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1.
Grasslands dominated by Stipa tenacissima are important ecosystems in the arid and semiarid climates of western Mediterranean, where they have originated by degradation of open dry forests and shrublands. At present, although the level of exploitation in these ecosystems is very low, succession processes seem to have stopped. In this study, we explore the role of microsite and post-dispersal seed predation on the seedling emergence of two species from advanced successional stages (Rhamnus lycioides, Quercus coccifera), three species from degraded stages (Cistus clusii, Helianthemum violaceum, Anthyllis terniflora), and of the dominant species (Stipa tenacissima). Seeds of these species were sown in plots divided into two microsites: (i) under tussocks of S. tenacissima and (ii) in the bare soil between tussocks. Soil moisture was significantly higher in the tussock microsites. The interaction observed between microsite and time, reflected the slower drying process under tussocks. Seed predation was in general high and differed significantly between species but not between microsites. Q. coccifera was especially affected by predation. The only germination detected in gap areas was that of H. violaceum. The results pointed to a direct facilitative effect of tussock grass on the germination in R. lycioides (20.4±8.0%) and S. tenacissima (41.3±5.7%) while the corresponding figures for bare ground were 0%. These results suggest that the recovery of potential vegetation in this ecosystem is not possible or, at least, is very slow because the residual shrub patches do not generate a sufficient number of seeds able to disperse to favorable germination microsites. S. tenacissima may act as facilitator of seedling recruitment if there is a supply of diaspores.  相似文献   

2.
Grazing effects on patchy dryland vegetation in northern Patagonia   总被引:1,自引:0,他引:1  
In this study the spatial patterns and dynamics of vegetation patches along a grazing gradient in the steppe ofLarrea divaricataandStipaspp. in NE Patagonia (Argentina) are described. A general effect of grazing is the reduction of total plant cover resulting from the decrease in cover of perennial grasses (Stipa speciosa, Poa ligularis, Stipa tenuis) and some tall shrubs (Chuquiraga hystrix, Bougainvillea spinosa, Lycium chilense). Dwarf shrubs (Nassauvia fueguianaandJunellia seriphioides) increase their cover under medium and/or high grazing pressures. Plant species are spatially grouped into patches which alternate with areas of bare soil. Eleven types of vegetation patch differing in the dominant plant functional type or species, floristic richness and size were identified with different relative frequency along the grazing gradient. Based on these results, it is postulated that grazing forces the replacement of large patches dominated by tall shrubs with high species richness, byLarrea divaricatapatches or small dwarf shrub patches with low species richness and the extinction of grass patches. This results from: (1) disruption of local balances of species deletions and additions; (2) fragmentation of large patches; and (3) formation of new vegetation patches. These changes lead to differing plant spatial organization and heterogeneity along the grazing gradient which may be described by characteristic arrays of vegetation patches.  相似文献   

3.
Grazing in arid Mediterranean ecosystems brings about changes in species diversity, but the best way to measure such changes is unclear. In this study we compared various methods in order to identify indicators that might be useful for the management and conservation of grazed arid Mediterranean ecosystems.Changes in community structure and composition were compared along a previously studied grazing gradient in Cabo de Gata-Nijar Natural Park. Plant diversity was measured by calculating diversity (Shannon information index), evenness (Pielou index) and richness (species number). Rarefaction curves were used to measure plant species richness and the results were compared with traditional richness values. Community structure was measured as the percentage of bare ground and the coverage of overall perennial grasses, Stipa tenacissima L., perennial shrubs and annual plants. Our results showed that the proportion of bare ground, the cover of perennial shrubs and the relative abundance of S. tenacissima are good indicators of grazing effects on vegetation. Overall plant community structure was more sensitive to the grazing gradient than were the diversity indices. Finally, the adequacy of standardized methods for comparing species richness (i.e. rarefaction curves) along a grazing gradient are discussed.  相似文献   

4.
Despite low growth rates, plants in arid areas have a strong ability to modify soil surface properties affecting ecosystem processes and community dynamics. But our knowledge on species effects on soil properties in these areas comes largely from observational studies, increasing the risk of confounding factors and precluding estimations of rates of change. We evaluated changes in soil surface properties underneath Acacia salicina, Pinus halepensis and Eucalyptus occidentalis in a 10-year-old common garden experiment established on a degraded Stipa tenacissima steppe in southern Tunisia. The three species tested improved soil properties compared to those of open areas. Acacia salicina ranked first as soil modifier as the soil underneath this species showed higher total organic carbon, total nitrogen, available phosphorus, soil CO2 efflux and infiltration rate, and lower soil hydrophobicity than soil in open areas. The richness of vascular plants was higher under A. salicina than under the other types of cover. This species showed higher ability to improve microsite conditions and foster succession. Short rotations of A. salicina could thus be employed for the restoration of degraded S. tenacissima steppes provided that other aspects of its ecology are controlled. Pinus halepensis represents a good alternative when native species are a priority, albeit facilitative ability is lower.  相似文献   

5.
We examined change in areal extent of mountain meadows and fields of deciduous shrubs and conifer saplings in the central western Cascade Range of Oregon, based on analysis of aerial photographs taken in 1946 and 2000. These nonforest vegetation patches are distinctive habitats in dominantly forested landscapes, such as the Cascades, and change in extent of these habitats is of interest to scientists and land managers. We mapped and dated even-aged forest stands of probable postfire origin around the nonforest patches, using tree-ring analysis and interpretation of aerial photographs. We used these and archival data to interpret possible influences of past wildfire and sheep grazing on the extent of nonforest patches. The total area of nonforest vegetation patches decreased from 5.5 percent of the study area in 1946 to 2.5 percent in 2000. Significantly more cases of contracted patches were observed for mesic and xeric meadows which have adjacent forest established after wildfire in the approximately 150 years preceding 1946. A higher proportion of mesic (47 percent, n = 47) than xeric (17 percent, n= 115) meadows contracted between 1946 and 2000. Broad-leaved shrub fields were unchanged, probably because of topo-edaphic controls, dense cover of shrubs, and snow effects; but all fields of conifer saplings underwent succession to forest. We observed no strong influences of sheep grazing on the extent of meadows.  相似文献   

6.
Presented are the results from investigating the belowground biomass of grass associations as an important factor of functional stability of mountain ecosystems. Field investigations were conducted in plant communities of the dry-steppe, meadow-steppe and alpine belts of Armenia at altitudes of 1200–3250 m above the sea level. It is shown that the conditions of vertical belts, topography and the slope aspect of the mountain landscapes under investigation have influence on belowground biomass accumulation and distribution in the profile of the root layer of soils.  相似文献   

7.
Savannas cover about 20% of the Earth's land area across a wide range of climatic conditions. As an important and distinct biome, savannas produce approximately 29% of global terrestrial net primary productivity. In these ecosystems the distribution of belowground resources remains poorly investigated and the relationship to the climatic conditions remains unclear. In the present study, vertical profiles of soil nutrients (chloride, nitrate, phosphate and sulfate) and nitrogen stable isotopes were analyzed at four sites along the Kalahari mega-transect, where a distinct rainfall gradient exists on a homogeneous soil substrate. The results show clear differences in nutrients and δ15N vertical distributions between wet and dry seasons. The results also show how the formation of “fertility islands” (i.e., the concentration of soil nutrients in the soils beneath tree canopies) is not necessarily coupled with belowground processes in that the distribution of soil nutrients at the surface does not match belowground patterns. The results also indicate that phosphorus may be a limiting nutrient in these savanna ecosystems with seasonal dynamics in its cycling.  相似文献   

8.
In this review we compile results on interactions above and below ground and food web functioning in an arid environment at the Baza Basin, in the Iberian southeast. Our data reveal that herbivory is difficult to estimate in our system. Some herbivores (rabbits) and granivores (Messor ants) create nutrient and detritus-rich patches, with important effects on the diversity and abundance of species both above and below ground. Fluctuations of prey availability, especially those caused by changes in their circadian rhythms, provoke seasonal migrations and omnivory in predators. We also present experimental evidence of the effects of interactions between aboveground and belowground macroarthropods on litter degradation; belowground detritivores are responsible for a high percentage of ground surface litter decomposition. We demonstrate that belowground herbivores and decomposers can dramatically change aboveground multitrophic interactions. Finally, we identify the biotic and abiotic factors controlling aboveground and belowground macrofaunal distribution at a broad scale. We conclude that trophic interactions in this semi-arid area are numerous and complex with many of the interactions involving more than two or three organisms. The interactions between above- and belowground organisms added complexity to this system. These habitats in which organisms deal with extreme abiotic conditions promote odd interactions resulting in an increase of biodiversity. Finally, despite the large research effort devoted to understanding food web structure and dynamics in this area recently, our state of knowledge is still far from providing a complete picture of interactions and their implications in the regulation and functioning of the system.  相似文献   

9.
The aim of this study is to assess the Iberian populations of Forsskaolea tenacissima L. according to its biogeographical interest, habitat, geographical range and conservation status. Results point out that they are restricted to gravel wadis of Tabernas Desert (SE Spain), are scarcely included in protected areas and represent historically isolated populations with relict behaviour. We also describe a new association, Senecioni-Forsskaoleetum tenacissimae. Conservation status of species is cause for concern and two conservation actions must be carried out. Firstly, protected areas should house Forsskaolea populations and secondly, phytosociological characterization of a community allows inventorying its habitat and directing conservation efforts to community level.  相似文献   

10.
Effects of soil and vegetation on runoff along a catena in semi-arid Spain   总被引:1,自引:0,他引:1  
Runoff and infiltration were investigated on abandoned fields of patchy vegetation in semi-arid Spain during 15 months of natural rainfall and by rainfall simulations. The aim was to ascertain sources and sinks of runoff and the effects of soils and plant cover. Soils of the catena developed from mica schists of the upper hillslopes, fan deposits of the lower hillslopes, and an alluvial terrace at the bottom. Runoff from natural events were from three sets of three pairs each of 10 × 2 m runoff plots. The pairs of each set had different densities of plant cover; the sets were vegetated with tussock grass, Stipa tenacissima, a shrub, Anthyllis cytisoides, and a bush, Retama sphaerocarpa. Nineteen natural rainfall events of intensities up to 18 mm/h produced 400 mm of rain during the study period. Because the rainfall threshold for runoff production was about 20 mm, only eight events produced runoff. The rainfall simulations used a sprinkler that produced 50 mm/h of rain for 30 minutes; runoff was recorded each minute in 0.24 m2 bounded plots.The depth and structure of the soil mantle provide the main controls on runoff rates, which are lowest on the lower fan deposits and highest on the thin upslope soils. The river-bank terrace, with a surface covered by crusts and mosses, also yields relatively high runoff. In general, vegetation density varies inversely with runoff. Nevertheless, shrub and bush litter favor runoff, as does a particular spatial distribution of individual plants on the hillslope. Settling of the upper few centimeters of soils of the alluvial fan following cessation of cultivation 15 to 40 years ago has produced a near-surface compacted layer favoring shallow subsurface runoff. Apparently contradictory results between runoff plots and rainfall simulations are the result of differing processes.  相似文献   

11.
Decreasing wind speed is one aspect of global climate change as well as global warming, and has become a new research orientation in recent decades. The decrease is especially evident in places with frequent perennially high wind speeds. We simulated decreased wind speed by using a steel-sheet wind shield in a temperate grassland in Inner Mongolia to examine the changes in physical environmental variables, as well as their impacts on the photosynthesis of grass leaves and net ecosystem exchange (NEE). We then used models to calculate the variation of boundary layer conductance (BLC) and its impact on leaf photosynthesis, and this allowed us to separate the direct effects of wind speed reduction on leaf photo- synthesis (BLC) from the indirect ones (via soil moisture balance). The results showed that reduced wind speed primarily resulted in higher moisture and temperature in soil, and indirectly affected net assimilation and water use efficiency of the prevalent bunch grass Stipa krylovii. Moreover, the wind-sheltered plant community had a stronger ability to sequester carbon than did the wind-exposed community during the growing season.  相似文献   

12.
The diets of euros or hill kangaroos (Macropus robustus), domestic sheep (Ovis aries) and feral goats (Capra hircus) in hilly, shrub rangeland in southern Australia were examined in a 12-year study. Levels of dietary overlap between these herbivores, their foraging in relation to resource availability, and the potential for competition in different conditions were also examined. The diet of euros was based around grasses. In dry seasons some shrubs were also eaten but in severe drought grass formed more than 80% of their diet. While grass was important to sheep in wetter conditions, they ate much shrub in dry conditions. Feral goats had broad diets but their preference for browse was high. In drought, euros had only a modest dietary overlap with sheep and goats. Data for dietary niche breadths and electivities pointed to only limited competition between the herbivores.  相似文献   

13.
Data from packrat middens are used to reconstruct the migration of single-leaf pinyon (Pinus monophylla) and Utah juniper (Juniperus osteosperma) in the White Mountain region of California and Nevada for the last 20,000 years. Today this region is characterized by arid conditions with dry summers. Pinyon woodlands, which are favored by dry summer conditions, are dominant at middle elevations of the White Mountains. The midden record indicates that during the late Pleistocene and early Holocene, juniper was dominant at elevations now occupied by desert shrubs. A late Pleistocene-early Holocene record of desert shrubs was found only in the double rainshadow of the Sierra Nevada and the White Mountains. Pinyon entered the area during the Holocene (ca. 9000 yr B.P.) and subsequently became more important than juniper. During the late Holocene, desert shrubs became established at the lower elevation sites that they now occupy. The late Pleistocene-early Holocene record indicates that there may have been a more zonal upper-level wind flow and associated penetration of Pacific moisture, as indicated by the occurrence of desert shrubs downwind of the north-south-trending mountain ranges. Zonal flow probably became less important during the early Holocene based on the increasing amount of pinyon. The increase in pinyon also may indicate a change from an even seasonal distribution of precipitation to the dry summer conditions presently found in the region. [Key words: biogeography, packrat middens, Great Basin paleoenvironments, Pinus monophylla, Juniperus osteosperma, White Mountains.]  相似文献   

14.
In three Larrea shrublands, cacti grow under Larrea canopy at northern and southern exposures and in open spaces were counted. Cacti distribution was related to environmental variables. Seven out of the 15 cacti present were found to be associated with shrubs, whereas the other species were found in the open spaces. The lower temperature and radiation occurring under shrubs at south exposures would explain the absence of cacti in these places. The higher organic matter, nutrients, and moisture in the soils under shrubs appeared to be correlated with higher cacti density. The presence of cacti seedlings only under shrubs could be attributed to the favourable environmental conditions for cacti seed germination and seedling survival that occurred in these places.  相似文献   

15.
Medium-sized kangaroo rats (Dipodomys spp. Gray) function as keystone species in the dynamics of desert bunchgrasses. We tested the hypothesis that kangaroo rat graminivory leads to reduced grass growth and inflorescence production, and that kangaroo rat grass consumption reflects their preferences for open microhabitats. We excluded kangaroo rats from grasses, and measured tiller loss, leaf and tiller growth and inflorescence production. We recorded the extent of tiller loss in grasses varying in tussock size or the extent of surrounding cover. Consumption of tillers by kangaroo rats was extensive, peaking in late summer. Grasses protected from kangaroo rat graminivory showed elevated leaf and tiller growth and inflorescence production. Grass tussocks protected by surrounding vegetation or large grass tussocks suffered lower graminivory than tussocks in the open or small tussocks. Our data demonstrate that the mechanism of the keystone role of kangaroo rats is through direct graminivory. We hypothesize that kangaroo rat graminivory may contribute to the persistence of desertified shrublands by limiting grass recruitment and abundance. Medium-sized kangaroo rats prefer open microhabitats and increase in abundance as such microhabitats increase, suggesting a positive feedback, with kangaroo rats increasing in abundance in overgrazed, desertified landscapes, and then contributing to their persistence through graminivory.  相似文献   

16.
祁连山高山灌丛生物量及其分配特征   总被引:2,自引:1,他引:1  
分布于青海云杉林上线的祁连山高山灌丛是祁连山水源涵养林效益最佳林型,研究高山灌丛生物量是生态系统生产力的重要体现,也是群落结构和功能的主要测度之一。以祁连山排露沟流域3 300~3 700 m高山灌丛为研究对象,采用标准地、样方收获法以及壕沟挖掘方法,对不同海拔高度10个固定样地灌丛生物量,器官生物量进行调查,分析了不同海拔梯度灌丛的叶、枝、须根、细根、粗根烘干重数据及其生物量分配特征。结果表明:祁连山高山灌丛总生物量为12 869.39±3 306.16 kg/hm2(平均值±标准差,n=10),其中器官分配以枝生物量所占比例最高,达32.21%,叶、须根、细根、粗根的比例分别为15.70%、14.06%、11.13%和26.90%。不同海拔梯度灌丛生物量器官分配比例差异较大,地上生物量平均为6 097.17kg/hm2,地下生物量平均为6 772.22kg/hm2,不同海拔根茎比在0.56~1.93之间变化,海拔3 500 m处根茎比达最大。祁连山高山灌丛生物量与海拔呈现显著的负相关(R2 = 0.898 7,p<0.01),随着海拔的升高,灌丛总生物量,地上以及地下生物量均呈现下降的趋势。研究结果可为内陆河流域生态环境的保护及其高山灌丛对全球气候变化的响应研究提供重要理论依据和应用资料。  相似文献   

17.
黄土高原西部荒漠草原植被恢复的土壤水分管理研究   总被引:3,自引:2,他引:1  
针对兰州地区荒漠草原带具有代表性的三种地带性植被:草本植被短花针茅、灌木植被珍珠猪毛菜、荒漠锦鸡儿,对其0~2.0 m的土壤含水量进行了比较,同时对草本和灌木植被的根系进行了测定。结果表明:天然植被对土壤含水量的影响在1 m以上。随着土壤深度的不断加深,土壤含水量呈减少趋势而后趋于稳定。短花针茅区1 m以上含水量比珍珠猪毛菜和荒漠锦鸡区平均高1.08和1.21个百分点;1 m以下土壤含水量变化情况是:珍珠猪毛菜区最高,荒漠锦鸡儿区最低。本文在分析了兰州地区荒漠草原区土壤水分动态的基础上,提出了这一地区植被恢复的对策。  相似文献   

18.
Pine plantations coexist with Stipa tenacissima grasslands in many semiarid western Mediterranean areas. We compared three microsites created by a 30-year-old Pinus halepensis plantation: below pine plantation line (BP), below canopy of pines (BC) and interline bare band (BA). They were evaluated in terms of soil properties, pine litter and suitability as recruitment niches for S. tenacissima. Next, in a manipulative experiment in growth chambers we tested the hypothesis that pine litter interferes with the seedling emergence of S. tenacissima. Three treatments in pots were compared: (a) soil from BA; (b) intact soil + litter from BP; and (c) soil + litter from BP, which was mixed in the laboratory (BPMX). In the field the main microsite differences were pine cover and litter cover and thickness. Seedling emergence was significantly greater in BA than in BP. Emergence and litter depth fits a linear regression model. In the growth chamber litter did not interfere with the emergence of S. tenacissima. However, seedlings grown without litter were 28% longer and their mass was 27% greater than in the litter treatments. The detected pine litter interference may be relevant for plant dynamics and might be considered in forestry management programs.  相似文献   

19.
The sustainability of ecosystem restoration of refuse dumps in open-pit coal mines depends on plant species selection, their configuration, and the optimal usage of water resources. This study is based on field experiments in the northern refuse dump of the Heidaigou open-pit coal mine in Inner Mongolia of China established in 1995. Eight plant configurations, including trees, shrubs, grasses, and their combinations, as well as the adjacent community of natural vegetation, were selected. The succession of the revegetated plants, soil water storage, the spatiotemporal distribution of plant water deficits degree and its compensation degree were also studied. Results indicated that the vegetation cover (shrubs and herbaceous cover), richness, abundance, soil nutrients (soil organic matter, N and P), and biological soil crust coverage on the soil surface are significantly influenced by the vegetation configurations. The average soil water storage values in the shrub+grass and grass communities throughout the growing season are 208.69 mm and 206.55 mm, which are the closest to that of in the natural vegetation community (215.87 mm). Plant water deficits degree in the grass and shrub+grass communities were the lowest, but the degrees of water deficit compensation in these configuration were larger than those of the other vegetation configurations. Differences in plant water deficit degree and water compensation among the different configurations were significant (P<0.05). Plant water deficit degrees were predominantly minimal on the surface, increased with increasing soil depth, and remained stable at 80 cm soil depth. The soil moisture compensation in the natural vegetation, shrub+grass, and grass communities changed at 10%, while that in other vegetation communities changed between 20% and 40%. Overall, we conclude that the shrub+grass and grass configuration modes are the optimal vegetation restoration models in terms of ecohydrology for future ecological engineering projects.  相似文献   

20.
We quantified soil nutrients and biological crust cover (bryophytes and lichens) under the canopies of three species of Mojave Desert shrubs and in interspaces between shrubs at three elevations to determine the effects of shrub species, soil crust, and elevation on islands of soil fertility. Means of pH, organic matter, total Kjeldahl nitrogen, nitrogen mineralization, and gravimetric soil moisture are significantly greater in soils under Ambrosia dumosa (Gray) Payne, Larrea tridentata Cov., and Coleogyne ramosissima Torr. than soils from adjacent interspace microhabitats. Although soil moisture and soil organic matter increase by a factor of 1.5 from the low elevation to the high elevation site, the ratio of shrub to interspace concentrations, or the difference in mean soil variables between shrubs and interspaces, is effectively constant and independent of elevation. Total bryophyte and lichen cover is relatively low (24.5%), however, there are 11 species of bryophytes and two species of lichens distributed across three elevations with the highest species richness and cover at the low-elevation site. Bryophyte and lichen cover is correlated with silt but is not related, consistently, to soil nutrients. Overall, the balance of processes controlling spatial aggregation of soil nutrients under shrubs is remarkably insensitive to potential differences in organic inputs among elevations, shrub species, and soil crust surfaces.  相似文献   

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