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1.
Inoceramid bivalves of the upper Albian and lower Cenomanian of the United States Western Interior are revised, Eleven species-level taxa and three genera are described. Two new species, Gnesioceramus mowriensis, characterizing the Mowry Shale of the early, but not the earliest, Cenomanian, and Posidonioceramus merewetheri, of the lower Cenomanian, and on new genus, Posidonioceramus, are recognised. The Western Interior inoceramid species from this interval are strongly endemic and are not good tools for long-distance correlations, although they are very effective in regional dating.In terms of the inoceramid biostratigraphy, middle and upper parts of the upper Albian can be referred to the Gnesioceramus Biozone, represented by G. comancheanus (Cragin) and G. bellvuensis (Reeside). These taxa are endemic to the Western Interior and some adjacent areas (Gulf Coast; Greenland?), but are closely allied to the cosmopolitan species, Gnesioceramus anglicus (Woods). At approximately the Albian-Cenomanian boundary, the endemic clade of ‘Inoceramus’ nahwisi appears, now referred to the newly erected Posidonioceramus, resulting in a distinct P. nahwisi biozone. This zone corresponds to the lower part of the ammonite Neogastroplites’ stratigraphic range. Gnesioceramids re-appear in the early Cenomanian. Close to base of the Cenomanian, for the first in the Western Interior, the genus Inoceramus, represented by Inoceramus irenensis Warren and Stelck, 1958, apparently immigrated into the Western Interior Basin.The Western Interior inoceramids do not allow for direct correlation to chronostratigraphic standard subdivision. The Albian-Cenomanian boundary, as earlier recognized on geochronologic correlations and confirmed, to some extent, based on ammonites, may approximately be located close to the appearance level of the genus Posidonioceramus. 相似文献
2.
Cenomanian rocks of Egypt are rich in moderately to well-preserved echinoids. From the large Cretaceous echinoid collection at the Cairo Geological Museum, a total of 27 regular species, in 12 genera, eight families, six orders and two subclasses, are revised and systematically described from these strata as exposed at several localities in Egypt. One new family (Pedinopsidae), one new subfamily (Lorioliinae), one new genus (Alternocidaris) and two new species of Goniopygus, G. subaequalis (Gebel Ekma) and G. macrotuberculatus (Gebel El Minsherah), are erected. Additionally, two species of Diplopodia, D. micropyga Fourtau, 1921 and D. halperti Fourtau, 1921, are transferred to the genus Pedinopsis. 相似文献
3.
Early and early middle Cenomanian turrilitid ammonoids from the upper part of the Albian–Cenomanian Aitamir Formation of the Koppeh Dagh (northeast Iran) are described, illustrated and placed in an integrated stratigraphic context. The Aitamir Formation represents a graded siliciclastic shelf system and the turrilitid faunas comprise eight different species. Representatives of Mariella and Hypoturrilites have been recorded from a lower Cenomanian horizon in the Ghorghoreh section, corresponding to the Mantelliceras mantelli ammonite biozone. Mariella bicarinata (Kner, 1852) and H. wiedmanni Collignon, 1964 are recorded for the first time from Iran and the larger palaeobiogeographical area, respectively. The horizons with Turrilites costatus and T. scheuchzerianus at Taherabad can be dated as early middle Cenomanian Acanthoceras rhotomagense Zone, T. costatus Subzone. Concentrations of ammonoids commonly occur above regressive sandstone units, at the transition into overlying deeper marine shale units and in their lower parts, respectively. Such intervals represent the early transgressive systems tract of depositional sequences and may thus be regarded as early transgressive, slightly condensed shell accumulations. The uppermost lower to lower middle Cenomanian succession at Taherabad is of striking bio-, event- and sequence stratigraphic similarity to contemporaneous sections in northwest Europe. It reflects deposition during falling and low sea-level associated with the latest early Cenomanian unconformity SB Ce 3 as well as the pulsating transgressive development during the early middle Cenomanian, including levels correlative to the arlesiensis, primus and Mid-Cenomanian events in northwest Europe. 相似文献
4.
The phylostratigraphy, taphonomy and palaeoecology of the Late Cretaceous neoselachian Ptychodus of northern Germany appears to be facies related. Ptychodus is not present in lower Cenomanian shark-tooth-rich rocks. First P. oweni records seem to relate to middle Cenomanian strata. P. decurrens appears in the middle to upper Cenomanian mainly in non-coastal environments of the shallow marine carbonate ramp and swell facies which isolated teeth were found partly in giant ammonite scour troughs on the Northwestphalian-Lippe High submarine swell in the southern Pre-North Sea Basin. They are recorded rare in deeper basin black shales facies (upwelling influenced, OAE Event II). P. polygyrus seems to be restricted to upwelling influenced basin and deeper ramp facies mainly of the uppermost Cenomanian and basal lower Turonian (OAE II Event). P. mammillaris is mostly represented during the lower to middle Turonian in the inoceramid-rich ramp and the near shore greensand facies along the Münsterland Cretaceous Basin coast north of the Rhenish Massif mainland. Finally, P. latissimus is recorded by two new tooth sets and appears in the upper Turonian basin swell facies and the coastal greensands. Autochthonous post-Turonian Ptychodus remains are unrecorded in the Santonian–Campanian of Germany yet. Reworked material from Cenomanian/Turonian strata was found in early Santonian and middle Eocene shark-tooth-rich condensation beds. With the regression starting in the Coniacian, Ptychodus disappeared in at least the Münster Cretaceous Basin (NW-Germany), but remained present at least in North America in the Western Interior Seaway. The Cenomanian/Turonian Ptychodus species indicate a rapid neoselachian evolution within the marine transgression and global high stand. A correlation between inoceramid shell sizes, thicknesses and their increasing size during the Cenomanian and Turonian might explain the more robust and coarser ridged enamel surfaces in Ptychodus teeth, if Ptychodus is believed to have preyed on epifaunistic inoceramid bivalves. 相似文献
5.
The fossil dendrobranchiate shrimp Macropenaeus was originally described from the Upper Cretaceous (Cenomanian) limestones of Hadjoula, northwest Lebanon. A new species, M. sidiaichensis sp. nov., was recovered from the Sidi Aïch Formation in the Northern Chotts Range, southern Tunisia. The Barremian occurrence of the genus in Tunisia suggests that Macropenaeus most likely originated in northern Africa and then migrated to Arabia in the Cenomanian. The association of the fossil shrimp with conchostracans, plant fragments, and fish indicates the interaction of freshwater and marine conditions that characterized the northern African margin as part of widespread coastal complex, paralic environments during the Barremian. 相似文献
6.
A detailed stratigraphic analysis was carried out on the Lower–Middle Cenomanian hemipelagic deposits of the Blieux section (Alpes-de-Haute-Provence; southeast France) in order to identify the Middle Cenomanian event I (MCE I) in the Vocontian Basin. These deposits are represented by five bundles composed of limestone–marl alternations that are separated by thick marly intervals. The Blieux section, which is well exposed, very thick, continuous and relatively rich in macrofauna, provides an ideal succession for an integrated approach. Biostratigraphy by ammonoids and sequence stratigraphy have been established for the whole succession whereas calcareous nannofossil and geochemical analyses have been carried out on a restricted interval across the Lower/Middle Cenomanian boundary. The uppermost part of the Mantelliceras mantelli Zone, the Mantelliceras dixoni Zone and the lower part of the Acanthoceras rhotomagense Zone have been recognized. The appearance of the genus Cunningtoniceras (C. inerme or C. cunningtoni) is used to place the base of the A. rhotomagense Zone and the Lower/Middle Cenomanian boundary. This boundary is also well characterized by the presence of nannofossil Subzone UC2C. Two orders of hierarchically stacked depositional sequences have been identified. Medium- and large-scale sequences correspond to 400 ky eccentricity cycles and to third-order cycles, respectively. The duration of the interval studied (from the uppermost part of the M. mantelli to the lower part of the A. rhotomagense zones) is estimated to be 2.8 my. Carbon-isotope values determined from bulk carbonate sediments show a first positive excursion (+0.6‰) corresponding to the MCE Ia, in the lower part of the A. rhotomagense Zone. A subsequent increase (+1.1‰) is recorded and could correspond to MCE Ib, but a sharp return to baseline values as expected in an excursion is not observed. The duration of the MCE I is estimated to be less than 400 ky. The Blieux section is correlated with some classical sections of the Anglo-Paris (Southerham, Folkestone, Cap Blanc-Nez) and Lower Saxony (Baddeckenstedt and Wunstorf) basins using ammonoid biostratigraphy, sequence stratigraphy, and chemostratigraphy. It is proposed as a candidate for the Middle Cenomanian GSSP (Global Boundary Stratotype Section and Point). 相似文献
7.
Pentanogmius Taverne (Actinopterygii: Tselfatiiformes) is a Late Cretaceous bony fish. Here, the diagnosis for the genus is emended and a new species, P. fritschi sp. nov., described. The new species is based on a nearly complete skeleton from the Britton Formation (upper Cenomanian–lower Turonian) of the Eagle Ford Shale in Dallas County, Texas, USA. The skeleton measures about 1.7 m in total length and represents the sole Cenomanian–Turonian example of Pentanogmius in North America. The most peculiar aspect of this new species is the morphology of the dorsal fin in which its anterior one-third is elongate to form a ‘hook-shaped sail.’ Pentanogmius fritschi sp. nov. was likely an active swimmer in open ocean environments that possibly fed opportunistically on a variety of relatively small pelagic fishes and invertebrates. The present stratigraphic record and anatomical evidence indicate the following phylogenetic hypothesis among the three North American Pentanogmius species: [P. fritschi sp. nov. [P. evolutus + P. crieleyi]]. 相似文献
8.
We re-define the Cretaceous bony fish genus Rhinconichthys by re-describing the type species, R. taylori, and defining two new species; R. purgatorensis sp. nov. from the lowermost Carlile Shale (middle Turonian), southeastern Colorado, United States, and R. uyenoi sp. nov. from the Mikasa Formation (Cenomanian), Middle Yezo Group, Hokkaido, Japan. Rhinconichthys purgatoirensis sp. nov. is designated on a newly discovered specimen consisting of a nearly complete skull with pectoral elements. Only known previously by two Cenomanian age specimens from England and Japan, the North American specimen significantly extends the geographic and stratigraphic range of Rhinconichthys. The skull of Rhinconichthys is elongate, including an expansive gill basket, and estimated maximum body length ranges between 2.0 and 2.7 m. Rhinconichthys was likely an obligate suspension-feeder due to its derived cranial morphology, characterized by a remarkably large and elongate hyomandibula. The hyomandibula mechanically acts as a lever to thrust the jaw articulation and hyoid arch both ventrally and anterolaterally during protraction, thus creating a massive buccal space to maximize filtering of planktonic prey items. Cladistic analysis supports a monophyly of suspension-feeding pachycormids including Rhinconichthys, but further resolution within this clade will require more information through additional fossil specimens. 相似文献
9.
Dominik Hennhoefer Aisha Al Suwaidi Cinzia Bottini Emina Helja Thomas Steuber 《Sedimentology》2019,66(2):536-555
The sedimentary record of the Arabian Shelf offers a unique opportunity to study the Cretaceous (Albian–Turonian) greenhouse climate from a palaeoequatorial perspective. In particular, hemipelagic to pelagic carbonate successions from the extensive Shilaif intra‐shelf basin have the potential to produce an excellent record of carbon cycle perturbations during this interval. This study presents a 269 m thick chemostratigraphic (carbonate δ13C and δ18O) record from the Middle Albian to Early Turonian of central Abu Dhabi (United Arab Emirates), representing over 14 Myr of uninterrupted carbonate sedimentation. The Mauddud to Shilaif formations represent outer ramp to basinal intra‐shelf carbonates with variations from laminated organic‐rich to clean bioturbated intervals. Isotopic evidence of the latest Albian Anoxic Event (Oceanic Anoxic Event 1d), Middle Cenomanian Event I and the Cenomanian–Turonian Anoxic Event (Oceanic Anoxic Event 2) are confirmed and biostratigraphically calibrated by means of calcareous nannofossils. The carbon isotope record allows correlation with other regional records and well‐calibrated records across the Tethyan Ocean and represents a significant improvement of the chronostratigraphic framework of the United Arab Emirates (Shilaif) and Oman (Natih) intra‐shelf basins. The study further confirms that low carbon isotope values corresponding to the two source rock intervals in the Shilaif Formation clearly precede the isotopic expressions of Oceanic Anoxic Event 1d and Oceanic Anoxic Event 2. 相似文献
10.
The Cenomanian–Turonian boundary interval is generally considered a critical time for planktonic foraminifera due to the environmental perturbations associated with Oceanic Anoxic Event 2. However, only the rotaliporids became extinct at the onset of the event, whilst several lineages evolved and/or diversified. This remarkable morphologic plasticity is often overlooked in the literature, partly because a number of stratigraphic sections have only been studied in thin-section due to the degree of lithification of the samples. Improved documentation of the morphological variability of planktonic foraminifera and better defined species concepts are required in order to improve biostratigraphy, particularly as Helvetoglobotruncana helvetica is an unreliable marker for the base of the Turonian. At the same time, detailed study of the planktonic foraminiferal response to OAE 2 demands a more profound knowledge of the assemblage composition.We present new biostratigraphic, taxonomic, and quantitative data for planktonic foraminiferal species from the Clot Chevalier section (Vocontian Basin, SE France), with the aim of (1) providing a detailed biostratigraphic analysis of the section, (2) documenting the morphological plasticity of specimens in this time interval and stabilizing species concepts, and (3) identifying promising markers to improve the resolution of the present biozonation and allow regional correlation. Samples were processed with acetic acid to extract isolated planktonic foraminifera. Assemblages were assigned to the upper Cenomanian Rotalipora cushmani Zone and to the uppermost Cenomanian–lowermost Turonian Whiteinella archaeocretacea Zone. Planktonic foraminiferal bioevents and assemblage composition identified at Clot Chevalier are compared with the well-studied Pont d'Issole section located ca. 15 km to the NE, highlighting similarities and differences in the species occurrences that may complicate the stratigraphic correlation between the two sections.The results of our study support the validity and common occurrence of species that have been misidentified and/or overlooked in the literature (i.e., Dicarinella roddai, Praeglobotruncana oraviensis, Marginotruncana caronae) and indicate that primitive marginotruncanids evolved before the onset of OAE 2, although species diversification occurred only after the event. Moreover, we believe that the first appearance of P. oraviensis might represent a promising bioevent for approximating the Cenomanian/Turonian boundary, after calibration with bio- and chemostratigraphically well-constrained sections. Finally, we describe three new trochospiral species, named “Pseudoclavihedbergella” chevaliensis, Praeglobotruncana pseudoalgeriana and Praeglobotruncana clotensis. 相似文献
11.
Sampling of Cenomanian fossil-rich horizons within the La Luna Formation of two localities in the Zulia and Trujillo states (northern Venezuela) yielded numerous shark teeth belonging to various species within the order Lamniformes (Mackerel sharks). Twelve lamniform species were identified including three new species (Squalicorax lalunaensis sp. nov., Squalicorax moodyi sp. nov., Acutalamna karsteni gen. et sp. nov.) and the genus Microcarcharias gen. nov. is proposed to accommodate with the peculiar morphology of the small-sized odontaspidid M. saskatchewanensis. Other taxa reported here include Cretoxyrhina mantelli, Cretolamna sp., cf. Nanocorax sp. and five Squalicorax species left in open nomenclature. This is the first report of chondrichthyans from the mid-Cretaceous of Venezuela and one of the few records of this group from the Cenomanian of South America. The composition of these assemblages suggests some degree of endemism in the La Luna Sea but also possible connexions with the Western Interior Seaway. One of the most striking features of these assemblages is the high anacoracid diversity (eight species) despite the corresponding outer shelf/upper slope palaeoenvironments of the La Luna Formation. The high diversity of these opportunistic predators is probably related to the high diversity of medium to large marine vertebrates that provided food resources. 相似文献
12.
The relatively complete and well preserved shell of a turtle, from the middle Cenomanian of Nazaré (Portugal), is studied here. It is recognized as a member of the crown group Pleurodira and, more specifically, of Bothremydidae. Pleurodira are one of the two lineages of modern turtles, their origin being in Gondwana. Pleurodira are very abundant in the uppermost Cretaceous record of Europe. However, this new finding is one of the few occurrences in the lower Upper Cretaceous of Laurasia. A single member of Bothremydidae had so far been identified in Portugal: the late Campanian-early Maastrichtian Rosasia soutoi. It was the only Cretaceous turtle identified in that country both at specific and at generic levels, being exclusive of Portugal. The taxon from Nazaré is identified as Algorachelus peregrinus, this form is also present in the contemporaneous beds in Spain, and is the oldest member of Bothremydidae in Laurasia. Algorachelus peregrinus is confirmed here to be a coastal form, which facilitated its spread. The two oldest known bothremydids from Laurasia, the European A. peregrinus and the North American Paiutemys tibert, are compared for the first time. They are recognized as closely related taxa. This study provides new data allowing a more precisely characterization of the oldest so far known dispersal event of Pleurodira in Laurasia, which was performed by an African lineage of Bothremydidae that reached the east coast of the Atlantic Ocean at least in the middle Cenomanian, and the west region of that Ocean at least in the late Cenomanian. 相似文献
13.
《Gondwana Research》2016,29(4):1579-1584
The subfamily Aleocharinae is a hyper-diverse group of Staphylinidae, accounting for nearly one-third of the diversity within the largest animal family on Earth. Here we describe Cretodeinopsis aenigmatica gen. et sp. nov., the first definitive Mesozoic aleocharine based on a well-preserved individual in the earliest Late Cretaceous (Cenomanian) amber from Myanmar (Burma). The new genus is clearly assigned to the ‘basal’ group of Aleocharinae based on its overall body shape, antennal insertions on the vertex of the head, and emarginate posterolateral margins of elytra. It is placed in the Recent tribe Deinopsini based on the strongly deflexed head, pectinate edge of abdominal segments III–V, tarsal structures, and non-pentamerous tarsi. The new discovery of the oldest known Aleocharinae from about 99 Mya represents a new aleocharine belonging to the ‘basal’ group of Aleocharinae, reinforcing the hypothesis that Gymnusa, Stylogymnusa and the tribe Deinopsini probably form a monophyletic lineage and suggest that the monophyly of Gymnusini is not supported based on the presence of 6 setigerous pores on the head. It indicates that the origin of the subfamily probably predates the Cenomanian. This new discovery is about 55 Ma older than the previously known Eocene-aged ones from Baltic amber, greatly broadening our knowledge on the early evolution of Aleocharinae. 相似文献
14.
15.
The Upper Cretaceous succession outcropping in the Anamas–Akseki Autochton, consists of approximately 500 m thick purely platform carbonate sediments. It begins with Cenomanian limestones intercalated with limestone breccias (Unit-1) containing mainly Pseudorhapydionina dubia, Pseudonummoloculina heimi, Spiroloculina cretacea (Assemblage I) and unconformably overlies the Lower Cretaceous (Barremian–Aptian) limestones with Vercorsella laurentii, Praechrysalidina infracretacea and Salpingoporella hasi. The Cenomanian limestones include foraminiferal packstone–wackestone, peloidal packstone–wackestone and mudstone microfacies deposited in restricted platform conditions. The Cenomanian succession is truncated by an unconformity characterised by locale bauxite deposits. Immediately above the unconformable surface, dolomitic limestones and rudistid limestones (Unit-2) are assigned to the upper Campanian based on the benthic foraminiferal assemblage (Assemblage II) comprising mainly Murciella gr. cuvillieri, Pseudocyclammina sphaeroidea, Accordiella conica, Scandonea samnitica and Fleuryana adriatica (smaller-sized populations). The upper Campanian limestones composed of dominantly foraminiferal-microbial packstone–wackestone microfacies deposited in shallow water environment with low energy, restricted circulation. The following limestones of the Unit-2 is characterised by sporadic intercalation of “open shelf” Orbitoides, Omphalocyclus, Siderolites assemblage (Assemblage III), assigned to the Maastrichtian, in addition to pre-existing “restricted platform” species. In the upper part of this biozone, the Rhapydionina liburnica/Fleuryana adriatica concurrent range subzone (Assemblage IIIb) is distinguished by the presence of Valvulina aff. triangularis, Loftusia minor as well as the nominal species. The Maastrichtian limestones with sporadically open marine influence consist of bioclastic (rudist-bearing) packstone–floatstone, foraminiferal packstone–wackestone with rudist fragments and peloidal/intraclastic packstone–wackestone microfacies deposited in shallow subtidal–subtidal (lagoonal) environments. The Upper Cretaceous succession passes upwardly into 70 m thick limestones and clayey limestones (Unit-3) which do not contain rudists and pre-existing foraminiferal assemblage with one exception Valvulina aff. triangularis. Variable amounts of ostracoda, discorbids, miliolids, dasycladacean algae and Stomatorbina sp. (Assemblage IV) occur into mud-rich microfacies suggesting restricted conditions with low water energy. A probable Danian age is proposed for the Unit-3 based on the occurrence of Valvulina aff. triangularis and Stomatorbina sp. which were previously recorded from Danian of peri-Tethyan platforms. 相似文献
16.
The taxonomic status of specimens referred to Tarchia kielanae, T. gigantea, and Minotaurasaurus ramachandrani has been contested. The holotype of T. kielanae, Zaklad Paleobiologii (Institute of Paleobiology)-Polish Academy of Sciences (ZPAL) MgD I/111, is redescribed. It has features common to both Palaeontological Institute-Russian Academy of Sciences (PIN) 3142/250, long considered Tarchia, and to M. ramachandrani but which are lacking in Saichania chulsanensis. The specimen PIN 3142/250 is not referable to Saichania but instead represents a new species, Tarchia teresae sp. nov. Similarly, the holotype skull of M. ramachandrani is not referable to T. kielanae, so Minotaurasaurus is a valid taxon. Tarchia is more derived than either Saichania or Minotaurasaurus while sharing features of both. 相似文献
17.
《Journal of African Earth Sciences》2007,47(1):1-8
The basal mudstones from the El-Nom borehole in the Gebel Abraq area in southern Egypt have yielded a diverse and relatively well preserved terrestrial palynoflora that includes Balmeisporites holodictyus, Crybelosporites pannuceus, Foveotricolpites gigantoreticulatus, Nyssapollenites albertensis, Retimonocolpites variplicatus and Rousea delicipollis. These suggest an Albian–Cenomanian age and deposition in a fluvio-deltaic environment; no marine phytoplankton is reported. The fern-dominated palynoflora and the overwhelming presence of kaolinitic clays suggest a warm, humid palaeoclimate. According to available knowledge, the mudstones in the Gebel Abraq area, equivalents of the so-called “Timsah Formation”, might be correlated with an older rock unit, the Maghrabi Formation, based on the new palynological age assessment. This new definition of local stratigraphy implies that the Bernice sheet of geological map of Egypt [Klitzsch, E., List, F., Pöhlmann, G., 1987. Geological map of Egypt, sheet NF 36 NE Bernice, 1: 500 000. Conoco and the Egyptian General Petroleum Corporation, Cairo] ought to be reconsidered. 相似文献
18.
One new genus with three new species of the family Prionoglarididae, Palaeosiamoglaris leinhardi Azar, Huang et Nel gen. et sp. nov.; P. burmica Azar, Huang et Nel sp. nov. and P. inexpectata Azar, Huang et Nel sp. nov. are characterised, illustrated and described from Cretaceous Burmese amber. Palaeosiamoglaris gen. nov. share several characters with the recent genus Siamoglaris Lienhard, 2004, endemic to the Oriental (Sino-Indian) Region. A new tribe Siamoglaridini Azar, Huang et Nel trib. nov. is established to accommodate the group Siamoglaris + Palaeosiamoglaris; the other Prionoglaridinae being attributed to Prionoglaridini. This discovery implicates a palaeobiogeographic repartition for this particular tribe (Siamoglarini) that is still the same currently. Most diagnostic features show a nearly stasis state, supporting the notion that these characters appeared at least by the early Cenomanian. A checklist of all prionoglaridids is given. 相似文献
19.
《Gondwana Research》2014,26(4):1357-1368
Oceanic Anoxic Event 2 (OAE 2) during the Cenomanian–Turonian (C/T) transition caused stepwise marine extinctions. Using organic compounds, stable carbon and oxygen isotopes, and foraminifera from three depth-transect sections in northern Spain, this study revealed repeated anoxic/euxinic events coinciding with warming and stepwise extinctions of planktonic and/or benthic foraminifera within intermediate to surface waters in the proto-North Atlantic during the C/T transition. Those short-duration euxinic events occurred four times: at 93.95 Ma, marked by the extinction of Rotalipora greenhornensis; at 93.90 Ma, marked by the extinction of Rotalipora cushmani; at the mid-time maximum of the plateau of the δ13C of carbonates (93.70 Ma); and at the time of the C/T boundary (93.55 Ma). Furthermore, the main benthic foraminiferal extinctions occurred during the first and second euxinic events in the upper slope, during the second and third euxinic events in the outer to middle shelf, and during the third and fourth events in the middle shelf. The main euxinic events in each section also showed a progression to the shallow shelf. The main anoxia–extinction events occurred in the upper slope and outer shelf then moved to the middle shelf. The shallowest section had relatively weak anoxia and a proportionally low extinction rate. These new findings indicate that foraminiferal extinctions started from the intermediate water and the continental slope and then moved to the continental shelf. This was the result of the repeated progression of euxinic–anoxic water from the upper slope to the middle shelf on the eastern continental margin of the proto-North Atlantic four times during a 400 kyr period, to the end of the Cenomanian. 相似文献
20.
We describe two new species of extinct biting midges in the fossil genus Protoculicoides: Protoculicoides hispanicus Szadziewski and Arillo, sp. nov. and Protoculicodes sanjusti Szadziewski and Arillo, sp. nov., from Albian (101–113 Ma) amber of San Just, Spain. Atriculicoides Remm, 1976 is recognized as a new junior synonym of Protoculicoides Boesel, 1937 (NEW SYNONYM). Devalquia Choufani et al., 2013 is regarded here as a new junior synonym of the extant genus Culicoides Latreille, 1809 (NEW SYNONYM). Metahelea roggeroi Choufani et al., 2013 from Upper Cretaceous French amber, is excluded from the tribe Heteromyiini and placed in the extant genus Stilobezzia Kieffer, 1911, tribe Ceratopogonini: Stilobezzia roggeroi (Choufani et al., 2013), comb. nov. The extinct tribe Atriculicoidini Szadziewski, 1996 is regarded as a new subfamily, Atriculicoidinae, stat. nov., that includes the fossil genus Protoculicoides, with 13 species reported from Albian, Turonian, Cenomanian, Coniacian, Santonian and Campanian ambers (78–113 Ma). Protoculicoides skalskii Szadziewski & Arillo, 1998 from Lower Cretaceous amber of Álava, Spain, Protoculicoides succineus Szadziewski, 1966 from Lower Cretaceous Lebanese amber and Protoculicoides burmiticus Szadziewski & Poinar, 2005 from Upper Cretaceous Burmese amber are transferred to the extinct genus Archiculicoides Szadziewski, 1996, comb. nov. Protoculicoides krzeminskii Choufani et al., 2014 from Lower Cretaceous Lebanese amber is transferred to the extinct genus Archiaustroconops Szadziewski, 1996, comb. nov. The Cretaceous subfamily Atriculicoidinae forms an unresolved trichotomy with the extant subfamilies Forcipomyiinae and Dasyheleinae, both of which date to the Eocene. The fossil record indicates that wings with macrotrichia in biting midges evolved during the mid-Cretaceous greenhouse climate. We suggest that this was an evolutionary adaptation to new atmospheric conditions with higher levels of CO2 in order for the antennal Johnston's organs of males to receive the vibrational sex signals produced by females during flight. 相似文献