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1.
A combined approach of field geology and numerical simulation was conducted for evaluating the tsunami impacts on the shelf sediments. The 2003 Tokachi-oki earthquake, M 8.0, that occurred on 25 September 2003 off southeastern Hokkaido, northern Japan, generated a locally destructive tsunami. Maximum run-up height of the tsunami waves reached 4 m above sea level. In order to estimate the tsunami impacts on shallow marine sediments, we compared pre- and post-tsunami marine sediments in water depths of 38–112 m in terms of grain size, sedimentary structure, and microfossil content. Decreases of fine fractions, especially finer than very fine sand, which led to coarsen the mean grain size, were detected in the inner shelf of the northern part of the study area. Foraminiferal assemblages also changed in the coarsened sediments. On the other hand, the other shelf sediments largely unchanged or slightly fined. We also simulated the tsunami wave velocity and direction, and grain size entrained by the modeled tsunami. The numerical simulation resulted in that the 2003 tsunami could transport very fine sand in water depths shallower than 45–95 m at the northern part of the study area. This is comparable with the actual grain-size changes after the tsunami had passed. However, some storms and tidal currents might also be possible to stir the surface sediments after the pre-tsunami survey, so we could not conclude that the grain-size changes had been caused only by the tsunami. Nevertheless, a combined approach of sampling and modeling was powerful for estimating the tsunami impacts under the sea.  相似文献   

2.
The great Indonesian earth quake (26 December 2004) triggered a tsunami wave across the Bay of Bengal and Indian Ocean basins and has brought a major havoc in several countries including India. The coastal segment between Thotapalli and Valiazhikal in Kerala state of southwest India, where considerably rich beach placer deposit with ilmenite percentage of more than 70% is concentrated, has been investigated to understand the impact of tsunami on coastal sediments. The grain size analysis flashes out the significant differences between the pre- and post-tsunami littoral environments. While the mineral grains collected during pre-tsunami period show well-sorted nature, the post-tsunami samples represent moderately to poorly sorted nature. Similarly, unimodal and bimodal distributions of the sediments have been recorded for pre- and post-tsunami sediments, respectively. Further, mineral assemblages corresponding to before and after this major wave activity clearly indicate the large-scale redistribution of sediments. The post-tsunami sediments register increasing trends of garnet, sillimanite and rutile. The total heavy mineral percentage of the post-tsunami sediment also shows an improved concentration, perhaps due to the large-scale transport of lighter fraction. Magnetite percentage of post-tsunami samples reflects higher concentration compared to the pre-tsunami samples, indicating the intensity of reworking process. X-ray diffraction patterns of ilmenite grains have confirmed the increased presence of pseduorutile, and pseudobrookite in post-tsunami samples, which could be due to the mixing of more altered grains. SEM examination of grains also confirms the significant alteration patterns on the ubiquitous mineral of placer body, the ilmenite. The reason for these textural, mineralogical and micromorphological changes in heavy minerals particularly in ilmenite, could be due to the churning action on the deeper sediments of onshore region or on the sediments entrapped in the near shelf region of the area, by the ∼ 6 m high tsunami waves.  相似文献   

3.
Offshore sediment characteristics of the 2004 tsunami were identified from a shallow core collected from the Chennai Coast, India. The depositional sequence clearly distinguishes four different processes: mixed facies (post-tsunami): 0–8 cm; tsunami return flow facies (TRFF): 8–20 cm; tsunami landward flow facies: 20–44 cm; and pre-tsunami facies: 44–64 cm, which all took place during and after the tsunami event. The coarse-grained nature and higher carbonate in the TRFF indicate that considerable sediment load was transported from the beach/land area to the offshore region during the return flow of tsunami waves. The relatively greater abundance of benthic foraminiferal species in the core sample suggests that the taxa were transported from deeper regions of the inner shelf regions of Bay of Bengal region. The depositional characteristics in this region can be utilized for future comparative studies from this region as well as in other offshore regions affected by tsunamis with sequence-based studies on local topography.  相似文献   

4.
Present study examined phosphorus dynamics through delineation of source as well as availability of phosphorus and its fractionation within the intertidal sediments of Pichavaram mangrove ecosystem. Twelve sediment samples and two cores were collected from the mangrove forest along with estuarine area (Vellar-Coleroon) during January 2005. Sediments were analyzed for total phosphorus and its fractionation using operationally defined chemical sequential extraction scheme (SEDEX). Dissolved phosphorus (in water) and total phosphorus (in sediments) concentrations were high in the Vellar region of Pichavaram mangrove area due to pollution load from nearby villages and agricultural fields. However, the spatial variation in dissolved phosphorus were insignificant (at significance level = 0.05). The results for the phosphorus fractionation (post-tsunami) were compared with earlier studies (pre-tsunami). It was observed that all phosphorus fractions (except adsorbed-phosphorus) showed a highly significant (at significance level = 0.05) increase in concentration after the tsunami event. There was significant decrease in the adsorbed phosphorus concentration as a result of tsunami. The changes were more pronounced for organic phosphorus which increased by almost twofold following the event. These variations were attributed to change in salinity, increase in dissolved oxygen as well as the retreat of tsunami water carrying the waste load. The vertical distribution of phosphorus through core sediments showed that mixing after tsunami had altered the different phosphorus fraction and its availability. Overall, the study indicated that the fluvial weathering along with litter degradation and anthropogenic sources controlled the biogeochemistry of phosphorus in this mangrove ecosystem. Observed changes in the concentrations are a result of altered physico-chemical characteristics caused by tsunami.  相似文献   

5.
The Indian Ocean tsunami event of December 26, 2004 not only left massive casualties and economic damages, but also raised concerns about the destruction and recovery of coastal ecosystems. This work aimed to analyze the spatial patterns and temporal trajectories of vegetation damage and recovery using a multisensor multitemporal remote sensing, dataset. Using the study area of Koh Phra Thong, Thailand as a case study, we demonstrate the capabilities of remote sensing analysis in assessing the consequences of an extreme flooding event on the dynamics of coastal vegetation. Field surveys and satellite mid-resolution multispectral satellite data covering the period from February 2003 to December 2009 were used to map flooded areas and coastal vegetation loss and recovery following the tsunami. Normalized Difference Reflectance change detection was performed to map the extent of flooded areas. Vegetation Fraction Cover derived using spectral unmixing techniques was used to study the multitemporal changes in coastal vegetation after the event. Vegetation change detection techniques were applied to characterize the vegetation cover changes in two different time frames: short-term changes (from 4?days to 1?year after the event), and long-term dynamics (up to 5?years after). Estimates of vegetation change (decline, recovery, and gain) were quantified and mapped, with extreme vegetation losses found directly after the tsunami (up to 79?% in flooded areas). After 1?year, different trends had developed, indicating that recovering vegetation had reached up to 55?% of pre-tsunami land cover, but with different trajectories for each vegetation type.  相似文献   

6.
After the 2004 Sumatra?CAndaman tsunamigenic earthquake, waters from the ocean moved upstream along rivers, bays, harbors, and lagoons and inundated many coastal and inland locations in the southern, eastern, and northern parts of Sri Lanka. The tsunami waters were observed to move upwards inland and then recede downwards to the ocean after varying inundation periods in different coastal areas. Subsequent massive tsunami waves came with the wave height varying from 3 to 8?m inland with speed of about 30?C40?kmph. The oceanic waves carrying heterogeneous sediments with water deposited them in coastal as well as inland locations about 1?km from the present coastline. Given the chaotic nature of tsunami oceanic waves, pre-tsunami deposits, such as beach sands, debris from coral reefs and buildings, parts of vehicles and ships, and tree trunks are found incorporated in authentic tsunami sediments. Thus, the texture, structure, and composition of sediments deposited by tsunami waters differed from one location to another. Therefore, in identifying paleo-tsunami sediments, care was taken to compare them with diagnostic unmixed uncontaminated recent tsunami sediments having characteristic textures and marine microfossil assemblages, such as foraminifera, radiolarians, and diatoms where preserved in coastal depressions. The radiocarbon ages of the carbonate and the organic fractions of these sediments are stratigraphically inconsistent, indicating mixing of sediments by the tsunami waves. The concentrations of organic carbon and nitrogen and their isotopic signatures confirm marine origin of these sediments.  相似文献   

7.
The Sumatra–Andaman Tsunami left distinctive sedimentological and geomorphological signatures in the area of Khao Lak. Fine-grained sediments, predominantly layers of cohesive, carbonate-rich, fine-sandy silt with thicknesses of 1–10 cm, erosionally overlying pre-tsunami sandy soils and sediments, represent the most common tsunami deposits in the study area. Petrographically, they differ significantly from other coastal sediments and affiliated soils. Due to their grain size and corresponding clay mineral content, muddy shelf sediments (sub-wave base) are indicated as a main source. The present results suggest that indications of shelf influence, although varying regionally, might contribute to the identification of fine-grained tsunami sediments and their differentiation from storm sediments. However, the observed differences of tsunami sediments to soils and other coastal sediments, especially with respect to carbonate mineralogy, might disappear in short geological time under conditions of intensive weathering and bioturbation. At Cape Pakarang, hundreds of boulders with up to 24 tons were deposited on the foreshore and upper shoreface. Applying Nott’s (Earth Planet Sci Lett 210:269–276, 2003) formulas, minimum flow velocities of 3.9 m/s are required to transport the largest boulders. The devastating tsunami effect of both, onshore flow and backflow, is documented by damaged human constructions. Geomorphological effects include intensive widening of estuary mouths and the development of erosional channels. Now, estuary mouths are reduced, and erosional channels cut off from the sea due to the formation of a post-tsunami beach ridge.  相似文献   

8.
对从2011年9月至2012年6月在黄河兰州段采集的24个水样进行了孢粉分析,并探讨了河流搬运花粉组合随时间变化的特征以及累积花粉的组合状况。结果表明,河流中花粉组合和花粉浓度随时间具有明显的变化,春夏时段花粉组合以榆属、杨属、柳属和桦属等乔木花粉为主,秋冬时段以蒿属、藜科等草本花粉为主,且花粉浓度和河流含沙量有较好的正相关关系。根据河水流量与花粉浓度积分计算得出样品采集时段的累积花粉通量,其组合特征与各时间点的花粉组合具有明显的差异,表明单个时间节点花粉组合由于影响因素较多不能够反映流域植被状况,而累积花粉组合可以指示泥沙和花粉源区的植被状况。本研究中黄河兰州段河水搬运花粉可能指示黄河兰州段以上至刘家峡水库之间黄土区的植被情况。  相似文献   

9.
The Indian Ocean tsunami flooded the coastal zone of the Andaman Sea and left tsunami deposits with a thickness of a few millimetres to tens of centimetres over a roughly one-kilometre-wide tsunami inundation zone. The preservation potential and the post-depositional changes of the onshore tsunami deposits in the coastal plain setting, under conditions of a tropical climate with high seasonal rainfall, were assessed by reinvestigating trenches located along 13 shore-perpendicular transects; the trenches were documented shortly after the tsunami and after 1, 2, 3 and 4 years. The tsunami deposits were found preserved after 4 years at only half of the studied sites. In about 30% of the sites, the tsunami deposits were not preserved due to human activity; in a further 20% of the sites, the thin tsunami deposits were eroded or not recognised due to new soil formation. The most significant changes took place during the first rainy season when the relief of the tsunami deposits was levelled; moderate sediment redeposition took place, and fine surface sediments were washed away, which frequently left a residual layer of coarse sand and gravel. The fast recovery of new plant cover stabilised the tsunami deposits and protected them against further remobilisation during the subsequent years. After five rainy seasons, tsunami deposits with a thickness of at least a few centimetres were relatively well preserved; however, their internal structures were often significantly blurred by roots and animal bioturbation. Moreover, soil formation within the deposits caused alterations, and in the case of thin layers, it was not possible to recognise them anymore. Tsunami boulders were only slightly weathered but not moved. Among the various factors influencing the preservation potential, the thickness of the original tsunami deposits is the most important. A comparison between the first post-tsunami survey and the preserved record suggests that tsunamis with a run-up smaller than three metres are not likely to be preserved; for larger tsunamis, only about 50% of their inundation area is likely to be presented by the preserved extent of the tsunami deposits. Any modelling of paleotsunamis from their deposits must take into account post-depositional changes.  相似文献   

10.
Coastal communities in the western United States face risks of inundation by distant tsunamis that propagate across the Pacific Ocean as well as local tsunamis produced by great (Mw?>?8) earthquakes on the Cascadia subduction zone. In 1964, the Mw 9.2 Alaska earthquake launched a Pacific-wide tsunami that flooded Cannon Beach, a small community (population 1640) in northwestern Oregon, causing over $230,000 in damages. However, since the giant 2004 Indian Ocean tsunami, the 2010 Chile tsunami and the recent 2011 Tohoku-Oki tsunami, renewed concern over potential impacts of a Cascadia tsunami on the western US has motivated closer examination of the local hazard. This study applies a simple sediment transport model to reconstruct the flow speed of the most recent Cascadia tsunami that flooded the region in 1700 using the thickness and grain size of sand layers deposited by the waves. Sedimentary properties of sand from the 1700 tsunami deposit provide model inputs. The sediment transport model calculates tsunami flow speed from the shear velocity required to suspend the quantity and grain size distribution of the observed sand layers. The model assumes a steady, spatially uniform tsunami flow and that sand settles out of suspension forming a deposit when the flow velocity decreases to zero. Using flow depths constrained by numerical tsunami simulations for Cannon Beach, the sediment transport model calculated flow speeds of 6.5?C7.6?m/s for sites within 0.6?km of the beach and higher flow speeds (~8.8?m/s) for sites 0.8?C1.2?km inland. Flow speed calculated for sites within 0.6?km of the beach compare well with maximum velocities estimated for the largest tsunami simulation. The higher flow speeds calculated for the two sites furthest landward contrast with much lower maximum velocities (<3.8?m/s) predicted by numerical simulations. Grain size distributions of sand layers from the most distal sites are inconsistent with deposition from sediment falling out of suspension. We infer that rapid deceleration in tsunami flow and convergences in sediment transport formed unusually thick deposits. Consequently, higher flow speeds calculated by the sediment model probably overestimate the actual wave speed at sites furthest inland.  相似文献   

11.
Sue Dawson   《Sedimentary Geology》2007,200(3-4):328-335
Distinctive diatom assemblages may be associated with tsunami sediments and may often contrast with the assemblages found within sediments underlying the tsunami deposit as well as those associated with the modern coastal environment. Sediments associated with the 1998 tsunami that destroyed much of the Sissano lagoon area in northern Papua New Guinea have been investigated. Surface sediments from three transects across the sediment spit near Warapu have been examined for diatom content and preservation. The preservation is variable, and the data show an, often chaotic, assemblage that can be attributed to the tsunami waves incorporating and depositing diatoms from distinctive habitat zones during their runup and subsequent backwash. The diatoms identified within the Warapu sediments indicate an origin from within the inter-tidal and offshore area rather than from the beach–sand spit complex. The sand deposits disclose a high percentage, in excess of 75%, of broken diatom valves, and a predominance of centric (circular) species due to preferential preservation. The study demonstrates that the application of diatom biostratigraphy to modern tsunami deposits can be used in conjunction with other stratigraphical lines of evidence to interpret the source and provenance of historical and palaeo-tsunami deposits.  相似文献   

12.
Natural catastrophes could damage island biodiversity and ecosystems, and their effects could become devastating if combined with human disturbances. In this study, we determined the effects of the tsunami occurred in Robinson Crusoe Island (Chile) on 27 February 2010 on an endangered soil–plant system. Using data of endemic Cabbage Trees (Dendroseris litoralis Skottsb.) and soil attributes taken before and after the 2010 event, we developed thematic maps to assess the changes in population size and soil substrate of Cabbage Trees caused by the tsunami. We determined that from 153 pre-tsunami (2009) standing Cabbage Trees, only 66 (43 %) survived in 2011, mostly in elevations above 25 m a.s.l. Before the tsunami, 86 (56 %) of Cabbage Trees were established in humus-rich soil sites whereas after the tsunami, this number declined to 53 (35 %). These results represent the first report of a severe population decline after a tsunami and indicate that tsunamis are an important source of species extinction in small oceanic islands not only by reducing the population size but also by reducing the quality of sites for plant growth.  相似文献   

13.
Trophic groups were used to investigate broad patterns and predict species interactions of macrobenthic assemblages in Calcasieu Estuary, Louisiana. Macrobenthic assemblages of the estuary were numerically dominated by deposit-feeding species. Surface-deposit feeders were the most abundant macrobenthos of the upper estuary, subsurface-deposit feeders dominated the lower estuary, and a trophically well-mixed assemblage inhabited the middle estuary. There were periodic shifts in species dominance within each region, but the shifts were without temporal pattern and seldom led to changes in trophic structure of the region. The macrobenthic assemblages of the upper estuary were dominated by several early-colonizing species which switched feeding modes with changes in water flow, suggesting that disturbances in the upper estuary had a direct effect on macrobenthic communities. Disturbances also may have eliminated macrobenthic species before they could establish a well-mixed community. The presence of trophically mixed communities in the middle estuary probably was indicative of fewer disturbances there. The dominance by subsurface-deposit feeders in the lower estuary probably indicates that sedimentary food in Calcasieu Lake was seldom utilized at the sediment-water interface. Rather, most food became buried and was available only to subsurface-deposit feeders.  相似文献   

14.
The 2004 tsunami deposits and probable paleotsunami deposits were studied at the southern Kho Khao Island, on Andaman Sea coast of Thailand. The 2004 tsunami laid down about 8?cm of fining upward medium sand and locally about 40?cm of massive coarse sand with common mud clasts. The sediments were characterized by the presence of marine foraminiferal assemblage; however, already after 5?years many of carbonate foraminiferal tests were partly or completely dissolved. The probable paleotsunami deposits form layer about 1?m thick. It consists of massive very coarse sand with common big shells and mud clasts. Its composition suggests a marine origin and the presence of mud clasts, and similarity to the 2004 tsunami deposits suggests that the layer was left by paleotsunami, which took place probably during the late Holocene, even though two shells within the layer gave 14C ages of 40,000?years or more.  相似文献   

15.
The structure of macro infaunal (>0.5 mm sieve size) assemblages was examined in samples of unconsolidated substrates collected during the summers of 1994–1997 at 208 stations throughout North Carolina estuaries. Numerical classification (cluster analysis) of stations resulted in 14 distinct site groups that reflected discernible habitatrelated patterns in species distributions. Multiple discriminant analysis, performed on synoptic abiotic variables (depth, salinity, dissolved oxygen, pH, percent silt-clay), showed that the separation of site groups was related primarily to salinity. Percent silt-clay had a secondary influence on the separation of sites within similar salinity zones. Species diversity among site groups generally decreased with decreasing salinity and increasing mud content of sediment. Nodal analysis showed a wide range in constancy and fidelity of species assemblages within site groups. Some assemblages dominated by euryhaline species had no particular affinity with any one site group. The strongest affinities, as evidenced by high values of both constancy and fidelity, were displayed by an assemblage of oligochaetes, insect larvae, gammaridean amphipods, and the clamCorbicula fluminea in tidal freshwater muds; and an assemblage of haustoriid amphipods, the bivalveDonax variabilis, the polychaeteParaonis fulgens, and unidentified echinoids at high-salinity sites in outer Pamlico Sound near ocean inlets. A series of stations with impaired benthic assemblages in polluted habitats emerged from the cluster analysis and was distinguishable from other site groups that reflected a greater influence of natural controlling factors (such as salinity and sediment type) on species distributions. These results suggest that the interaction of both anthropogenic and natural environmental controlling factors is important in defining the structure of these infaunal assemblages.  相似文献   

16.
An objective of the Mid-Atlantic Integrated Assessment Program (MAIA) of the U.S. Environmental Protection Agency is to develop an index for assessing benthic community condition in estuaries of the mid-Atlantic region of the United States (Delaware Bay through Pamlico Sound). To develop such an index, natural unimpaired communities must first be identified and variability related to natural factors accounted for. This study focused on these two objectives; Lnansó et al. (2002) describe the index. Using existing data sets from multiple years, classification analyses of species abundance and discriminant analysis were employed to identify major habitat types in the MAIA region and evaluate the physical characteristics that structure benthic infaunal assemblages. Sampling was restricted to soft bottoms and to the index development period, July through early October. The analyses revealed salinity and sediment composition as major factors structuring infaunal assemblages in mid-Atlantic estuaries. Geographical location was a secondary factor. Nine habitat classes were distinguished as a combination of 6 salinity classes, 2 sediment types, and the separation of North Carolina and Delaware-Chesapeake Bay polyhaline sites. The effect of sediment types on faunal assemblages was restricted to polyhaline sites, which were separated into two sediment groups above and below 90% sand content. Assemblages corresponding to each of these 9 habitats were identified in the context of widely recognized patterns of dominant taxa. Differences between North Carolina and Delaware-Chesapeake Bay polyhaline assemblages were attributed to the relative contributions of species and not to differences in species composition. No zoogeographic discontinuities could be identified. Our results reinforce the findings of recent studies which suggest that, with respect to estuarine benthic assemblages, the boundary between the Virginian and the Carolinian Provinces be moved to a new location south of Pamlico Sound.  相似文献   

17.
Disturbances influence ecological communities over a wide range of scales. We investigated the effects of localized hypoxic disturbances on an estuarine fish assemblage at several spatial (m2 and 10s km2) and temporal (days, seasons, years) scales in a multivariate framework (temperature, salinity, depth, dissolved oxygen). We examined whether seasons, years) scales in a multivariate framework (temperature, salinity, depth, dissolved oxygen). We examined whether there were consistent changes in fish and crustacean estuarine assemblage characteristics along environmental gradients and whether these relationships were altered by hypoxic disturbances. We also investigated at what scale dissolved oxygen concentration may be influencing the structure of motile estuarine assemblages and whether the size of the hypoxic zone altered its effects on the estuarine assemblage. Hypoxic disturbances altered fish and crustacean assemblages along the depth gradients that were present during well-oxygenated periods. Species diversity, richness, and catch rates were lower in hypoxic patches than in oxygenated areas. Dissolved oxygen concentration remained an important explanatory variable for patch-level assemblage dissimilarity, species richness, and diversity when data were aggregated across seasons. When we examined the data at a larger scale, by aggregating information across the study area, we did not detect influences of hypoxia on assemblage structure. Fish moved out of local hypoxic zones, but remained within the estuary even in years with extensive hypoxia. There was no effect of size of the hypoxic distrubance on whether organisms responded to hypoxia or on diversity or richness of the study site. These results suggest that these local disturbances play an important role in structuring motile species assemblages at a patch-level within an estuary, but regional factors such as recruitment and migration are important in influencing species assemblages for the entire estuary over months and years.  相似文献   

18.
The structure of the demersal fish assemblage in Sepetiba Bay, southeast Brazil (22°54′–23°04′S, 43°34′–44°10′W) was described and related to environmental variables. A 3-yr long trawl survey (from July 1993 to June 1996) was conducted using a stratified random sampling strategy at 158 stations. One hundred and seven species of fishes in 80 genera and 44 families were identified. Gerreidae and Ariidae made up 30.5% and 28.6%, and 19.8% and 28.1% of the total number and weight, respectively. Sciaenidae, Engraulididae, Sparidae, and Carangidae were other prominent families. Three assemblages existed in the bay: one inhabited shallow, less saline and less transparent water in the inner zone (depth <5 m, transparency <2 m, salinity <30 psu, and slightly higher temperature); another in the outer zone with the opposite situation (depth >10 m, transparency >3 m, salinity >30 psu, and slightly lower temperature); and a third assemblage in the middle zone likely to prefer average values or show no clear preferences. Most species decreased in abundance over the 3-yr period of study. No seasonal change was detected in the assemblage structure, but many species were rare, indicating high spatial and temporal patchiness. Canonical correlation analyses were used to describe and compare the fish assemblages in the three different habitats. The outer zone had the greatest number of species recorded, lowest abundance, highest diversity and evenness, and was comprised mainly byCynossion leiarchus, Prionotus punctatus, Anchoa tricolor, Haemulon steindachneri, Diplectrum radiale, Etropus crossotus, andSphoeroides greeleyi; the inner zone had the lowest number of species recorded but the greatest number of species per sample, highest abundance, lowest diversity and evenness, being comprised mainly byDiapterus rhombeus, Genidens genidens, Cathrops spixii, Chloroschombrus chrysurus, Sciadeichthyes luniscutis, Cetengraulis edentulus, andArchorsargus rhomboidalis; while the third group had the lowest number of species per sample, being comprised by species widely distributed asGerres aprion, G. genidens, Gerres gula, andMicropogonias furnieri. Depth, followed by transparency and salinity, were the primary factors influencing assemblage distribution.  相似文献   

19.
 Cold-seep communities have relatively low diversity, are dominated by one or two taxa present in high density and high biomass in comparison with the surrounding fauna, and are restricted to aphotic habitats. Their associated heterotrophic fauna are usually distinctive from the fauna of their surroundings. In contrast, a more commonplace chemoautotrophically based community occurs in shallow photic habitats. The associated heterotrophic fauna includes many of the species typical of the surrounding communities and typically dominates abundance, whereas the species with chemoautotrophic symbionts dominate biomass. All modern seep assemblages are deeper than 550 m. Many fossil seep assemblages occurred in water as shallow as the mid-shelf (<200 m). In contrast, communities where species with chemoautotrophic symbionts are biomass dominants, but not numerical dominants, are common in shallow waters at present but rarely reported in the geological record. We suggest that the absence of cold-seep communities on the continental shelf presently is due to a combination of predation and competitive exclusion by primary consumers limiting the presence of species dependent on chemoautotrophic symbionts. We suggest that cold-seep assemblages are more common at shelf depths in the fossil record for two reasons: (a) The biases of preservation have accentuated their distribution by transforming communities where species with chemoautotrophic symbionts dominate by biomass, but not numerically, into cold-seep-appearing assemblages. (b) The importance of predation pressure and oligotrophy has varied, with decreased predation pressure accompanying increased oligotrophy favoring cold-seep communities. We suggest that the paucity of shallow-water assemblages with species harboring chemoautotrophic symbionts as biomass dominants in the fossil record is based on the reliance of paleoecological analysis on numerical abundance data when energy flow analyses are required to identify these assemblages. The distinctiveness of the fossil seep assemblage is intensified by taphonomic processes that bias the assemblage against small individuals and epifaunal species, so that diversity declines, the small heterotrophic component of the assemblage is significantly reduced, and the epifaunal component is minimized. The final assemblage is usually dominated by the better-preserved large infaunal clams which perchance are also the species with chemoautotrophic symbionts. In contrast, preservation does not enhance the distinctiveness of these chemoautotroph-harboring species in shallow water. Received: 16 April 1998 / Accepted: 29 June 1999  相似文献   

20.
Quantitative plant assemblage data from coal balls, miospores, megaspores, and compression floras from the Calhoun coal bed (Missourian) of the Illinois Basin (USA) are used to interpret spatial and temporal changes in plant communities in the paleo-peat swamp. Coal-ball and miospore floras from the Calhoun coal bed are dominated strongly by tree ferns, and pteridosperms and sigillarian lycopsids are subdominant, depending on geographic location within the coal bed. Although the overall composition of Calhoun peat-swamp assemblages is consistent both temporally and spatially, site-to-site differences and short-term shifts in species dominance indicate local topographic and hydrologic control on species composition within the broader context of the swamp. Statistical comparison of the Calhoun miospore assemblages with those from other Late Pennsylvanian coal beds suggests that the same basic species pool was represented in each peat-swamp landscape and that the relative patterns of dominance and diversity were persistent from site to site. Therefore, it appears that the relative patterns of proportional dominance stayed roughly the same from one coal bed to the next during Late Pennsylvanian glacially-driven climatic oscillations.  相似文献   

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