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1.
Tunisia, Ukraine, Russia, the North Sea area and elsewhere have yielded knowledge that the Early Cretaceous planktonic foraminifera are predominantly members of the Praehedbergellidae. The taxonomy and phylogeny of this family have been revised and refined. New generic and specific taxa have been proposed, new phylogenetic lineages have been described, and many type specimens have been imaged by scanning electron microscopy for the first time.Gorbachikella(Hauterivian–Early Aptian) gave rise toPraehedbergella(Late Hauterivian–Late Aptian), evolvingBlefuscuiana,Lilliputianella, and the planospiralBlowiellain the Barremian. The last gave rise toGlobigerinelloidesin the Aptian, from which the macroperforateAlanlordella(and its descendantPlanomalina) evolved in the Albian.Blowiellawas ancestral to taxa with radially elongated chambers (Claviblowiella, Leupoldina, Schackoina).  相似文献   

2.
The Khami group within the petroliferous Zagros Basin of Iran includes the Surmeh, Hith, Fahliyan, Gadvan and Dariyan formations. The Dariyan Formation, previously known as Orbitolina limestone is very signif-icant as a reservoir rock in the Zagros Basin. In this study, the age of the Dariyan Formation is examined based on biostratigraphic and paleontological investigations. 236 m of Dariyan Formation sediments in the sections at Kuh-e Rahmat （north） and Nour-Abad （west） of Iran were selected to study its biostratigraphy. According to the lithostratigraphic investigations, the Dariyan Formation in these sections is subdivided into three： lower, middle and upper, which include thin-bedded to medium-bedded and massive lime-stone. Micropaleontologic study of the Dariyan Formation has led to the recognition of two biozones in the Nour-Abad area and one biozone in the Kuh-e Rahmat stratigraphic column. Diagnostic foraminifera in these sections are as follows： Globigerinel oides algerianus Cushman and Ten Dam, Globigerinelloides fer-reolensis Moullade, Globigerinelloides maridalensis （Bolli）, Globigerinelloides paragottisi Verga and Permoli Silva, Hedbergel a aptiana Bartenstein, Hedbergel a excelsa Longoria, Hedbergel a luterbacheri Longoria, Hedbergella occulta Longoria, Hedbergella ruka （Banner et al.）, Hedbergella similis Longoria, Hedbergella trocoidea （Gandolfi）, Muricohedbergella delrioensis （Carsey）, Pseudoplanomalina cheniourensis Sigal, Con-icorbitolina conica d＇Archiac, Iraqia sp., Mesorbitolina parva Douglass, Mesorbitolina pervia Douglass, Meso-rbitolina ovalis Gorog and Arnaud-Vanneau, Mesorbitolina subconcava Leymerie, Mesorbitolina texana （Roemer）, Mesorbitolina ovalis-pervia Cherchi and Schroeder, Debarina hahounerensis Fourcade et al., Marssonel a trochus （d＇Orbigny）, Nezzazata sp., Orbitolina sp., Textularia sp.  相似文献   

3.
The Upper Barremian to Aptian Almadich Formation (Inner Prebetic Domain of the Betic Cordillera) is composed of hemipelagic sediments deposited on a distal carbonate ramp in the southern Iberian Palaeomargin. Within this facies we have found a thick interval of blue to black shales and marls that is interpreted as deposited under oxygen-depleted conditions. We think that this interval, dated as early Aptian, represents the local record of Ocean Anoxic Event 1a. The integrated biostratigraphic analysis of a section in the Almadich Formation, by means of planktonic foraminifera, calcareous nannofossils and ammonites, enables us to recognize most of the biostratigraphic units based on these three fossil groups and to correlate between them. The Sartousiana, Sarasini, Weissi, Deshayesi and Furcata (ammonite) Zones were identified for the Upper Barremian–Lower Aptian interval. By means of calcareous nannofossil biostratigraphy the Micrantholithus hoschulzii, Hayesites irregularis and Rhagodiscus angustus Zones, plus several additional biohorizons, were identified. A quantitative study performed on a set of 27 Lower Aptian samples has enabled the precise identification of the ‘nannoconid crisis’, the lower limit of which clearly precedes the main anoxic event, and its correlation with other bioevents. Planktonic foraminifera occur consistently throughout the Lower to Upper Aptian of the Cau section and are moderately well preserved. This fact allows us to use the most recent taxonomic framework, based on wall texture, to identify the Blowiella blowi, Schackoina cabri, Globigerinelloides ferreolensis, Globigerinelloides algerianus, Hedbergella trocoidea andTicinella bejaouaensis Zones. Coincident with the anoxic episode, the planktonic foraminiferal assemblages are composed of a significant number of forms with elongated chambers and/or tubulospines assigned to the genera Claviblowiella,Lilliputianella , Leupoldina and Schackoina. Most of the planktonic foraminiferal and nannofossil taxa are illustrated.  相似文献   

4.
Reexamination of the Barremian–Aptian planktonic foraminifers from three sections (Verkhoirechie, mountain Krasnaya, and Marino) allowed the biostratigraphic scheme for Southwest and Central Crimea to be refined and updated. The following standard zones are recognized in the studied sections: Blowiella blowi (upper Barremian), Hedbergella excelsa (upper Barremian–lower Aptian), Leupoldina cabri (lower Aptian), H. luterbacheri, Globigerinelloides ferreolensis, Gl. barri, Gl. algerianus, Hedbergella trocoidea, Paraticinella rohri (upper Aptian). Beds with Hedbergella ruka are recognized in the B. blowi Zone. Foraminifers from the Partizanskoe section, representing the lower Aptian L. cabri and H. luterbacheri zones, are studied. The recognized strata are correlated with ammonite and nannoplankton zones and paleomagnetic data.  相似文献   

5.
It is now generally accepted that the Oceanic Anoxic Event 1a [OAE 1a] correlates with the lower part of the Leupoldina cabri planktonic foraminiferal Zone. Its calibration against the standard ammonite scale, however, seems to be more problematic. This is due, in part, to the fact that ammonites are scarce and/or of little diagnostic value from a biochronological viewpoint in the lower Aptian pelagic successions where the black shale horizons are better developed.We have been able to characterize OAE 1a geochemically in the relatively shallow water deposits of the eastern Iberian Chain (Maestrat Basin, eastern Spain), where ammonite faunas are rich. The interval corresponding to this event is dominated by the genera Roloboceras and Megatyloceras, accompanied by Deshayesites forbesi and Deshayesites gr. euglyphus/spathi. This assemblage is characteristic of the middle/upper part of the Deshayesites weissi Zone. The first occurrence of the species Deshayesites deshayesi (d'Orbigny), which marks the base of the overlying zone, takes place in our sections some metres above the OAE 1a interval.In the historical stratotype region of Cassis-La Bédoule (southern Provence Basin, southeastern France), the OAE 1a interval is also characterized by the presence of Roloboceras and Megatyloceras. Nevertheless, it has usually been correlated with the D. deshayesi Zone. In our opinion, this discrepancy is due to divergences in the taxonomic assignments of the deshayesitids present in these beds. In fact, the specimens attributed by French authors [Ropolo, P., Conte, G., Gonnet, R., Masse, J.P., Moullade, M., 2000. Les faunes d'Ammonites du Barrémien supérieur/Aptien inférieur (Bédoulien) dans la région stratotypique de Cassis-La Bédoule (SE France): état des connaissances et propositions pour une zonation par Ammonites du Bédoulien-type. Géologie Méditerranéenne 25, 167–175; Ropolo, P., Moullade, M., Gonnet, R., Conte, G., Tronchetti, G., 2006. The Deshayesitidae Stoyanov, 1949 (Ammonoidea) of the Aptian historical stratotype region at Cassis-La Bédoule (SE France), Carnets de Géologie / Notebooks on Geology Memoir 2006/01, 1–46.] to D. deshayesi and D. dechyi can be reinterpreted as belonging to D. forbesi.Following this reinterpretation, the Roloboceras beds (equivalent of OAE 1a) of Cassis-La Bédoule would also correspond to the D. weissi Zone. This age is additionally corroborated by data from southern England [Casey, R., 1961a. The stratigraphical palaeontology of the Lower Greensand. Palaeontology 3, 487–621; Casey, R., 1961b. A Monograph of the Ammonoidea of the Lower Greensand, part III. Palaeontographical Society, London, pp. 119–216], and by our recent observations in Le Teil (Ardèche Platform, southeastern France), where the Roloboceras faunas are also associated with Deshayesites consobrinus and Deshayesites gr. euglyphus, taxa that are characteristic of the D. weissi Zone.  相似文献   

6.
The Penninic Ocean was a side tract of the Central Atlantic Oceanic System intercalated between the European and the Austroalpine plates. Its closure started in the Early Cretaceous, as subduction of the oceanic crust beyond the Austroalpine plate. The sedimentary change on the Austroalpine shelf from pelagic carbonates into deep-water siliciclastics correlated with the denudation of the accretionary wedge resulting from that subduction. Within the Bajuvaric Unit of the Upper Austroalpine, this transition is reflected by the lithostratigraphic boundary between the older Schrambach and the younger Tannheim Formation. This boundary is well exposed in a newly discovered site at Sittendorf, southwest of Vienna. This new outcrop yields an extraordinarily rich planktonic foraminifera assemblage characterized by typical Aptian species belonging to Blowiella, Globigerinelloides, Hedbergella, Leupoldina, and Praehedbergella. A detailed biostratigraphic analysis based on thin-section investigations precisely dated the lithostratigraphic boundary within the lower part of the early Aptian Leupoldina cabri Acme Zone, having an approximate age of 123 Ma. Along with the biostratigraphic analyses, the gamma-log outcrop measurement was a powerful tool in interpreting the stratigraphy and the tectonic setting in the outcrop, which intersects one smaller-scale isoclinal fold.  相似文献   

7.
Palaeotemperatures during the late Barremian–early Aptian (Early Cretaceous) on the Russian Platform have been determined on the basis of oxygen isotope analysis of aragonitic bivalve molluscan and ammonoid shells and belemnite rostra with well-preserved microstructure from the Ulyanovsk area. Those obtained from the planispiral and heteromorph ammonoid shells from the lower Aptian Volgensis–Schilovkensis, Deshayesi–Tuberculatum, and Deshayesi–Renauxianum zones range from 26.7 to 33.2 °C, from 29.2 to 33.1 °C, and from 27.0 to 29.5 °C, respectively. A heteromorph Helicancylus? cf. philadelphius shell from the uppermost lower Aptian Bowerbanki Zone was secreted in highest temperature conditions (32.8–35.2 °C). In contrast, upper Barremian molluscs (bivalve Cyprina sp. and belemnite Oxyteuthis sp.) of the Ulyanovsk area show significantly lower palaeotemperatures: 16.9–18.5 °C and 7.9–17.8 °C, respectively, which is in accordance with known palaeogeographic and palaeobotanical evidences, showing that a distinct climatic optimum seems to have occurred during the late early Aptian, when warm Tethyan water penetrated into the basin. Marked changes in calculated growth temperatures for investigated molluscs from the Russian Platform were most likely connected with both the general warming trend during the late Barremian–early Aptian and local palaeonvironmental conditions. New data from the Bowerbanki Zone of the Russian Platform provide evidence on existence of the positive carbon isotope anomaly (2.4–6‰) at the end of the lower Aptian. There were apparently the three positive C-isotope anomalies during the late Barremian–early Aptian. The onset of mid early Aptian Oceanic Anoxic Event (OAE) 1a seems to coincide with both the beginning of significant warm conditions (followed by short-term cooling) and the abrupt decline in heavy carbon isotope concentrations in marine carbonates, which partly were the likely consequences of the intensive release of CO2 (biased by volcanic activity) and/or dissociation of methane gas hydrate.  相似文献   

8.
Sediments of Early Aptian age in Bulgaria can be assigned to four different facies: platform carbonates (Urgonian complex), shallow-water siliciclastics, hemipelagic and flyschoid siliciclastics. The taxonomic analysis of the ammonite faunas of 18 sections from these four different facies resulted in a revision of the existing ammonite zonation scheme so far applied in Bulgaria and adjoining areas. A new biostratigraphic scheme, which bridges the western and eastern Tethys, is thereby proposed for the Lower Aptian of Bulgaria.The Upper Barremian Martelites sarasini Zone is characterized in its upper part by the Pseudocrioceras waagenoides Subzone in the shallow-water sections and by a horizon with Turkmeniceras turkmenicum in the deep-water settings. The Upper Barremian/Lower Aptian boundary is fixed by the first appearance of Paradeshayesites oglanlensis. For the Lower Aptian the following ammonite zones were established (from bottom to top): The Paradeshayesites oglanlensis Zone, the Deshayesites forbesi Zone (= formerly Paradeshayesites weissi Zone) including the Roloboceras hambrovi Subzone in the upper part, the Deshayesites deshayesi Zone including the Paradeshayesites grandis Subzone in the upper part and the Dufrenoyia furcata Zone. The Lower–Middle Aptian boundary has been defined by the appearance of species belonging to the genera Epicheloniceras and Colombiceras.The Lower Aptian ammonite faunas of Bulgaria, allow an interregional correlation with other areas of the Tethyan Realm. The presence of Turkmeniceras in the Upper Barremian enables a correlation with the Transcaspian region, whereas Roloboceras, Koeneniceras and Volgoceratoides found in the middle part of the Lower Aptian are more typical representatives of the ammonite faunas in northern Europe (England, Germany, Volga region).The analysis of the ammonite successions in combination with sedimentological observations enable us to conclude that the marls and marly limestones of the Lower Aptian studied here also cover the interval of the Oceanic Anoxic Event 1a. An interval of thin-laminated clays, rich in organic matter, was identified in the upper part of the D. forbesi Zone (Roloboceras hambrovi Subzone). This interval is characterized by a total lack of benthic faunas.  相似文献   

9.
In 2009 two wells were drilled with 100% core recovery at Roquefort-La Bédoule (Bouches-du-Rhône, SE France), the historical Bedoulian stratotype. Here we present holostratigraphic results based on a detailed study of the cored sediments. Our work confirms that the La Bédoule area offers one of the best records for the period spanning the late Bedoulian, the anoxic event OAE1a and the Bedoulian/Gargasian (lower-upper Aptian substages) transition. New data provide a refined succession of micropaleontogical events already well correlated with ammonites from previous fieldwork and, thus, improve the cross-calibration of bioevents with high-resolution isotope stratigraphy. Methods of the quantitative micropalaeontology applied on benthic foraminifera such as tritaxias help testing their probable orbitally triggered cyclicity, which might be used to precise estimates of duration of events such as OAE1a, the Dufrenoya furcata ammonite Zone, the Globigerinelloides ferreolensis planktonic foraminiferal zone and the C7 isotopic stage.The lithologic, biotic and possibly isotopic changes seen at the level of and around bed 170 (top of “Niveau Blanc” sensu auctorum) are strong arguments to use this key-level as the boundary between the two Aptian substages (or stages in an alternative classification) and to support the proposal of La Bédoule as a potential locality for the GSSP of the Gargasian Substage (or of historical Aptian sensu stricto, in the alternative classification).  相似文献   

10.
Quantified organic-walled dinoflagellate cyst (dinocyst) assemblages are presented for two sedimentary successions deposited in neritic environments of the Tethys Ocean during the Barremian and Aptian in an attempt to reconcile established dinocyst biostratigraphic schemes for Tethyan and Austral regions. One section is at Angles, southeast France (the Barremian stratotype section); the other is at Deep Sea Drilling Project Site 263, off northwest Australia. We also construct a carbon isotope record for Site 263 using bulk organic carbon.Both sections contain abundant, well-preserved dinocyst assemblages. These are diverse, with 89 taxa identified at Angles and 103 taxa identified at Site 263. Of these, more than 93% are cosmopolitan. When combined with other work at Angles and Site 263, we found that nine dinocysts have their first occurrence (FO) or last occurrence (LO) at both locations. These dinocyst events are, in alphabetical order: LO of Cassiculosphaeridia magna, FO of Criboperidinium? tenuiceras, LO of Kleithriasphaeridium fasciatum, LO of Muderongia staurota, FO of Odontochitina operculata, LO of Phoberocysta neocomica, FO of Prolixosphaeridium parvispinum, FO of Pseudoceratium retusum var. securigerum, and FO of Tehamadinium sousense. Although these events support a Barremian–Aptian age for both sections, their stratigraphic order is not the same in the sections. The δ13Corg record at Site 263 displays a characteristic series of changes that have also been recorded in other carbon isotope curves spanning the Late Barremian–Early Aptian. Such independent dating (along with ammonite zones at Angles) suggests that three of the nine dinocyst events are approximately isochronous at Angles and Site 263: the LO of K. fasciatum in the mid Barremian, the FO of P. retusum var. securigerum and the FO of C.? tenuiceras in the earliest Aptian; the other six dinocyst events are diachronous. Dinocyst assemblages at Site 263 can be loosely placed within existing Australian zonation schemes, providing much-needed calibration. Our data suggest that the Muderongia testudinaria Zone ends in sediments of mid Barremian age, the succeeding Muderongia australis Zone extends into the Early Aptian, and the younger Odontochitina operculata Zone begins in Early Aptian deposits. The boundary between the M. australis and O. operculata zones, and the Ovoidinium cinctum (as Ascodinium) Subzone, positioned at the top of the M. australis Zone when present, could not be recognized incontrovertibly. Interestingly, however, this horizon broadly correlates with the onset and extent of the Selli Event, a time of major biogeochemical change.  相似文献   

11.
Neotrocholina Reichel, 1956 is one of the most important benthic foraminifera in Early Cretaceous. Some of the species of this genus are indexes in biostratigraphy especially for this interval (e.g., Neotrocholina friburgensis: Late Barremian–Early Aptian; Neotrocholina aptiensis: Early Aptian). In order to conduct accurate paleontological investigations, sampling from the carbonate units of the Tirgan Formation in Kopet Dagh sedimentary basin is done. According to the occurrence level in the studied stratigraphic sections as well as biometric interpretations, two species of this genus (N. friburgensis and Neotrocholina valdensis) are pointed out.  相似文献   

12.
Radiolarian biostratigraphic research has been carried out along two continuous sections through the Xialu Chert, one of the accreted sheets included in the Yarlung-Zangbo Suture Zone. Six radiolarian zones have been identified as follows: Laxtorum(?) jurassicum Zone (Aalenian), Tricolocapsa plicarum Zone (Bajocian–lower Bathonian), Stylocapsa(?) spiralis Zone (upper Callovian–Oxfordian), Hsuum maxwelli Zone (Kimmeridgian), Pseudodictyomitra carpatica Zone (upper Tithonian–lower Valanginian), and Turbocapsula costata Zone (Aptian).A reconstructed stratigraphy of the Xialu Chert, based on lithological succession and radiolarian dating, indicates that the chert exhibits a long depositional history, at least from early Middle Jurassic (Aalenian) to late Early Cretaceous (Aptian). The separation of the Lhasa Block from the northern Gondwana margin must, therefore, be dated before the Aalenian. The absence of calcareous sediments in the Xialu Chert indicates that the oceanic basin was deeper than the CCD throughout the depositional history. The transition from chert to siliceous mudstone is recorded sometime in Early Cretaceous, most probably around the Barremian/Aptian boundary. This means that the oceanic plate had already started being consumed at a trench by that time. The accretion of the Xialu Chert occurred after the Aptian time.  相似文献   

13.
Two cored boreholes in the central part of the North West German Basin recovered a unique section of Upper Barremian to Lower Aptian strata. Calcareous nannofossils show a distinctive shift from boreal endemic assemblages in the Barremian to cosmopolitan ones in the Aptian. This onset of new cosmopolitan species (e.g., Chiastozygus litterarius, Flabellites oblongus, Rhagodiscus angustus, Braarudosphaera sp., Eprolithus sp.) is spread over an interval of 25m, starting well below the early Aptian "Fischschiefer", a dark laminated shale rich in organic matter. These changes in the composition of calcareous nannofossils indicate that major palaeoceanographic changes occurred before the deposition of the Fischschiefer.The distribution pattern of calcispheres allows the differentiation of two sedimentary successions, separated by the Fischschiefer. The lower succession, which includes the sediments below the base of the Fischschiefer, indicates a boreal-pelagic environment. The lower part of the upper successions which includes the Fischschiefer, represents warmer inner shelf conditions. This corresponds to the presence of rich Tethyan-derived nannoconid assemblages and the presence of the planktonic foraminifera Hedbergella in the Fischschiefer. The topmost part of the upper succession (middle Aptian, Hedbergella marl) suggests a change to a pelagic warmer water environment. Two new taxa are introduced: the calcareous dinoflagellate cyst Obliquipithonella laqueata n. sp. and the foraminiferan Choanaella fortunate n. gen. n. sp.  相似文献   

14.
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15.
The Aptian/Albian boundary, as currently placed, presents problems which have not yet been solved and possibly never will be. No detailed definition of recent years can be applied on a world-wide scale, which will be needed for any definition acceptable to the Subcommission on Cretaceous Stratigraphy. The only satisfactory and workable solution is to choose a completely new position. It is suggested that this should be at the base of the Zone of Lyelliceras lyelli. This would change the start of the Albian from appproximately 112 Ma to 109 Ma but it would be a standard which should be recognizable on all five continents.  相似文献   

16.
《Geodinamica Acta》2013,26(5):349-361
The most widespread blocks within the Cretaceous ophiolitic mélange (North Anatolian ophiolitic mélange) in Central Anatolia (Turkey) are pillow basalts, radiolarites, other ophiolitic fragments and Jurassic-Cretaceous carbonate blocks. The pillow basalts crop out as discrete blocks in close relation to radiolarites and ophiolitic units in Cretaceous ophiolitic mélange.

The geochemical results suggest that analyzed pillow basalts are within-plate ocean island alkali basalts. The enrichment of incompatible elements (Nb, Ta, Light REE, Th, U, Cs, Rb, Ba, K) demonstrates the ocean island environment (both tholeiites and alkali basalts) and enriched MORB. Dated calcareous intrafills and biodetrital carbonates reveal an age span of Callovian—Early Aptian. The thin-shelled protoglobigerinids, belonging to the genus Globuligerina, in the calcareous intrafills between pillow basalt lobes indicates a Callovian—Barremian age interval, most probably, Valanginian to Late Barremian. The volcanic and radiolarite detritus-bearing orbitolinid—Baccinella biodetrital carbonates dated as Late Barremian-Early Aptian in age, were probably deposited around atolls and have a close relationship with the ocean island pillow basalts.

The results collectively support the presence of a seamount on the Neo-Tethyan oceanic crust during the Valanginian—Late Barremian and atolls during the Late Barremian-Early Aptian interval. The presence of an oceanic crust older than that seamount along the Northern Branch of Neo-Tethys is conformable with the geodynamic evolution of the Tethys.  相似文献   

17.
The chronostratigraphic framework of the non-marine deposits of the Central Tunisian Lower Cretaceous (Kebar Formation) is reviewed from a biostratigraphic viewpoint. The outcrops located in the Jebel Kebar, Jebel Ksaïra and Jebel Koumine localities provided charophyte assemblages belonging to two biochronozones: Ascidiella cruciata-Pseudoglobator paucibracteatus (upper Barremian–lower Aptian) and Clavator grovesii lusitanicus (upper Aptian–lower Albian). Clavatoraceans from the upper Barremian–lower Aptian in the Tunisian Atlas are reported here for the first time. The assemblage consists of Atopochara trivolvis var. triquetra, Ascidiella iberica var. inflata, Globator maillardii var. trochiliscoides, Globator maillardii var. biutricularis, Echinochara peckii var. lazarii, Clavator harrisii var. harrisii and Clavator harrisii var. reyi. In addition, a new characean species, Mesochara magna nov. sp. Trabelsi and Martín-Closas, is described herein. The results show that the Kebar Formation is diachronous in Central Tunisia, with a more complete record to the north (Jebel Koumine) than in the type locality (Jebel Kebar). Barremian–Aptian diapiric activity is proposed as the factor that controls the diachronous nature of this formation.The late Barremian–early Aptian charophyte assemblages from the Kebar Formation display strong affinities with the contemporaneous floras of the European basins, thus suggesting that intense supraregional floristic exchanges occurred between the Tethyan islands scattered throughout what is now Western Europe and North Africa. The biogeographic distribution of these charophytes leads to the hypothesis that the peri-Tethyan Archipelago acted as an effective bridge for the intercontinental exchanges of these plants between Laurasia and Gondwana.  相似文献   

18.
The 4th Kilian Group meeting (Dijon, France, 30th August 2010) focused on the Aptian and Albian Stages. For the Aptian, a two-fold division of the stage was adopted for the Mediterranean area with a boundary between the Dufrenoyia furcata and Epicheloniceras martini Zones. The main changes to the zonal scheme concern the Lower Aptian with: the introduction of a Deshayesites luppovi Subzone in the upper part of the Deshayesites oglanlensis Zone; the replacement of Deshayesites weissi by Deshayesites forbesi as new index-species of the second interval zone; the introduction of a Roloboceras hambrovi Subzone in the upper part of the D. forbesi Zone; and the subdivision of the D. furcata Zone into the D. furcata and Dufrenoyia dufrenoyi Subzones. For the Albian, the upper part of the Douvilleiceras mammillatum Zone (Lower Albian) is now characterized by a Lyelliceras pseudolyelli Subzone. The main amendments concern the Upper Albian. The base of this substage is defined by the base of the Dipoloceras cristatum Zone. Above it, the Upper Albian zonal scheme comprises in stratigraphic order the Mortoniceras pricei, Mortoniceras inflatum, Mortoniceras fallax, Mortoniceras rostratum, Mortoniceras perinflatum and Arrhaphoceras briacensis Zones.  相似文献   

19.
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20.
Prejanopterus curvirostra, from the Early Cretaceous of La Rioja province, was the first pterosaur genus and species described from Spain. The material comprises disarticulated cranial and postcranial remains from several individuals. The fossil-bearing bed is assigned to the lacustrine Leza Formation (eastern Cameros Basin, NW margin of the Iberian Range). This unit is regarded as either Berriasian-Valanginian or Barremian-Aptian. Prejanopterus curvirostris (specific name emended) was originally diagnosed on the basis of several characters of which the most significant was a lateral curvature of the rostrum. Re-examination of the holotype (rostrum) and paratype (partial rostrum with teeth) indicates that there is no genuine sideways bend of the preserved premaxilla-maxilla segments, but a slight dorsal curvature. Prejanopterus is characterized by a unique combination of characters: an emended diagnosis is provided. In contrast with previous estimates, the wing span of Prejanopterus was probably not much (if ever) in excess of 2 m. A phylogenetic analysis suggests that Prejanopterus is a basal pterodactyloid positioned between Pterodactylus and Cycnorhamphus-Gallodactylus. Prejanopterus represents the first evidence of Pterodactylidae in the Early Cretaceous of the Iberian Peninsula.  相似文献   

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