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1.
The marine sediments of the area of Verde Peninsula-Jabali Island(39°28′S/62°19′W-40°28′S/62°11′W) Holocene in age(3-2 ky),and modern beaches contain a significant amount of bioeroded mollusc shells.Fifteen sites were analyzed,in which 20.11%of the mollusc shells(2168 valves) presented bioerosion traces,in 54 species(30 bivalves and 24 gastropods).Fourteen ichnogenera were reported:Entobia,Maeandropoiydora,Iramena,Caulostrepsis,Pennatichnus,Pinaceocladichnus,Trypanites,and Gastrochaenolites(Domichnia),Gnathichnus and Radulichnus(Pascichnia),Finichnus and Centrichnus(Fixichnia),Oichnus(Praedicnia)(macrobioerosion),y Semidendrina(microbioerosion),the latter is first reported in mollusc shells in Argentina.Eleven ichnospecies were identified Finichnus peristroma,Maeandropoiydora sulcans,Gnathichnus pentax,Pinaceocladichnus onubensis,Caulostrepsis taeniola,Centrichnus eccentricus,Radulichnus inopinatus,Oichnus simplex,Oichnus paraboloides,Oichnus gradatus,and Gastrochaenolites torpedo(lithic remains).The dominant ichnogenera in the Holocene deposits are Iramena,Entobia and Oichnus.The same ichnogenera are constant with different abundance in the modern beaches,and increasing representation of Pinaceocladichnus and Pennatichnus.The dominant ichnofacies in the Holocene deposits is Trypanites,revealing a benthonic marine community composed of cheilostome bryzoans,clionaid sponges,predator gastropods,regular echinoids,polychaete annelids,bivalves,thallophytas and fungi.Generally,the area was described as a sublittoral,low-energy,stable environment with high rate of oxygenation,and sandy bottoms,with rocky bottoms at Villalonga locality.  相似文献   

2.
The ecological significance of algal and seagrass wrack subsidies has been well-documented for exposed-coast sandy beaches but is relatively unstudied in lower-energy and mixed-sediment beaches. In marine nearshore environments where beaches are fringed with riparian vegetation, the potential for reciprocal subsidies between marine and terrestrial ecosystems exists. Within the marine-terrestrial ecotone, upper intertidal “wrack zones” accumulate organic debris from algae, seagrass, and terrestrial plant sources and provide food and shelter for many organisms. Human modification also occurs within this ecotone, particularly in the form of armoring structures for bank stabilization that physically disrupt the connectivity between ecosystems. We conducted detailed wrack and log surveys in spring and fall over 3 years at 29 armored–unarmored beach pairs in Puget Sound, WA, USA. Armoring lowered the elevation of the interface between marine and terrestrial ecosystems and narrowed the width of the intertidal transition zone. Armored beaches had substantially less wrack overall and a lower proportion of terrestrial plant material, while marine riparian zones (especially trees overhanging the beach) were an important source of wrack to unarmored beaches. Armored beaches also had far fewer logs in this transition zone. Thus, they lacked biogenic habitat provided by logs and riparian wrack as well as the organic input used by wrack consumers. Results such as these that demonstrate armoring-associated loss of connectivity across the marine-terrestrial ecotone may be useful in informing conservation, restoration, and management actions.  相似文献   

3.
Sandy beaches are characterised by heterogeneously distributed food sources both in time and space. The major energy supply is derived from marine subsidies in the form of beach-cast macrophyte wrack. Wrack patches are short-lived, and their position on the beaches varies with tidal and seasonal cycles as well as weather conditions. Little is known about how sandy beach inhabitants orient themselves towards, and colonise, wrack patches. In a series of field studies on islands off Vancouver Island (British Columbia, Canada), colonisation patterns of wrack patches by beach fleas and sand hoppers (Amphipoda: Talitridae), the most abundant macrofaunal detritivores, were studied. As indicated by colonisation of experimental patches of wrack and wrack surrogates that were either visible or buried in the sand, beach fleas (Traskorchestia traskiana) rely on olfactory cues for locating freshly deposited wrack patches in their patchy and dynamic habitat. Dense colonisation of freshly deposited algal wrack generally occurred within less than 1 h but depended upon the tidal height of wrack patches. Beach flea density in freshly deposited wrack patches increased with increasing tidal height. By contrast, sand hoppers (Megalorchestia californiana) colonised freshly deposited wrack patches in densities that decreased with tidal height. Discussing these interspecific differences, we provide a primer for future detailed studies on transport of matter along the marine–terrestrial gradient of sand beaches.  相似文献   

4.
We investigated the role of sandy beaches in nearshore nutrient cycling by quantifying macrophyte wrack inputs and examining relationships between wrack accumulation and pore water nutrients during the summer dry season. Macrophyte inputs, primarily giant kelp Macrocystis pyrifera, exceeded 2.3 kg m−1 day−1. Mean wrack biomass varied 100-fold among beaches (range = 0.41 to 46.43 kg m−1). Mean concentrations of dissolved inorganic nitrogen (DIN), primarily NOx−-N, and dissolved organic nitrogen (DON) in intertidal pore water varied significantly among beaches (ranges = 1 to 6,553 μM and 7 to 2,006 μM, respectively). Intertidal DIN and DON concentrations were significantly correlated with wrack biomass. Surf zone concentrations of DIN were also strongly correlated with wrack biomass and with intertidal DIN, suggesting export of nutrients from re-mineralized wrack. Our results suggest beach ecosystems can process and re-mineralize substantial organic inputs and accumulate dissolved nutrients, which are subsequently available to nearshore waters and primary producers.  相似文献   

5.
The Atlantic ribbed mussel, Geukensia demissa, is found in salt marshes along the North American Atlantic Coast. As a first step to study the possibility of future cultivation and harvest of ribbed mussels for nutrient removal from eutrophic urban environments, the feeding behavior of ribbed mussels in situ was studied from July to October 2011. Two locations approximately 80 km apart were used as study sites: Milford Harbor (Connecticut; 41°12′42.46″N, 73°3′7.75″W) and Hunts Point (Bronx, New York; 40°48′5.99″N, 73°52′17.76″W). Total particulate matter was higher at Hunts Point than at Milford Harbor, but the organic content was higher at Milford than at Hunts Point. The relatively low quantity of organic content in Hunts Point seston resulted in a much higher production of pseudofeces by mussels. Mussel clearance and absorption rates were higher at Milford Harbor than at Hunts Point. Nevertheless, mussels at both sites had the same absorption efficiency, suggesting that mussels are able to adapt to conditions at both locations. Ribbed mussels decreased clearance rate when the seston quantity was high at both sites. At Hunts Point, ribbed mussels increased the gut transit time of ingested particles when the amount of inorganic particulates in the water increased. This study does not quantify nutrient removal capacity of G. demissa; however, the environmental tolerance demonstrated here, and current lack of commercial harvest, suggests that this species may be a good candidate for nutrient bioextraction in highly impacted urban environments.  相似文献   

6.
Sandy beach surf zones serve as alternative nursery habitats for juvenile Chinook salmon (0 age) during their early marine residency, a period considered critical due to high and variable mortality rates. Despite the importance of early marine residence, the extent of juvenile salmon surf zone use and movement along sandy beaches is not well understood. Juvenile Chinook salmon distribution and movement were studied in shallow surf zone habitats by sampling from 2006 to 2010 with a beach seine 11 beaches adjacent and distant to four estuary mouths in Oregon and Washington, USA. The estuary of origin of each juvenile was determined using genetic stock identification methods and coded wire tags. Surf zones sampled were within littoral cells, which are stretches of the coastline bordered by rocky headlands, and included estuaries with and without Chinook salmon populations. Juvenile salmonids were only collected at littoral cells with Chinook-inhabited watersheds. Most juveniles (95 %) were present at sandy beaches adjacent (<500 m from estuary mouth) to their estuary of origin. Few Chinook salmon (5 %) were collected at littoral cells that contained non-natal estuaries. These results indicate that juvenile Chinook salmon inhabiting surf zones mostly use beaches adjacent to their estuaries of origin, but some juveniles may reside in beaches distant from their point of ocean entry.  相似文献   

7.
A detailed survey of the upper and middle Nova Scotian continental slope at 42°50′N and 63°30′W indicates a complex morphology dominated by mass movements on various scales and an immature turbidity current channel. The range of sediment facies is diverse including hemipelagic and turbidite muds, turbidite sands and gravelly sandy muds of debris flow origin. Deformed units, interpreted as slump deposits are also observed. Several facies associations, related to discrete morphological environments, are recognized. Thick turbidite sand units with minor intervening mud beds are characteristic of the high-relief uppermost slope and channel margin. Thinner turbidite sands, deformed slump beds and various mud facies are associated with small-scale, hummocky mid-slope topography. Sand beds are more abundant in the depressions than on intervening hummocks indicating the preferred transport paths of small turbidity currents. At the lower end of the main turbidity current channel, frequent turbidite sand beds with relatively minor mud beds are deposited on a depositional lobe. In areas unaffected by mass movements, alternating bioturbated mud and sandy muds make up the core sequences. A local model of sedimentation is proposed for this area and illustrates that simple models of continental slope sedimentation only apply to a limited range of settings.  相似文献   

8.
We apply Fourier and wavelet analyses to the precipitation and sunspot numbers in the time series (1901–2000) over Australia (27°S, 133°E), Canada (60°N, 95°W), Ethiopia (8°N, 38°E), Greenland (72°N, 40°W), United Kingdom (54°N, 2°W), India (20°N, 77°E), Iceland (65°N, 18°W), Japan (36°N, 138°E), United States (38°N, 97°W), South Africa (29°S, 24°E) and Russia (60°N, 100°E). Correlation analyses were also performed to find any relation among precipitation, sunspot numbers, temperature, and cloud-cover at the same spatial and temporal scale. Further correlations were also performed between precipitation with electron and proton fluence at the time interval, 1987–2006. All these parameters were considered in annual and seasonal scales. Though correlation study between precipitation and other parameters do not hint any linear relation, still the Fourier and wavelet analyses give an idea of common periodicities. The 9–11 year periodicity of sunspot numbers calculated by Fourier transform is also confirmed by wavelet transform in annual scale. Similarly, wavelet analysis for precipitation also supports the short periods at 2–5 years which is verified by Fourier transform in discontinuous time over different geographic regions.  相似文献   

9.
We experimentally evaluate the effect of migratory shorebirds on the benthic fauna of three southwestern Atlantic Argentinean stop-over and wintering sites: Bahía Samborombon (35°30′–36°22′S, 57°23′W), Mar Chiquita coastal lagoon (37°40′S, 57°26′W), and Bahía Blanca (38°48′–39°25′S, 50°–62°25′W). The experiments consisted of exclusion ceilings and controls (both 1 m2), with 10 replicates each and aligned at the same tidal level. During December 1994–May 1995 these experiments were done, twice in Bahía Samborombon, twice at two sites (Sotelo and Celpa) 4 km apart in Mar Chiquita coastal lagoon, and three times in Bahía Blanca. Three polychaete species (Laeonereis acuta, Nephtys fluviatilis, andHeteromastus similis) were found in similar densities in both areas of the Mar Chiquita lagoon.L. acuta andN. fluviatilis were affected in Sotelo, but there was no treatment effect in Celpa. In Sotelo the most abundant shorebirds were Hudsonian godwit (Limosa haemastica), American golden plover (Pluvialis dominica), and White-rumped sandpiper (Calidris fuscicollis), and in Celpa the most abundant shorebirds were White-rumped sandpiper, two-banded plower (Charadrius falklandicus), and lesser yellowlegs (Tringa flavipes). The largest polychaete densities were in the upper sediment layer (0–6 cm), which was also the most affected layer. Three polychaete species were found in Bahía Samborombon (L. acuta, H. similis, andNeanthes succinea), but onlyH. similis showed a treatment effect. The most abundant shorebirds in Samborombon were white-rumped sandpiper and two-banded plover. No effect was detected in Bahía Blanca, where the most abundant shorebirds were white-rumped sandpiper and American golden plover. In all cases, the species affected by shorebirds were the most abundant species. From the two sites of Mar Chiquita, there was a treatment effect only in Sotelo, which was also the area with higher shorebirds counts. However, much lower densities observed in Samborombon (similar to the unaffected area of Mar Chiquita) also produced a significant decrease in infaunal abundance. These evidences also suggest that just selection of study sites may produce different views of the interaction between shorebirds and benthic species.  相似文献   

10.
The American horseshoe crab, Limulus polyphemus, is found along the Atlantic and Gulf of Mexico coasts in genetically isolated populations. Eggs are laid in shoreline beaches that expose developing embryos to combinations of environmental stressors. Whether populations of L. polyphemus differ in multi-stressor tolerance had never been tested. We assessed the multi-stressor tolerance of L. polyphemus embryos from a population in Delaware Bay (DE) and determined whether these differed from the multi-stressor tolerance of embryos from a more southerly Florida Gulf Coast (FGC) population. We monitored the field sediment temperatures and determined multi-stressor tolerance of DE embryos, then compared these to published data for FGC embryos. For multi-stressor tolerance, we assessed development success of embryos in 2-week exposures to 36 full-factorial combinations of temperature (20, 25, 30, 35 °C), salinity (5, 15, and 34 ppt), and ambient O2 (5, 13, and 21% O2), followed by 2 weeks in recovery conditions. Sediment temperatures in the DE site ranged from 9.5 to 46 °C, with extended periods exceeding 35 °C. Development success was similar between the DE and FGC populations in 14 of 26 multi-stressor combinations. The DE embryos were generally more successful in conditions that included high temperature or moderate hyposalinity, whereas the FGC embryos were generally more successful in conditions that included extreme hyposalinity. This suggests that although multi-stressor tolerances are generally similar between the two populations, specific differences exist that correlate more with differences in nest microenvironment than latitude.  相似文献   

11.
The authors have made a comparative study of surface deformations deduced from the analysis of recent faults, and deformation in depth deduced from, focal mechanisms associated with shallow earthquakes. The initial results obtained in the Paphos region of Cyprus are presented.Study of striations on centi- to decametric-scale fracture surfaces which affect Plio-Quaternary beaches in the Paphos region has enabled the authors to determine statistically the position of the principal axes of Plio-Quaternary deformation: lengthening X (strike N32°, dip 12°NE), shortening Z (N297°, 23°NW) and intermediate Y (N138°, 65°SE).On the 10th of September 1953 a superficial earthquake (depth 6 km) of 6.5 magnitude occurred near Paphos (location 34.9°N, 32.2°E) whose focal mechanism has been studied. Fault-plane solutions were obtained using 36 data points relating to the initial movements of the P-waves measured by longand short-period seismographs. Unfortunately the distribution of data with respect to the focus is such that it does not permit a single, but rather three possible solutions. Two of these are compatible with the model of deformation deduced from surface study. While the most probable solution favours a normal fault (solution no. 3), the system of deep strike-slip faults (solution no. 2) is equally compatible with the existence of predominantly normal surface faults.The preliminary results constitute part of a more regional study of the eastern Mediterranean. The authors consider it fundamental to establish this type of correlation when studying neotectonics.  相似文献   

12.
The characteristics of the recent tsunami deposits threw light on some sources of deposition on the beach of the Red Sea. The studied area is delineated by latitudes 25°6′ N and 25°9′ N and longitudes 34°50′ E and 34°53′ E; it covers an elongate area of about 12 km2 along the Red Sea coast, North Marsa Alam City (4 km). The area is bounded by Wadi Asalay to the north and Wadi Sifayn from the south. For evaluation of the area, a lot of information allows us to interpret the conditions prevailing during deposition of the sediments especially at the coast. To achieve the target, five wells were drilled to study core samples, well logging measurements, and 69 vertical electrical sounding stations were carried out. The studied area and adjacent areas were geologically surveyed to note geological features related to Paleo-earthquakes. From geological and geophysical studies, the dominant rock types at the western portions of the studied area are sandstone, sandy clay, clay, clayey sandstone, and gravels; at the middle portion of the studied area, the rocks are hard, but the eastern side of the area, especially at the beach of the Red Sea, several cycles of depositions of coral reefs occurred with intercalations of clastic deposits such as clay, sand, sandstone, conglomerate, gravels, pebbles, and a lot of fossils and shell fragments. The rocks are characterized by heterogeneous properties and ill-sorted. The area includes large numbers of faults due to highly tectonism of the area. The results indicated that the area has lateral variation of sediments. The carbonate rocks at the beach contain clastic fragments, and carbonate blocks are included within clastic rocks. With increasing the distance from the beach to the west, the sediments are less heterogeneous. The beach of the Red Sea was subjected to Paleo-tsunami waves due to highly Paleo-seismic activity inside the Red Sea and left their signature in geological column especially at the beach. The observation of some geological features such as Paleo-liquefaction and landslides indicate that the area subjected to strong earthquakes related to rifting of the Red Sea.  相似文献   

13.
《Geochimica et cosmochimica acta》1999,63(23-24):4005-4012
Previous studies have shown that 187Os/188Os in seawater has become increasingly radiogenic over the last 40 Ma in a manner analogous to strontium. This rapid rise in the marine 187Os/188Os over the last 17 Ma has been attributed to an increase in the bulk silicate weathering rates resulting from the rise of the Himalayas and/or selective weathering and erosion of highly radiogenic organic rich ancient sediments. The key test of this hypothesis is the 187Os/188Os and the total osmium concentration of the Himalayan rivers. We report the concentration and isotopic composition of osmium in the Ganges, the Brahmaputra, and the Indus rivers. The 187Os/188Os of the Ganges close to its source (at Kaudiyal, 30°05′N, 78°50′E) is 2.65 and [Os] = 45 fM/kg. A second sample of the lower reaches of the Ganges at Patna (25°30′N, 85°10′E) gives 187Os/188Os =1.59 and [Os] = 171 fM/kg. The 187Os/188Os of the Brahmaputra at Guwahati (26°10′N, 91°58′E) is 1.07 and [Os] = 52 fM/kg. A sample of the Indus (Besham, 34°55′N, 72°51′E) has a 187Os/188Os of 1.2 and [Os] = 59 fM/kg. We infer that the Himalayas do not provide either a high flow of osmium or a highly radiogenic osmium component to the oceans. The overall trend for osmium and strontium could be explained by a regularly increasing input of global continental weathering sources but the Himalayas themselves appear not to be the dominant source.  相似文献   

14.
Pollen and dinoflagellate cysts from marine sediments in core A180-48 (15°19′N, 18°06′W; 2450 m water depth; 530 cm length) are used to reconstruct palaeoenvironmental conditions of nearshore tropical west Africa during the last deglaciation. High concentrations and influxes of pollen and dinoflagellate cysts between 11 000 and 10 000 yr BP are interpreted as reflecting an increase in continental trade-wind circulation and related coastal upwelling at 15°N latitude. The sea-surface temperature difference between glacial and interglacial times was not as strong as previously suggested. Together with local (fresh) ground-water input, this smaller temperature difference may explain the persistence of the Rhizophora mangrove and Guinean gallery forests near the shore until their massive extension during the early Holocene humid maximum around 9500 yr BP. Pollen data from the core are compared with data from Rosilda N110-Z, from the continental shelf at the same latitude.  相似文献   

15.
Characteristics of Beachrocks: A Review   总被引：1，自引：0，他引：1  
Beachrocks are calcareous rocks that have been quite recently formed and are found in the intertidal zone on sandy beaches. The authors are developing an artificial rock with self-healing capabilities that can be used in coastal areas to repair the infrastructure of coastal walls. Because beachrocks are formed naturally on beaches, they merit study as a model for artificial rocks. We conducted a bibliographical study in order to gather information about the characteristics of beachrocks, which may aid in the manufacturing of artificial rocks. This study investigated their geographic distribution, formative periods, physical properties, and origins. The study found that over 90 % of beachrocks are found between the latitudes 40°N and the Tropic of Capricorn, and that their formative periods range from 26,000 years to just a few decades ago. We also learned that the cement component of beachrocks mainly consists of either calcium carbonate or silica. The dynamic characteristics of these components have been studied in only three papers, which note that their uniaxial compression strength ranges from as low as 6.0 to 42.0 MPa. Since very limited data are available on beachrocks in Japan where the artificial rock technology will first be deployed, there is a need for collecting additional data on their mechanical properties, elemental composition ratios, and mineral content.  相似文献   

16.
This study is focused on the analyses of a Chaschuil section (27° 49′ S–68° 04′ W), north of the Argentina Famatina Belt, where Ordovician explosive-effusive arc volcanism took place under subaerial to subaqueous marine conditions. In analyzing the profile, we have recognized an Arenigian succession composed by dominant volcaniclastic lithofacies represented by volcaniclastic debris flow, turbidity current and minor resedimented syn-eruptive pyroclastic depositsand lavas. The upper portions of succession are represented by volcanogenic sedimentary lithofacies with fossiliferous levels. Great volumes of the volcaniclastic deposits are strongly controlled in their transport by mass flow processes. These representative deposits provide significant data in relation to the coeval volcanic events for recognizing a continuous explosive volcanism together a minor effusive activity and the degradation of volcanic edifices. Likewise mass flow deposits give indications of the high rate of sedimentation, strong slope control and instability episodes in the basin, typical of those volcanic environments. That substantial information was the key to understand the features and evolution of the Arenigian basin in the north of the Famatina System.  相似文献   

17.
Glaciers are retreating and thinning in the high altitude of the Himalayas due to global warming, causing into formation of numerous glacial lakes. It is necessary to monitor these glacial lakes consistently to save properties and lives downstream from probable disastrous glacial lake outburst flood. In this study, image processing software ArcGIS and ERDAS Imagine have been used to analyse multispectral image obtained by Earth resource satellite Landsat for delineating the glacial lakes with the help of image enhancement technique like NDWI. Landsat data since 1972 through 2013 have been used and maximum seven glacial lakes (L1–L7) have been detected and delineated in Dhauliganga catchment, they are situated above 4000 masl. The Glacial Lake L2 (Lat 30°26′45″E and Long 80°23′16″N) is the largest whose surface area was 132,300 m2 in Sept 2009, and L6 (Lat 30°23′27″E and Long 80°31′52″N) is highly unstable with variation rate ?55 to +145 % with increasing trend. Additionally, glacial lakes L2 (Lat 30°26′45″E and Long 80°23′16″N) and L6 (Lat 30°23′27″E and Long 80°31′52″N) have been identified as potentially hazardous. These lakes may probably burst; as a result, huge reserve of water and debris may be released all on a sudden. This may transform into hazardous flash flood in downstream causing loss of lives, as well as the destruction of houses, bridges, fields, forests, hydropower stations, roads, etc. It is to note that Dhauliganga river considered in this study is a tributary of Kaliganga river, and should not be confused with its namesake the Dhauliganga river, which is a tributary of Alaknanda river.  相似文献   

18.
Two new sulfide fields (Yubileinoe, 20°09′ N, and Surprise, 20°45.4′ N) were discovered between 20°01′ and 20°54′ N within the Russian Application Area of the Mid-Atlantic Ridge (MAR). The Yubileinoe field is located at a depth of 2300–2550 m in the near-top area of the first rift ridge, which is a boundary of the western wall of the rift valley. This new field and the Zenith-Victory field, which was previously discovered in the eastern wall, occur symmetrically relative to the rift valley of this MAR segment. The Surprise field at a depth of 2800–2850 m is situated in the eastern wall of the rift valley, on the slope of the volcanic uplift. After the discovery of these inactive sulfide fields, the number of hydrothermal fields within the Russian Application Area reached ten.  相似文献   

19.
Pollen evidence from a 350-cm section of a fen in a moraine belt at Rucañancu (39°33′S, 72°18′W) bears on the controversy regarding interpretation of late-glacial and Holocene climate in midlatitude Chile. Earlier pollen studies, indicating a cooling trend between approximately 11,000 and 10,000 yr B.P., disagreed with observations of glacier fluctuations which show continuous glacier wastage and, by inference, warming after 12,500 yr B.P. and possibly earlier, up until Neoglaciation, beginning after 6850 yr B.P. Fossil beetle assemblage data in this time range support the interpretation of climate made from the observed glacier behavior. At Rucañancu, a pollen assemblage containing upper montane podocarp (Podocarpus andinus) in quantities reaching 34% and dating between 10,440 and 10,000 yr B.P. implies a cold climate with summer temperatures possibly 5–8°C lower than today's. Holocene warming began afterward, later than the glacier and beetle records indicate, and continued until at least 8350 yr B.P., as suggested by the sequence of assemblages dominated by Myrtaceae, by Aextoxicon punctatum, and by Gramineae. A subsequent assemblage of Nothofagus obliqua type implies an increase of moisture until 6960 yr B.P., following which N. dombeyi type, under a cool and humid Neoglacial climate, became dominant.  相似文献   

20.
Beach-cast wrack of marine origin is considered a spatial subsidy to the marine-terrestrial transition zone. We found that the wrack line on sand and gravel beaches of Vancouver Island was frequented by intertidal purple shore crabs,Hemigrapsus nudus (Dana 1851) and densely colonized by detritivorous talitrid amphipods. Amphipods spend the day buried in sand and forage on beach wrack during the night.H. nudus were found in supratidal wrack putches immediately after nightly high tides in field censuses, but spent most of the day and ebb tides either submerged subtidally or hidden underneath intertidal rocks and boulders. In feeding trials, intertidal shore crabs were capable of preying on talitrid amphipods. We considerH. nudus an omnivore feeding on both fresh and decaying macroalgae as well as animal prey. Although living supratidally, amphipods were significantly preferred over intertidal littorine snails by foraging shore crabs. Handling time of amphipods was significantly shorter than for littorine snails. While amphipods had a reduced risk of predation byH. nudus when buried in the sand, foraging undern eath wrack patches did not reduce predation pressure on amphipods by shore crabs. Rates of amphipod consumption by shore crabs were higher at darkness than daylight. In addition to an apparent day-night rhythm, tidal height and time elapsed since previous high tide had a significant influence on shore crab density wrack. We conclude that beach-cast wrack acts as a spatial subsidy by virtue of providing a valuable food source to talitrid amphipods, which are in turn consumed by shore crabs that ride the nightly high tide into supratidal wrack patches to reduce the risk of passing bare sand on theiry way to a feeding habitat rich in valuable prey.  相似文献   

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