structure of Iguanodon bernissartensis, and similarities to Early Cretaceous dinosaur footprints found elsewhere in the world that are attributed to iguanodontids. This finding fills a noticeable gap in the iguanodontian fossil record, and is consistent with the clade's attainment of global distribution during the Early Cretaceous.  相似文献   

7.
A new ootype collected from the Upper Cretaceous Lijiacun Formation in the Shangdan Basin, Shaanxi Province is described in this paper. Based on general external shape, size, eggshell thickness and honeycomb‐like eggshell microstructure, eggs are referable to the oofamily Faveoloolithidae. Compared with other members of Faveoloolithidae, specimens described in this paper show special characteristics: adjacent pores are usually separated by two eggshell units between which often develop interspaces; columnar eggshell units are relatively closely arranged in radial view. According to these characteristics, we erect a new oogenus and a new oospecies: Duovallumoolithus shangdanensis oogen. et oosp. nov. The new discovery expands the diversity of Faveoloolithidae.  相似文献   

8.
山东临沭地区早白垩世晚期恐龙足迹化石特征*   总被引:1,自引:0,他引:1       下载免费PDF全文
主要记述了山东临沭地区早白垩世晚期(大盛群沉积时期)、生活于干旱炎热气候条件下滨浅湖环境的恐龙所留下的大规模足迹化石群。初步研究表明,这些足迹以蜥脚类恐龙足迹占绝大多数,见少量鸟脚类、兽脚类和疑似甲龙类足迹(甲龙类足迹在中国罕见),足迹总体行进方向以向西和南西为主。通过对其中的2条蜥脚类和1条鸟脚类行迹及其行为学的分析认为其均处于慢行状态,而鸟脚类行迹的运动速度极慢,可能处于一种特殊的慢行状态。临沭地区早白垩世晚期大盛期如此大规模足迹化石群的发现,说明中国乃至东亚地区早白垩世普遍存在着以恐龙为代表的(包括鸟类在内)陆生脊椎动物群。足迹化石的研究将对进一步了解晚中生代陆地生物群的演化有着重要意义。对比其他地区早白垩世晚期地层中发现的恐龙足迹化石特征后认为,足迹的保存与当时的沉积环境有密切的关系,因此对恐龙足迹的研究又能进一步为恢复古生态、古地理环境等提供更丰富的信息。  相似文献   

9.
The Coll de Nargó area (Lleida Province, south-central Pyrenees) has yielded thousands of dinosaur eggs distributed in more than 30 levels across 370 m of Upper Cretaceous Tremp Formation deposits. Two stratigraphic units yielding dinosaur eggshells, eggs and clutches, can be distinguished. Multiple eggshells were collected from up to 75 different in situ clutches, located in consecutive stratigraphical levels throughout the Mas de Pinyes section. Four different oospecies have been recognized: Megaloolithus aureliensis, Megaloolithus siruguei, Megaloolithus cf. baghensis and Cairanoolithus cf. roussetensis. In absence of typical biochronological markers, fossil eggshells may have potential as a tool for dating. Based on these ootaxa, three oozones can be established along the stratigraphic section. The lower assemblage is composed by M. aureliensis, M. siruguei and Cairanoolithus cf. roussetensis; the middle one consists only of M. siruguei, while the upper oozone is characterized by M. cf. baghensis. Results suggest that the age of Coll de Nargó rocks ranges from the latest Campanian to the beginning of the Late Maastrichtian. Also reported is the first evidence of the oogenus Cairanoolithus outside of France, where it was regarded as an endemic ootaxon. The Coll de Nargó area is considered to be one of the most important dinosaur nesting areas in Europe recurrently used by several dinosaurs during a long time span. Finally, the presence of fossilized eggshell membrane in several specimens from Coll de Nargó contributes to understanding the taphonomic process of megaloolithid eggs in this area.  相似文献   

10.
The spotty nature of the terrestrial fossil record for the Mesozoic hinders a more complete understanding of dinosaur diversity. For stegosaurs (Ornithischia), the plated dinosaurs, only a few and fragmentary remains are reported from the Early Cretaceous of Europe. A recent revision concluded that only a partial vertebra of the nomen dubium Craterosaurus (?Aptian, England) could be considered as stegosaurian. Here we report on a stegosaur tooth from the Early Cretaceous (Berriasian) Purbeckian deposits of Cherves-de-Cognac (Charente), southwestern France. This tiny tooth was examined in detail using microtomography. Comparisons being limited by the rarity of stegosaur tooth rows material (e.g., from the skull of the holotype of Stegosaurus stenops) and dental material, notably from Europe, we observed new material of cf. Stegosaurus armatus and Hesperosaurus mjosi from the Upper Jurassic Morrison Formation of Wyoming (USA). The tooth shows the most similarities to the Late Jurassic genera Stegosaurus and Hesperosaurus, but differs in having a distinctive downwardly arched (V-shaped) cingulum on the ?lingual face (maxillary tooth hypothesis). It is referred to as Stegosauria indeterminate, a medium-sized quadrupedal herbivore that inhabited an emerged land between the Armorican Massif and the Massif Central. This finding is the first evidence of a stegosaur from the Early Cretaceous of France and a welcome addition to the meagre European record of that time. In addition, it is the second stegosaurian tooth crown reported from Europe. The assemblage of ornithischians of Cherves-de-Cognac shares some similarities with that of the Early Cretaceous (Berriasian) of the Purbeck Limestone Group, southern England. The relative rarity of ornithischian osteological remains in both Purbeckian environments suggests that most of these dinosaurs were mainly inhabitants of inland terrestrial palaeoenvironments.  相似文献   

11.
Eighty two dinosaur trackways were newly discovered in Upper Cretaceous lacustrine deposits on islands in the vicinity of Yeosu, Korea. Most dinosaur tracks occur in marginal lake deposits with polygonal desiccation cracks. The dinosaur tracks at the Yeosu site include 65 ornithopod trackways, 16 theropod trackways and one sauropod trackway. The prevalence of ornithopod tracks and the limited occurrence of sauropod tracks at the Yeosu site evidently reflect decreased sauropod diversity in the Upper Cretaceous. All ornithopod trackways represent bipeds, and most of the ornithopod tracks are similar to Caririchnium from other sites of the Korean peninsula. All fossil wood specimens collected in the study area represent conifers (three species of cupressaceous and two species of taxodiaceous conifers, and a new species) except for one, which is a discotyledon. It is thus inferred that the southwestern part of the Korean Peninsula was primarily covered with mesic forests with taxodiaceous trees during the Late Cretaceous. The K–Ar age of the Yeosu tracksite is determined as 81–65 Ma (Camapnian to Maastrichtian). It indicates that the Yeosu track site contains the last records of dinosaurs living in Asia. Consequently, semi-arid palaeoclimatic conditions, together with a large lake as a persistent water source and rich vegetation of gymnosperm trees as food, resulted in the preservation of abundant dinosaur tracks in the Upper Cretaceous on the Korean Peninsula.  相似文献   

12.
Two closely associated egg types occur at the same locality in the Upper Cretaceous (Maastrichtian) St. Mary River Formation in north central Montana. These specimens represent the first fossil eggs described from this formation. At least fifteen small ovoid eggs or egg portions are scattered through a 25 cm interval of rock. Five significantly larger, round eggs overlie these smaller eggs and are in close proximity to one another on a single bedding plane. The best preserved egg of the smaller size measures 36 mm × 62 mm and exhibits the prismatic, two-layered eggshell structure of a theropod egg. The dispersed distribution and inconsistent angles of these small eggs likely resulted from disturbance by subsequent nesting activity and/or possibly nest predation. At least twelve additional small prismatic eggs also occur at this site. We assign the small eggs as a new oogenus and oospecies, Tetonoolithus nelsoni, within the Prismatoolithidae. The large round eggs measure 130 mm in diameter and the eggshell displays substantial diagenetic alteration. These eggs likely belonged to a hadrosaur due to their similarity in egg size, shape, and eggshell thickness to Maiasaura eggs from the stratigraphically lower Two Medicine Formation. Eggs at different stratigraphic levels at this site indicate that conditions favorable to both dinosaur species persisted for an extended period of time. However, determining whether these dinosaurs occupied the nesting site at the same or different years remains beyond the resolution of the rock record.  相似文献   

13.
在晚白垩世期间,哈萨克斯坦东部的斋桑盆地形成了一个湖盆,一直延续到现在。古斋桑湖中最老的沉积是Tayzhuzgen组,由厚达136m的泥岩、粉砂岩、页岩、石英砂岩、砾岩和淡水石灰岩红层组成。按照当地的地层名称,Tayzhuzgen组包括Manrak、Tayzhuzgen、Aulisbulak、Aktobe、Dysum bay和Kiin Kerish层。在斋桑湖以南的Tayzhuzgen河附近一个地点的Tayzhuzgen组下部,发现了恐龙蛋壳碎片,蛋壳至少包括有两个主要类型:圆形蛋类和长形蛋类。恐龙蛋壳指示了Tayzhuzgen组下部的时代为晚白垩世(可能是马斯特里赫特期)。在本组的较高层位,发现了晚?古新世的植物化石,因此斋桑盆地的白垩系-古近系界线在Tayzhuzgen组内部。在Tayzhuzgen组中,白垩纪与古新世化石层之间的地层间隔只有约50m,由此推测在Tayzhuzgen组内部可能存在一个或多个不整合,或是本组的中部为凝缩沉积,即在白垩系-古近系界线处的沉积速率非常缓慢。Tayzhuzgen组顶部有一个显著的不整合,那里中或晚始新世(Arshantan,Irdinmanhan或Egilian陆生哺乳动物"期")的地层存在于含哺乳动物化石剖面的底部。斋桑盆地的白垩系-古近系界线位于古斋桑湖沉积剖面的下部。这不是一个完整的界线剖面,可能是一个包含有一个或多个不整合和/或一个由慢速沉积造成的凝缩剖面。  相似文献   

14.
记述了产自河南汝阳盆地早白垩世晚期相互关联的禽龙类9个中前部尾椎椎体化石。由于化石只保存不完整的尾椎部分,其属种难以断定。但它的发现说明,汝阳巨型蜥脚类恐龙动物群中,除了植食性的大型蜥脚类恐龙外,也存在大型鸟脚类恐龙——禽龙类。保存的尾椎椎体系列与发现于比利时的早白垩世贝尼萨尔禽龙相比,不论是大小还是形态构造上均极为相似,从而判断这9个连续的尾椎体分别为第7到第15尾椎。进而推断,河南汝阳盆地产出巨型蜥脚类恐龙动物群化石的沉积地层形成于早白垩世晚期。  相似文献   

15.
白垩纪四足动物足印的生物地层学、生物年代学与遗迹相   总被引:3,自引:2,他引:1  
从全球范围来看,白垩纪四足动物的足印多数是非鸟恐龙与鸟类留下的痕迹;少量足印来自翼龙、鳄鱼、龟、哺乳动物和其他四足动物。白垩纪的足迹化石以东亚(尤其是中国和朝鲜)和北美西部的最为人所知。南美(主要是阿根廷和巴西)也有一定数量广泛分布的足迹化石,欧洲、非洲与澳大利亚的白垩纪足迹组合则鲜为人知。以白垩纪四足动物的足印记录为基础,我们对两个全球足印生物年代重新进行了检查。早白垩世生物年代以蜥脚类与鸟脚类的足迹为特征。晚白垩世生物年代中的蜥脚类足迹较少,但是鸭嘴龙、暴龙和角龙的足迹增多了。另外,白垩纪足印化石的记录中记载了许多重要的生物地层学信息,如北美白垩纪中期蜥脚类恐龙的消失,以及白垩纪末恐龙的绝灭。越来越多来自东亚的白垩纪足印记录使我们对更精细的地方性白垩纪足印生物年代学有了初步印象。因此,以地方性四足恐龙(包括鸟类)遗迹属的地层分布为基础,可以识别出三个或四个足印生物年代。种类丰富并具有地方性特色的东亚的白垩纪鸟类动物的遗迹群,可能指示白垩纪时东亚存在着一个独特而繁盛的鸟类动物群。以足印化石为基础的这一假说有待进一步的验证。  相似文献   

16.
安徽齐云山晚白垩世恐龙蛋矿物和元素组成及古环境分析   总被引:1,自引:0,他引:1  
姜琴  何情  郑刘根  胡毅 《岩矿测试》2017,36(4):340-349
安徽齐云山新发现的恐龙蛋化石保存完好,是研究该地区晚白垩世恐龙动物群生活环境及绝灭的关键。本文采用粉晶X射线衍射法(XRD)、电感耦合等离子体发射光谱/质谱法(ICP-OES/MS)分析齐云山恐龙蛋壳、蛋核及围岩的宏观矿物及主量、微量元素组成,以探讨齐云山晚白垩世古环境和古气候特征。结果表明:恐龙蛋壳主要矿物组成为方解石和石英,且蛋内外物质矿物组成相似,主要为石英和长石;而元素组成包括常量元素Ca、Na、Mg、K、Al和Fe及微量元素Sr、Zn、Ba、As、Ti、V、Cr、Co、Ni、Cu、Mn、Pb、La、U和Ir,其中Sr在蛋壳中含量的平均值为6199.3×10~(-6),高于河南西峡及广东南雄大部分恐龙蛋壳中的Sr含量,蛋壳中Ir含量的平均值为59.6×10~(-12),Sr和Ir与背景值相比均有明显异常,分析认为是恐龙与外界进行物质交换时吸收有害元素引起的。而As、Mn和Pb的富集及Zn的不足可能使恐龙机体中毒进而灭绝。围岩中Fe含量的增加表明齐云山地区晚白垩世为干旱炎热的气候条件,与Fe呈正相关性的Ir元素随Fe含量的增加而富集,且越靠近K/T界线,围岩中Sr和Ir元素含量越高,这是导致该地区恐龙动物群绝灭的原因之一。此结论为研究齐云山地区恐龙动物群的生活环境提供了依据。  相似文献   

17.
山东沭河裂谷带由北至南发育8个早白垩世晚期(大盛期)的恐龙足迹群化石点,其中5处为本研究首次报道。沭河裂谷带足迹化石点多,分布广,层位多,足迹数量和造迹恐龙种类丰富。郯城新发现的单个蜥脚类巨型足迹直径近1.0m,可能仅次于前人描述过的兰州西南盐锅峡下白垩统河口群内中国最大的蜥脚类足迹化石。沭河裂谷带的恐龙足迹产出于大盛群田家楼组,地质年代为110~100Ma,属于早白垩世晚期Aptian晚期—Albian期。恐龙足迹一般为凹形印痕,造迹者以蜥脚类恐龙为主,其次为鸟脚类和兽脚类。足迹多保存在发育波痕、泥裂、雨痕等沉积构造的细砂—粉砂岩沉积物层面。足迹和行迹特征显示造迹恐龙为群居和在栖息地漫步行走状态。足迹原位保存特征及赋存地层沉积学研究表明,早白垩世中晚期气候处于由温暖潮湿变得炎热干旱的过渡期,湖泊、三角洲广布,恐龙栖息于湖滨环境。沭河裂谷带早白垩世晚期恐龙足迹群的发现和综合研究进一步证实,中国早白垩世恐龙动物群有着更广阔的地理分布和相似的古地理环境。  相似文献   

18.
The Late Cretaceous deposits of the continental Tremp Formation in the Vallcebre Syncline (South-eastern Pyrenees) provide an extensive egg record of dinosaurs. The parataxonomical study and analysis of multiple eggshell samples, the precise stratigraphical control of several sections and the time calibration of the abundant egg levels enable the establishment of a robust oospecies succession. The successive occurrence of three megaloolithid oospecies (Megaloolithus siruguei-Megaloolithus mamillare-Megaloolithus sp.) in the Early and Late Maastrichtian is well correlated with the magnetic polarity time scale throughout the chron 31. The replacement of Megaloolithus siruguei with Megaloolithus mamillare occurs around the reversal of chrons 31r-31n. A comparison with oospecies successions from Arc basin localities (France) allows the age calibration for such oospecies replacement to be confirmed. This age refinement implies that some of the stated boundaries for the proposed oospecies assemblages may change and that an in-depth revision of the age and magnetic calibration of some south European egg localities is required.  相似文献   

19.
The scarcity of diagnostic skeletal elements in the latest Cretaceous theropod record of the Ibero-Armorican domain (southwestern Europe) prevents to perform accurate phylogenetic, paleobiogeographic, and diversity studies. In contrast, eggs and eggshells of theropod dinosaurs are relatively abundant and well known in this region from which several ootaxa have been described. Here, we describe the first Late Maastrichtian theropod ootaxon (Prismatoolithus trempii oosp. nov.) from SW Europe and demonstrate that oological record can be used as a proxy for assessing diversity of egg-producers and may help to complement their scarce bone record. The performed analyses indicate that the theropod taxa and ootaxa reach their diversity maxima during the Late Campanian and start to decrease near the Campanian–Maastrichtian boundary at both global and regional scales. The oological diversity of theropods in the Ibero-Armorican domain is consistent with the theropod diversity identified at high taxonomic level. Two distinct assemblages of theropod ootaxa can be recognized in the latest Cretaceous of the Ibero-Armorican domain. Their temporal transition can be correlated with other dinosaur faunal changes recorded in the region. This faunal turnover took place around the Early–Late Maastrichtian boundary, involving ornithopods, sauropods, ankylosaurs and, according to the present results, theropods as well.  相似文献   

20.
吕君昌 《地球学报》2016,37(2):129-140
在江西赣州地区晚白垩世地层中发现的窃蛋龙类化石——赣州华南龙,属于进步窃蛋龙类,其独特的头部特征为研究窃蛋龙类恐龙的颅面演化、古地理分布及古生态环境提供重要信息;在辽西早白垩世义县组地层中发现的大型、前肢短的驰龙类恐龙——孙氏振元龙,为大型、短前肢类型的驰龙类提供羽毛形态学方面的重要信息,为研究驰龙类的多样性、鸟类羽毛以及飞行起源提供了重要依据;在辽西发现的喙嘴龙类翼龙——朝阳东方颌翼龙,是该地区晚侏罗世地层中发现的第一件翼龙标本,不仅填补了时代上的空白,同时对于印证晚侏罗世喙嘴龙类的辐射演化具有重要作用。这些重要发现对于研究窃蛋龙类的演化、驰龙类羽毛演化及鸟类羽毛起源等热点与难点问题提供了重要的参考依据,尤其在发现华南龙的基础上,命名了赣州恐龙动物群。  相似文献   

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