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1.
Fourteen midwater trawl collections to depths of 450 m to 1,400 m were taken at eleven stations in the Bering Sea and adjoining regions of the northern North Pacific by the R/V Hakuho Maru during the summer of 1975. A total of 29 kinds of fishes were identified. Mesopelagic fishes of the families Myctophidae, Gonostomatidae and Bathylagidae predominated in the catches, contributing 14 species (94%) of the fishes caught.Seventeen species of fishes were caught in the Bering Sea, and all of these are known from nearby areas. The mesopelagic fish fauna of the Bering Sea is similar to that in adjoining regions of the northern North Pacific Ocean: endemic species are rare or absent. Stenobrachius nannochir was usually the most common mesopelagic fish in our catches.Stenobrachius leucopsarus is a diel vertical migrant that is usually the dominant mesopelagic fish in modified Subarctic waters of the northeastern Pacific. The change in dominance fromS. nannochir in the western Bering Sea toS. leucopsarus in the eastern Bering Sea is related to differences in oceanographic conditions.  相似文献   

2.
The biomass, abundance, and vertical distribution of micronekton, including enidarians, mysids, euphausiids, decapods, thaliaceans, and fishes, were studied on the basis of samples collected with an 8-m2 opening-closing rectangular midwater trawl (RMT-8, mesh size: 4.5 mm) at three stations in the subarctic Pacific (the western subarctic gyre, the central Subarctic, and the Gulf of Alaska) and one station in the oceanic Bering Sea. The total biomass in the 0–1000 m water column ranged from 2.9 to 5.1 gDW m–2. Except for primary consumers that showed highly variable biomass (thaliaceans and euphausiids), biomass was highest in the oceanic Bering Sea followed by the central (boundary between eastern and western gyres), western gyre, and eastern Gulf of Alaska. The biomass compositions by higher taxa were basically similar between regions: fishes were most dominant, followed by enidarians at all stations, except for the marked predominance of thaliaceans in the Gulf of Alaska. High biomasses of gelatinous animals (31% of overall dry weight), occasionally comparable to those of fishes and crustaceans, suggest their potential importance in the subarctic Pacific. Characteristics in vertical patterns of micronekton biomass common in all stations were: (1) a mesopelagic peak around 500–600 m both day and night, (2) a layer of low biomass in the cold intermediate water and/or in the upper mesopelagic zone, (3) a nighttime shift of biomass to upper layers, and (4) an highly variable biomass of epipelagic/interzonal migrants (euphausiids and thaliaceans).  相似文献   

3.
根据第4次中国北极科学考察在白令海与楚科奇海进行的鱼类拖网调查资料,分析了白令海与楚科奇海鱼类生物的种类组成、优势种、物种多样性和区系特征,探讨了鱼类生物对北极气候快速变化的响应。结果表明,白令海与楚科奇海两个海域共鉴定鱼类生物14科41种;主要优势种类为粗壮拟庸鲽(Hippoglossoides robustus)、北鳕(Boreogadus saida)、短角床杜父鱼(Myoxocephalus scorpius)、斑鳍北鳚(Lumpenus fabricii)、粗糙钩杜父鱼(Artediellus scaber);从适温性来看,冷水性种类最多,有35种,冷温性种类6种;从栖息地生态类型来看,底层鱼类、近底层鱼类和中上层鱼类分别为35、5和1种;Shannon-Wiener多样性指数平均为1.21,呈现南高北低的特点,整体多样性水平不高;气候变化引起部分北极、亚北极海区鱼类出现不同程度的纬向和纵向移动,由此将引起北极渔业资源分布格局的变化。  相似文献   

4.
Bacterial biomass and production rate were measured in the surface (0–100 m) and mesopelagic layers (100–1,000 m) in the subarctic Pacific and the Bering Sea between July–September, 1997. Depth profiles were determined at stations occupied in oceanic domains including the subarctic gyres (western, Bering Sea, and Gulf of Alaska) and a boundary region south of the gyres. In the surface layer (0–100 m), both bacterial biomass and production were generally high in the western and Bering Sea gyres, with the tendency of decrease toward east. This geographic pattern was consistent with the dominant regime of phytoplankton biomass at the time of our survey. A significant portion of variation in bacterial production was explained by the concentration of chlorophyll a (r 2 = 0.340, n = 60, P < 0.001) and, to the greater extent, by the concentration of semilabile total organic carbon (SL-TOC = TOC at a given depth—TOC at 1,000 m, r 2 = 0.488, n = 59, P < 0.0001). Temperature significantly improved the regression model: temperature and chlorophyll jointly explained 60% of variation in bacterial production. These results support the hypothesis that bacteiral growth is largely regulated by the combination of temperature and the supply of dissolved organic carbon in subarctic surface waters. In the mesopelagic layer (100–1,000 m), the geographic pattern of bacterial production was strikingly different from the surface phytoplankton distribution: the production was high in the boundary region where the phytoplankton biomass was lowest. Bacterial growth appeared to be largely controlled by the supply of organic carbon, as indicated by the strong dependency of bacterial production on SL-TOC (r 2 = 0.753, n = 75, P < 0.0001). The spatial uncoupling between surface phytoplankton and mesopelagic bacterial production suggests that the supply rate of labile dissolved organic carbon in the mesopelagic zone does not simply reflect the magnitude of the particulate organic carbon flux in the subarctic Pacific.  相似文献   

5.
The mesopelagic community of fishes and squids are fundamental in the diet of apex predators, but in most cases their life histories and habitat requirements are poorly understood. In May 1999, a pilot study was conducted to identify mesopelagic nekton, describe dominant physical characteristics of their habitat, and compare their relative abundances over several study sites in the southeastern Bering Sea. Biological samples were collected at 250, 500, and 1000 m depths with an open pelagic rope trawl lined with 1.2-cm mesh in the codend. Net type, mesh size, and trawling techniques were designed to parallel those of extensive Russian research surveys in the western Bering Sea, permitting direct comparisons between study results. Forty-three species of fish and 15 species of cephalopods were identified, including a new species of gonatid squid and a range extension for Paraliparis paucidens, a snailfish never before observed in Alaskan waters. Faunal biomass was high with over 25,000 (1400 kg) fish and squid collected in only 13 trawls. Concentrations of fish in this area surpass published records from the western Bering Sea and North Pacific Ocean by an order of magnitude, driven primarily by Leuroglossus schmidti, a deep-sea smelt. Generally, specimens were of high quality, and new size records were established for several species of fish and squid. The physical environment as determined from altimetry, satellite-tracked drifters, and water properties (temperature and salinity) was typical of the last decade for this area. Spatial patterns in species distribution were observed, but further research is needed to determine whether these are a factor of mesoscale variability or of habitat characteristics.  相似文献   

6.
Abundances and biomasses of planktonic ciliates and copepod nauplii, major components of the microzooplankton community, were investigated in the subarctic North Pacific and the Bering Sea in summer of 1997. Their regional variation was illustrated by demarcating the entire area into five regions. Ciliates always predominated both in abundance (>94%) and biomass (>78%) over nauplii. Regional means of ciliates in the water column were higher in the Alaskan Gyre (120 × 106 cells/m2) and the Western Subarctic Gyre (110 × 106 cells/m2) in terms of abundance, and rich in the Bering Sea Gyre (360 mgC/m2) and the Western Subarctic Gyre (340 mgC/m2) in terms of biomass. By contrast, standing crops of ciliates were poor in the Oyashio Region (67 × 106 cells/m2; 170 mgC/m2) and the Transition Region (64 × 106 cells/m2; 160 mgC/m2). The values of biomass reported here are generally in agreement with the values reported previously from the Bering Sea Gyre and the Alaskan Gyre but are considerably higher than the previous value found in the Western Subarctic Gyre. No significant correlations could be found between chlorophyll a crop and standing crops of ciliates and copepod nauplii over the entire subarctic North Pacific and the Bering Sea during this summer.  相似文献   

7.
The food habits of the dominant myctophid Stenobrachius leucopsarus were examined in the central basin of the Bering Sea in relation to oceanographic conditions, in summer 2002 and 2003 and spring 2006. S. leucopsarus exhibited an ontogenetic and seasonal dietary shift. In spring, small fish (≤40 mm) preyed mainly on Neocalanus flemingeri/plumchrus whereas large fish fed mainly on Neocalanus cristatus. In summer, small fish preyed mainly on Metridia pacifica whereas large fish fed mainly on euphausiids (Thysanoessa spp.). In the summer of 2003, when water temperature in the epipelagic layer (≤100 m) was warmer, reflecting the prevalence of the Alaskan Stream, small-sized S. leucopsarus showed a higher stomach content index, perhaps reflecting the greater abundance of M. pacifica. Thus, the present study shows that the physical variability in the epipelagic layer affects not only diets but also feeding performance of micronekton.  相似文献   

8.
Using the data obtained from CTD stations and hydrochemical measurements (oxygen, silicates, and phosphates) performed by the Pacific Scientific Research Fishery Center (TINRO Center) in 2001–2004, vertical structures of water masses were considered for the western Bering Sea and for the deep-water depression of the Sea of Okhotsk. It was shown that definite values of the Si/P molar ratio were characteristic for the water mass boundaries within which linear relationships between these two elements were observed. The lower boundaries of cold intermediate layers in both seas are characterized by a value of Si/P = 23. The ratio for the main halocline (the layer of nutrient concentration jump) is equal to 32, while that for the intermediate layer is equal to 43 (47 in the Sea of Okhotsk). In the Bering Sea, linear relationships between the concentrations of these elements are determined by mixing of waters of different origin. The deep convection, regeneration of phosphates in the lower part of the surface layer, and the significant oxygen deficiency in the intermediate layer determine the doubled inclination of their ratio compared to the Redfield’s parameter. At the same time, in the Sea of Okhotsk, the determining role in linear relationships between the elements considered is played by the aeration of intermediate layer with near-bottom shelf waves, and by tidal mixing.  相似文献   

9.
A comparative analysis was conducted on climate variability in four sub-arctic seas: the Sea of Okhotsk, the Bering Sea shelf, the Labrador Sea, and the Barents Sea. Based on data from the NCEP/NCAR reanalysis, the focus was on air–sea interactions, which influence ice cover, ocean currents, mixing, and stratification on sub-seasonal to decadal time scales. The seasonal cycles of the area-weighted averages of sea-level pressure (SLP), surface air temperature (SAT) and heat fluxes show remarkable similarity among the four sub-arctic seas. With respect to variation in climate, all four seas experience changes of comparable magnitude on interannual to interdecadal time scales, but with different timing. Since 2000 warm SAT anomalies were found during most of the year in three of the four sub-arctic seas, with the exception of the Sea of Okhotsk. A seesaw (out of phase) pattern in winter SAT anomalies between the Labrador and the Barents Sea in the Atlantic sector is observed during the past 50 years before 2000; a similar type of co-variability between the Sea of Okhotsk and the Bering Sea shelf in the Pacific is only evident since 1970s. Recent positive anomalies of net heat flux are more prominent in winter and spring in the Pacific sectors, and in summer in the Atlantic sectors. There is a reduced magnitude in wind mixing in the Sea of Okhotsk since 1980, in the Barents Sea since 2000, and in early spring/late winter in the Bering Sea shelf since 1995. Reduced sea-ice areas are seen over three out of four (except the Sea of Okhotsk) sub-arctic seas in recent decades, particularly after 2000 based on combined in situ and satellite observations (HadISST). This analysis provides context for the pan-regional synthesis of the linkages between climate and marine ecosystems.  相似文献   

10.
Major features of four marine ecosystems were analyzed based on a broad range of fisheries-associated datasets and a suite of oceanographic surveys. The ecosystems analyzed included the Gulf of Maine/Georges Bank in the Northwest Atlantic Ocean, the Norwegian/Barents Seas in the Northeast Atlantic Ocean, and the eastern Bering Sea and the Gulf of Alaska in the Northeast Pacific Ocean. We examined survey trends in major fish abundances, total system fish biomass, and zooplankton biomasses. We standardized each time series and examined trends and anomalies over time, using both time series and cross-correlational statistical methods. We compared dynamics of functionally analogous species from each of these four ecosystems. Major commonalities among ecosystems included a relatively stable amount of total fish biomass and the importance of large calanoid copepods, small pelagic fishes and gadids. Some of the changes in these components were synchronous across ecosystems. Major differences between ecosystems included gradients in the magnitude of total fish biomass, commercial fish biomass, and the timing of major detected events. This work demonstrates the value of comparative analysis across a wide range of marine ecosystems, suggestive of very few but none-the-less detectable common features across all northern hemisphere ocean systems.  相似文献   

11.
Based on trawl surveys in the Bering Sea and Chukchi Sea during the 2010 Chinese National Arctic Research Expedition, fish biodiversity characteristics, such as fish composition, dominant species, biodiversity, and faunal characteristics were conducted. We also discussed the responses of fishes to the quick changes in Arctic climate. The results showed that a total of 41 species in 14 families were recorded in these waters. The dominant species were Hippoglossoides robustus, Boreogadus saida, Myoxocephalus scorpius, Lumpenus fabricii, and Artediellus scaber. There were 35 coldwater species, accounting for 85.37%, and six cold temperate species, occupying 14.63%. The habitat types of fish could be grouped as follows: 35 species of demersal fishes, five benthopelagic fishes, and one pelagic fish. The Shannon–Wiener diversity index(H)(range between 0 and 2.18, 1.21 on average) was not high, and descended from south to north. Climate change has caused some fishes to shift along their latitudinal and longitudinal distribution around the Arctic and Subarctic areas, and this could lead to the decline of Arctic fishery resources.  相似文献   

12.
The article summarizes and analyzes published data on the distribution of sea-ice and open-ocean diatoms in 42 cores of bottom sediments from the northwestern part of the Subarctic Pacific that accumulated during the last glacial maximum (LGM). Based on micropaleontological records, the extent of winter sea ice during the LGM could be limited to the Okhotsk and Bering seas. During the warm season, the surface water masses from the open Subarctic Pacific spread widely in the marginal seas.  相似文献   

13.
Size distributions of Neocalanus cristatus, N. flemingeri and N. plumchrus were investigated in the eastern and the western subarctic gyres and three marginal seas of the North Pacific during the diapause period to examine the geographical variation in body size of Neocalanus species and to clarify the origin of the large biennial N. flemingeri which has been observed in the Oyashio region. There were significant among region variations in body sizes for all three species of Neocalanus. Generally, the body sizes of the copepods were larger in the marginal seas and marginal areas of the open ocean. In the open ocean, the body sizes increased westward. These patterns of variation in the body sizes roughly correlated with local food availability. Distribution of biennial N. flemingeri was restricted to the Sea of Japan, the Okhotsk Sea and the Oyashio region. The large-sized biennial N. flemingeri were abundantly observed in the Okhotsk Sea, and the medium-sized biennial individuals were observed in the Sea of Japan. These facts strongly suggest that the large biennial N. flemingeri in the Oyashio region are advected from the Okhotsk Sea.  相似文献   

14.
Results of the long-term study of the spatial and vertical distribution of the parasitic anadromous Arctic lamprey Lethenteron camtschaticum (Tilesius, 1811) (Petromyzontidae) in the North Pacific and data on its size composition are given. This species is most frequent in the northwestern Sea of Japan and the western Bering Sea. The maximum concentrations are noted in waters of southern Primorye, southwestern Sakhalin, the northwestern part of the Sea of Okhotsk, and the northern part of the Bering Sea, which is probably explained by the increased number of its victims, Pacific salmon. Near the bottom, Pacific lampreys are extremely few and are primarily encountered at depths less that 400 m, and in the Pelagic zone, in the 100-m layer. The catches have contained Arctic lampreys having a total length of 15–79 cm. The lampreys of several size groups in the catches may indicate that L. camtschaticum spends not less than four years in the sea. No relationship has been found between the body length and the capture depth. Analyzed are the relationships between the body length and weight and the body length and the condition factor. The seasonal dynamics of these indices are considered.  相似文献   

15.
Vertical profiles of tritium in seawater were determined for samples collected during the period from 1988 to 1990 at fourteen stations in the northwestern North Pacific (the Oyashio region) including the Okhotsk Sea and the Bering Sea. The profiles usually had a maximum in the surface layer and decreased gradually with depth down to 1,000 m. The water column inventory of tritium averaged 63% of the total atmospheric input in this region.The horizontal distribution of tritium showed a maximum in the region facing the Okhotsk Sea near 45°N for every isopycnal surface of 0 ranging from 26.60 to 27.40. The ages of the intermediate water were calculated for the respective isopycnal surfaces in the maximum region. This calculation assumed that the intermediate water was formed by the isopycnal mixing of two water masses—the Okhotsk Sea and the Bering Sea Component Waters, which had been produced in wintertime by the diapycnal mixing of the surface and the deep waters in the respective marginal seas. The results show that the intermediate water in this region was formed in the late 1980's for the water which has 0 of 26.60 to 26.80 and about 1970 for the water which has 0 of 27.00 to 27.40. Although we have estimated the mean ages of the intermediate water, the horizontal profile of dissolved oxygen suggests that the Okhotsk Sea Component Water is younger than the mean age.  相似文献   

16.
The sea-surface bioproductivity changes over the last 25 kyr were inferred from published data on 30 sediment cores from the open Northwest Pacific (NWP), Sea of Okhotsk, Bering Sea and Sea of Japan accounting for the glacioeustatic sea-level changes. A novel method was developed to compare the variations of several independent productivity proxies relative to the present-day values. During the Last Glacial Maximum, the bioproductivity in the Sea of Okhotsk and the western Bering Sea (BS) was lower than at present, whereas the southern and southeastern Bering Sea and the open NWP are characterized by enhanced bioproductivity. During the early deglacial stage, an increase in bioproductivity was estimated only for the southeastern Bering Sea. High and fairly high bioproductivity was estimated for Heinrich 1 in the open NWP, above the Umnak Plateau and on the Shirshov and Bowers Ridges in the Bering Sea. The high productivity in the Bering Sea, Sea of Okhotsk and NWP during the Bølling/Allerød was caused by the global warming and enhanced nutrient supply by meltwater from the continent. During the Early Holocene, high productivity was estimated for almost the entire NWP. The Late Holocene sea-surface bioproductivity was generally lower than that of the Early Holocene. Proposed factors that have controlled the sea-surface bioproductivity during the last 25 kyr include: the location of the sea ice margin, the river runoff, gradual flooding of the Bering Sea and the Sea of Okhotsk shelf areas, the water mass exchange between the marginal seas and the open NWP, the eolian supply and the deep vertical mixing of the water column.  相似文献   

17.
Processes relating to the formation of dense shelf water and intermediate water in the Okhotsk Sea were studied by examining oxygen isotope ratios (δ18O), salinity, and temperature. The salinity and δ18O of the cold dense shelf water on the northern continental shelf showed peculiar relationship. The relationship indicates that 3% of the mixed-layer water, having salinity of 32.6, froze and the remaining 97% became dense shelf water of salinities of more than 33.2 (σθ>26.7) during the sea ice formation. The salinity–δ18O relationship also shows that 20% of the Okhotsk Sea Intermediate Water at the σθ=26.8 level was derived from the dense shelf water. The remaining 80% came from the Western Subarctic Pacific water modified by diapycnal mixing of water affected by the surface cooling and freshening within the Okhotsk Sea. The mixing with dense shelf water contributes to only 26% of the temperature difference or 8% of the salinity difference between the original Pacific water and the Okhotsk Sea Intermediate Water at σθ=26.8. This result suggests that the cold and less saline properties of the Okhotsk Sea Intermediate Water are produced mainly by diapycnal mixing, rather than by mixing of the Pacific water with the dense shelf water.  相似文献   

18.
Basing upon the total of 1,081 samples collected by a large plankton net (160 cm in mouth diameter) in the Pacific Ocean, the geographical and vertical distribution ofGonostoma gracile were studied. The species is distributed in the water masses of the Kuroshio, the Kuroshio Extension, the North Pacific Current, the Oyashio, the North Pacific Subarctic Water and the western North Pacific Central Water. The center of distribution lies in the Kuroshio area off Japan. Vertically, the species occurs between the depths of 200 and 1,000 m, mainly 300–700 m, both during daytime and at night. A part of population might come up to 0–200 m at night, although its biomass is negligibly small in comparison to that remaining in 300–700 m layer. Postlarvae are found in 300–700 m layer, mainly at 300–500 m, and apparently do not undertake diurnal vertical migration.  相似文献   

19.
We survey the recent progress in studies of North Pacific Intermediate Water (NPIW) in SAGE (SubArctic Gyre Experiment), including important results obtained from related projects. Intensive observations have provided the transport distributions relating to NPIW and revealed the existence of the cross-wind-driven gyre Oyashio water transport that flows directly from the subarctic to subtropical gyres through the western boundary current as well as the diffusive contribution across the subarctic front. The anthropogenic CO2 transport into NPIW has been estimated. The northern part of NPIW in the Transition Domain east of Japan is transported to the Gulf of Alaska, feeding the mesothermal (intermediate temperature maximum) structure in the North Pacific subarctic region where deep convection is restricted by the strong halocline maintained by the warm and salty water transport originating from NPIW. This heat and salt transport is mostly balanced by the cooling and freshening in the formation of dense shelf water accompanied by sea-ice formation and convection in the Okhotsk Sea. Intensive observational and modeling studies have substantially altered our view of the intermediate-depth circulation in the North Pacific. NPIW circulations are related to diapycnal-meridional overturning, generated around the Okhotsk Sea due to tide-induced diapycnal mixing and dense shelf water formation accompanied by sea-ice formation in the Okhotsk Sea. This overturning circulation may possibly explain the direct cross-gyre transport through the Oyashio along the western boundary from the subarctic to subtropical gyres. This revised version was published online in July 2006 with corrections to the Cover Date.  相似文献   

20.
A numerical study using a 3-D nonhydrostatic model has been applied to baroclinic processes generated by the K 1 tidal flow in and around the Kuril Straits. The result shows that large-amplitude unsteady lee waves are generated and cause intense diapycnal mixing all along the Kuril Island Chain to levels of a maximum diapycnal diffusivity exceeding 103 cm2s−1. Significant water transformation by the vigorous mixing in shallow regions produces the distinct density and potential vorticity (PV) fronts along the Island Chain. The pinched-off eddies that arise and move away from the fronts have the ability to transport a large amount of mixed water (∼14 Sv) to the offshore regions, roughly half being directed to the North Pacific. These features are consistent with recent satellite imagery and in-situ observations, suggesting that diapycnal mixing within the vicinity of the Kuril Islands has a greater impact than was previously supposed on the Okhotsk Sea and the North Pacific. To examine this influence of tidal processes at the Kurils on circulations in the neighboring two basins, another numerical experiment was conducted using an ocean general circulation model with inclusion of tidal mixing along the islands, which gives a better representation of the Okhotsk Sea Mode Water than in the case without the tidal mixing. This is mainly attributed to the added effect of a significant upward salt flux into the surface layer due to tidal mixing in the Kuril Straits, which is subsequently transported to the interior region of the Okhotsk Sea. With a saline flux into the surface layer, cooling in winter in the northern part of the Okhotsk Sea can produce heavier water and thus enhance subduction, which is capable of reproducing a realistic Okhotsk Sea Mode Water. The associated low PV flux from the Kuril Straits to the open North Pacific excites the 2nd baroclinic-mode Kelvin and Rossby waves in addition to the 1st mode. Interestingly, the meridional overturning in the North Pacific is strengthened as a result of the dynamical adjustment caused by these waves, leading to a more realistic reproduction of the North Pacific Intermediate Water (NPIW) than in the case without tidal mixing. Accordingly, the joint effect of tidally-induced transport and transformation dominating in the Kuril Straits and subsequent eddy-transport is considered to play an important role in the ventilation of both the Okhotsk Sea and the North Pacific Ocean. This revised version was published online in July 2006 with corrections to the Cover Date.  相似文献   

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