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The Lower and Upper Cretaceous deposits from the Brazilian marginal basins present a rich and diversified fauna of ostracodes in continental, marine and mixohaline paleoenvironments. While the Cretaceous ostracodes from the northeastern region have already been the subject of many taxonomic and biostratigraphical studies, data from the southeastern basins are still scarce and restricted to the record of a few species. The present study represents a comprehensive contribution to the systematic knowledge of the Aptian–Santonian ostracodes from the Brazilian southeastern marginal basins. A total of 1045 cutting samples were analyzed from 10 wells drilled offshore in Santos, Campos and Espírito Santo basins. Thirty-eight ostracode taxa were recorded from different depositional environments, including the following six new species: Amphicytherura fragilis sp. nov., Brachycythere multidifferentis sp. nov., Fossocytheridea ballentae sp. nov., Fossocytheridea elegans sp. nov., Nigeroloxoconcha itanhaensis sp. nov. and Paracypris eniotmetos sp. nov.  相似文献   

3.
The North American fossil record of dinosaur eggshells for the Cretaceous is primarily restricted to formations of the middle (Albian–Cenomanian) and uppermost (Campanian–Maastrichtian) stages, with a large gap in the record for intermediate stages. Here we describe a dinosaur eggshell assemblage from a formation that represents an intermediate and poorly fossiliferous stage of the Upper Cretaceous, the Santonian Milk River Formation of southern Alberta, Canada. The Milk River eggshell assemblage contains five eggshell taxa: Continuoolithus, Porituberoolithus, Prismatoolithus, Spheroolithus, and Triprismatoolithus. These ootaxa are most similar to those reported from younger Campanian–Maastrichtian formations of the northern Western Interior than they are to ootaxa reported from older middle Cretaceous formations (i.e., predominantly Macroelongatoolithus). Characteristics of the Milk River ootaxa indicate that they are ascribable to at least one ornithopod and four small theropod species. The taxonomic affinity of the eggshell assemblage is consistent with the dinosaur fauna known based on isolated teeth and fragmentary skeletal remains from the formation, although most ornithischians and large theropods are not represented by eggshell. Relative to the Milk River Formation eggshell, similar oospecies occurring in younger Cretaceous deposits tend to be somewhat thicker, which may reflect an increase in body size of various dinosaur lineages during the Late Cretaceous.  相似文献   

4.
Thirteen new species referable to four genera, of which one is new, from the Cretaceous of Russia and Mongolia are established herein and assigned to the family Pelecinidae. Among the four genera, Protopelecinus gen. nov., including four new species, is referred to the subfamily Pelecininae, while Iscopinus Kozlov, including three new species, Eopelecinus Zhang, Rasnitsyn and Zhang, including five new species, and Scorpiopelecinus laetus sp. nov. are assigned to the subfamily Iscopininae. Of these new taxa, eight, namely Protopelecinus regularis, P. furtivus, Iscopinus simplex, ?I. suspectus, Eopelecinus exquisitus, E. scorpioideus, E. rudis and Scorpiopelecinus laetus, are from the Lower Cretaceous Zaza Formation of Baissa, Transbaikalia, Russia; two, E. minutus and E. fragilis, are from the basal Lower Cretaceous Tsagan-Tsab Formation of Khutel-Khara, Mongolia; two, P. dubius and P. deformis, are from the Lower Cretaceous (Aptian?) of Bon Tsagan, Mongolia; and one, I. separatus, is from the Upper Cretaceous (Cenomanian) Ola Formation of Obeshchayushchiy, Russia. A key to fossil pelecinid wasps is provided and a morphological analysis shows that the Pelecinidae might be paraphyletic with respect to the Proctotrupidae. The Chinese insect fauna from both the Yixian and Laiyang formations is dominated by Eopelecinus and Sinopelecinus whereas the Siberian + Mongolian fauna from the Zaza and Tsagan-Tsab formations is dominated by Eopelecinus and Iscopinus. Hence, Eopelecinus is common to both. The differences between the two faunas are likely to be the result of geographical variation in populations.  相似文献   

5.
A diverse fauna of wasps of the extinct parasitoid family Stigmaphronidae (Ceraphronoidea) are recorded in Early Cretaceous (Lower Albian) amber from Spain. Seven new species in five genera are described and figured based on 51 specimens, representing more material than in all the world’s other amber deposits combined. New species include: Elasmophron mari sp. nov., Libanophron sugaar sp. nov., Hippocoon basajauni sp. nov., Burmaphron jentilak sp. nov., B. sorginak sp. nov., B. iratxoak sp. nov., and Tagsmiphron olentzero sp. nov. The significance of the fauna is discussed and compared with that of other Cretaceous amber deposits, in particular the tremendous richness of the Spanish fauna is contrasted with the complete absence of stigmaphronids in the slightly younger and nearby French amber. Whether this stark difference represents particularly favorable conditions for these parasitoids, or their hosts, in the Cretaceous Spanish archipelago, or whether it is owing to taphonomic factors is discussed.  相似文献   

6.
The Upper Cretaceous La Cova limestones (southern Pyrenees, Spain) host a rich and diverse larger foraminiferal fauna, which represents the first diversification of K-strategists after the mass extinction at the Cenomanian–Turonian boundary.The stratigraphic distribution of the main taxa of larger foraminifera defines two assemblages. The first assemblage is characterised by the first appearance of lacazinids (Pseudolacazina loeblichi) and meandropsinids (Eofallotia simplex), by the large agglutinated Montsechiana montsechiensis, and by several species of complex rotalids (Rotorbinella campaniola, Iberorotalia reicheli, Orbitokhatina wondersmitti and Calcarinella schaubi). The second assemblage is defined by the appearance of Lacazina pyrenaica, Palandrosina taxyae and Martiguesia cyclamminiformis.A late Coniacian-early Santonian age was so far accepted for the La Cova limestones, based on indirect correlation with deep-water facies bearing planktic foraminifers of the Dicarinella concavata zone. Strontium isotope stratigraphy, based on many samples of pristine biotic calcite of rudists and ostreids, indicates that the La Cova limestones span from the early Coniacian to the early-middle Santonian boundary. The first assemblage of larger foraminifera appears very close to the early-middle Coniacian boundary and reaches its full diversity by the middle Coniacian. The originations defining the second assemblage are dated as earliest Santonian: they represent important bioevents to define the Coniacian-Santonian boundary in the shallow-water facies of the South Pyrenean province.By means of the calibration of strontium isotope stratigraphy to the Geological Time Scale, the larger foraminiferal assemblages of the La Cova limestones can be correlated to the standard biozonal scheme of ammonites, planktonic foraminifers and calcareous nannoplankton. This correlation is a first step toward a larger foraminifera standard biozonation for Upper Cretaceous carbonate platform facies.  相似文献   

7.
A relatively low diversity coral fauna comprising eight zooxanthellate, three azooxanthellate, and one unidentified species is described from a Late Cretaceous rocky shore at Ivö Klack, southern Sweden. All species, except the solitary azooxanthellate Paracyathus? sp., are represented by one or two specimens only, indicating low preservation potential similar to the aragonite-shelled gastropod fauna from the same locality. The fauna comprises one out of two northernmost zooxanthellate forms known and adds important environmental information to the fauna and depositional conditions of the rocky shore at Ivö Klack.  相似文献   

8.
A thin phosphate-granule conglomerate within the Upper Cretaceous (middle Campanian) Rattlesnake Mountain sandstone member of the Aguja Formation preserves a diverse chondrichthyan and osteichthyan fauna. This highly fossiliferous deposit (the ‘Ten Bits Microsite’) yielded about 5000 teeth, spines, and denticles in a small amount of matrix (c. 150 kg). About 30 identifiable species of sharks, rays, and bony fishes are recognized. Two of the three most abundant chondrichthyan species at Ten Bits (Scapanorhynchus texanus and Ischyrhiza mira) are also the most common species in other middle to late Campanian marine vertebrate faunas along the Gulf and Atlantic Coastal Plain. The myliobatiform rays Brachyrhizodus and Rhombodus that occur at Ten Bits also appear to be characteristic of the Gulf and Atlantic Coast, as are lamniform sharks such as Cretalamna and Serratolamna. These taxa are entirely absent or extremely rare in Western Interior Campanian faunas. In contrast, some small orectolobiform sharks (Cantioscyllium, Chiloscyllium, Columbusia) and small rays (Protoplatyrhina) found at Ten Bits are common in shallow water faunas of the Western Interior and Texas Coastal Plain, but rarely reported from the eastern Gulf or Atlantic Coast. The common Western Interior ray Myledaphus bipartitus does not occur at Ten Bits or in any Gulf or Atlantic Coast fauna. Ptychotrygon agujaensis is abundantly represented in the Ten Bits fauna, but unknown in correlative marine faunas. Although Ptychotrygon occurs in all Western Interior, Gulf and Atlantic Coastal Plain faunas, it is represented elsewhere by apparently endemic species at each collection site. The differences between Western Interior, Gulf, and Atlantic Coastal Plain faunas probably reflect latitudinal variation in water temperature or salinity, or different oceanic water circulation patterns between the Western Interior Seaway and the Gulf or Atlantic Coast that restricted the distributions of some marine fish species. The Ten Bits fauna shares typical species with both Western Interior and Gulf and Atlantic Coast faunas, reflecting its position at the border between these provinces; however, the dominant taxa found at Ten Bits are the same as those found on the Gulf and Atlantic Coast, and indicate that western Texas was more closely allied biogeographically with that province than with the Western Interior of North America. One species tentatively identified in the Ten Bits fauna on the basis of a single tooth, Igdabatis cf. I. indicus, is otherwise known only from southern Europe and Asia, although a similar large myliobatid ray also occurs rarely in Texas Coastal Plain faunas. These occurrences indicate that western Texas may have been near the northern limit of the range for some tropical Tethyan marine vertebrate species.  相似文献   

9.
This paper discusses a well-represented fossil record of cartilaginous fishes (Chondrichthyes) from southern South America. The recovered samples allow the recognition of three assemblages with chronostratigraphic and paleogeographic value: i) typical Maastrichtian sharks and rays with affinities to eastern Pacific fauna, including the taxa Ischyrhiza chilensis, Serratolamna serrata, Centrophoroides sp. associated to Carcharias sp., and Dasyatidae indet.; ii) a scarce reworked assemblage of Paleocene–Early Eocene age including the taxa Otodus obliquus and Megascyliorhinus cooperi; iii) a rich assemblage with reworked taxa of Early to Middle Eocene age, together with autochthonous deposited Middle to Late Eocene taxa with close affinities to paleoichthyofaunas recovered from the North Atlantic, represented by Carchariashopei’, Odontaspis winkleri, Carcharoides catticus, Macrorhizodus praecursor, Carcharocles auriculatus, Striatolamia sp., Striatolamia macrota, Hexanchus agassizi, Notorhynchus sp., Myliobatis sp., Abdounia sp., Pristiophorus sp., Squatina sp., cf. Rhizoprionodon sp., Ischyodus sp., and one new species, Jaekelotodus bagualensis sp. nov. The studied samples include for the first time taxa with well established chronostratigraphic resolutions as well as taphonomic information that help clarifying the age of the fossil-bearing units. In addition, they provide relevant information about the evolution of the Magallanes (=Austral) Basin from the Upper Cretaceous to the Paleogene, suggesting a probable connection with the Quiriquina Basin of south-central Chile during the latest Cretaceous. Finally, the studied assemblages indicate a latitudinal pattern of distribution that provides valuable data on the environmental evolution and temperature of southern South America during the Paleogene.  相似文献   

10.
The coal-bearing, alternating marine and non-marine Longzhaogou Group in eastern Heilongjiang, northeastern China, has long been considered as Jurassic, or mainly Jurassic, in age. However, recent studies have demonstrated that the ammonites and dinoflagellate cysts are of Early Cretaceous age. This has now been confirmed by new radiolarian evidence. The radiolarian fauna recovered from the upper Qihulin Formation of the Longzhaogou Group consists of nine poorly preserved species referable to nine genera. Novixitus is a Cretaceous genus, and the specimens of Archaeodictyomitra sp. and Xitus sp. recovered resemble A. vulgaris Pessagno and X. spicularius (Aliev), respectively.  相似文献   

11.
Thirteen decapod crustacean species, eight of which are new, from the mid-Cretaceous (late Albian) limestones of the Koskobilo quarry in northern Spain are described, illustrated, and discussed. They include: Graptocarcinus texanus; Navarrara betsieae gen. nov., sp. nov.; Acareprosopon bouvieri n. comb.; Laeviprosopon hispanicum sp. nov.; L. planum sp. nov.; L. edoi sp. nov.; L. crassum sp. nov.; Viaia robusta; Cretamaja granulata gen. nov., sp. nov.; Koskobilius postangustus gen. nov., sp. nov.; Navarrahomola hispanica; Glytodynomene alsasuensis; and Albenizus minutus gen. nov., sp. nov. Cretamaja and Koskobilius represent the oldest known spider crabs. In total, 36 species are now known from the Koskobilo locality based on 1078 specimens. To discover the magnitude of the diversity of the decapod fauna of Koskobilo, a comprehensive overview of decapod-rich localities and formations from the Cretaceous worldwide was compiled. It appears that Koskobilo is the most diverse decapod fauna from a single locality currently known from the Cretaceous. A rarefaction analysis shows that the maximum number of species is nearly reached. The number of genera, 26, is also unsurpassed for the Cretaceous. Forty-two species are found from localities within the Eguino Formation to which sediments from the Koskobilo quarry are ascribed, which is also unprecedented for a single formation within the Cretaceous. Evidence suggests that the most diverse decapod faunas from the Cretaceous are found in coral-associated limestones. This is consistent with evidence from the Recent, where decapod diversity is high in coral reefs compared to other habitats. This also suggests that the decapod peak diversity in Koskobilo is largely ecological in nature and not caused by a preservational bias. This is one of the most comprehensive studies on Cretaceous decapod diversity so far. Field work in coral-associated strata is expected to yield more decapod-rich faunas.  相似文献   

12.
Upper Cretaceous deposits in Mongolia, Chinese Inner Mongolia, and, more recently, southern China, have yielded individually rich and taxonomically diverser lizard assemblages. Here we describe the remains of a new terrestrial lizard, Asprosaurus bibongriensis gen. et sp. nov., from the Upper Cretaceous of South Korea. It represents the first record of a Mesozoic lizard from the Korean Peninsula and, although incomplete, is exceptional in its very large size. Characters of the mandible support attribution to crown-group Anguimorpha, with the closest similarities being to monstersaurs, the group represented today by the venomous North American Beaded lizard and Gila monster, genus Heloderma. This group is well-represented in the Upper Cretaceous fossil record in of eastern Asia, and the remains of large monstersaurs have been recovered from several dinosaur egg localities, suggesting dietary preferences similar to those of the living genus. The new Korean lizard, recovered from the Boseong Bibong-ri Dinosaur Egg Site, fits the same pattern.  相似文献   

13.
The stratigraphic distribution and taxonomic diversity of belemnites from the Volgian and Ryazanian deposits exposed in the eastern foothills of the North and Subpolar Urals, are studied and analysed. Studies of new collections from the Mauryn’ya and Yatriya rivers have revealed within the Laugeites groenlandicus-basal Surites analogus ammonite zones, the East Siberian Lagonibelus napaensis, Cylindroteuthis knoxvillensis, and Liobelus russiensis belemnite zones (the latter in the beds’ rank), the Lagonibelus gustomesovi and Arctoteuthis porrectiformis Beds, and local beds with Boreioteuthis explorata and Simobelus compactus. Since in the boundary beds of the Volgian and Ryazanian stages a great number of belemnite species known from the Tordenskjoldberget Member, Kong Karls Land, Svalbard, were found, it is inferred that the accumulation of the member started not in the Valanginian as many researchers believe but immediately at the beginning of the Cretaceous. The dynamics of belemnite species diversity in the northwestern margin of the West Siberian marine basin correlates well with climatic events. An increase of species number in the terminal Volgian-beginning of the Ryazanian corresponds to a temperature elevation in the Siberian paleoseas, and the subsequent reduction of species diversity in the second half of the Ryazanian is correlated with a gradual cooling. The peak of belemnite taxonomic diversity falls at the beginning of the Cretaceous when up to 15 species belonging to eight Cylindroteuthididae genera occurred concurrently. The new species Cylindroteuthis ornata sp. nov., Acroteuthis pseudoconoides sp. nov., Pachyteuthis eximia sp. nov., and Simobelus compactus sp. nov. are described.  相似文献   

14.
The mid-Cenomanian fish beds of Nammoura, Mont-Liban district, Lebanon contain a diverse fauna of aquatic and terrestrial vertebrates, a few crustaceans and moderately well-preserved plant remains of which a single species,Sapindopsis anhouryi , was previously described by Dilcher & Basson (1990). We add 11 species of ferns, gymnosperms and angiosperms of which Nammouria cretacea gen. et sp. nov, Nupharanthus cretacea gen. et sp. nov., Sapindopsis libanensis sp. nov. and Nammourophyllum altingioides gen. et sp. nov. are new taxa. The florule differs markedly from both Early Cretaceous and Turonian plant assemblages of the Middle East, thereby representing a distinct stage of the regional floristic evolution. Its phytogeographic affinities are with contemporaneous floras of North America, Central Europe and the Crimea. A combination of such features as xeromorphism, the prevalence of compound leaves, and the presence of deciduous angiosperm components and gymnosperms may indicate climatic conditions similar to those of the present day Mediterranean.  相似文献   

15.
The following new fossil trichopteran insects are described or revised from the Purbeck Limestone Group and Wealden Supergroup of southern England: Pteromixanum inviolatum gen. et sp. nov., P. ruderatum gen. et sp. nov., P. purbeckianum (Handlirsch) gen. et comb. nov. and P. poxwellense gen. et sp. nov. (Necrotauliidae);Purbimodus minor gen. et sp. nov., P. medius gen. et sp. nov., P. rasnitsyni gen. et sp. nov. and P. saxosus gen. et sp. nov. (Vitimotauliidae);Palaeoludus popovi gen. et sp. nov. (Dysoneuridae);Palaeotarsus desertus gen. et sp. nov. (Plectrotarsidae);Eucrunoecia ridicula gen. et sp. nov. (Lepidostomatidae);Palaeocentropus placidus gen. et sp. nov. (Calamoceratidae) and Helicophidae gen. et sp. incertae sedis. The general composition of the fauna is compared with Early Cretaceous faunas of Asia. The early appearance of several living families is noted.  相似文献   

16.
A previously unknown and morphologically distinct pentadactyl mammal track was recovered from the Upper Cretaceous (Maastrichtian) Laramie Formation at the Fossil Trace site, a National Natural Landmark which is the type locality for Schadipes crypticus, the only named mammal track known from North America. The track is different, and larger than S. crypticus, and thus is evidence of a diversity of mammal trackmakers at this site. Although Cretaceous mammal tracks are very rare, preliminary indications are that those currently known are all morphologically distinct and therefore indicative of a global diversity of different trackmakers, as the body fossil record suggests. Lack of well-preserved mammal trackways with morphologically distinct manus and pes footprints hampers efforts to name diagnostic ichnotaxa.  相似文献   

17.
Seven coleoid jaws recovered from Santonian to lower Campanian (Upper Cretaceous) strata in Hokkaido, Japan were taxonomically studied. Based on the comparison with the jaws of modern and fossil coleoids, six of the seven jaw fossils are referred to the following two genera and three species, including one possible new species: Nanaimoteuthis jeletzkyi and N. yokotai of the order Vampyromorpha, and Paleocirroteuthis sp. nov. (?) of the order Cirroctopodida. The other single lower jaw is seemingly similar to those of modern octopods and teuthids with respect to the shape of the inner lamella, but its order-level assignment could not be determined because of its imperfect preservation. N. jeletzkyi has been described in the Upper Cretaceous fore-arc basin deposits in Hokkaido (Yezo Group) and Vancouver Island, Canada (Nanaimo Group), whereas N. yokotai occurs only in the Yezo Group. These findings, complemented by previous reports of coleoid jaws, gladii, and phragmocones from the Yezo and Nanaimo Groups, demonstrate that a highly diversified, non-belemnitid coleoid fauna including large teuthids had already appeared during the post-Albian Late Cretaceous, in the North Pacific region.  相似文献   

18.
A single confidently dated species of cephalopod is so far known in the Tremadocian of the southern Central Andean Basin (NW Argentina and southern Bolivia). This species belongs to the Eothinoceratidae and has a strong affinity mainly with Avalonia. During the Floian, a notable increase in diversity took place, with the appearance of a variety of families represented by several genera, in particular, within the Family Eothinoceratidae. In addition to the previously described species from southern Bolivia, we evaluate the other records of that family from the Central Andean Basin, and propose the following new taxa: Saloceras sikus sp. nov., Saloceras quena sp. nov., Mutveiceras gen. nov., and Mutveiceras cienagaensis sp. nov. We also describe Margaritoceras diploide, Margaritoceras sp., and Mutveiceras sp. From a palaeogeographic perspective, the cephalopod fauna shows affinities mainly with those of England, Wales, and the Montagne Noire (cold water Gondwana and peri‐Gondwana). As with other cephalopod faunas of mid to high palaeolatitudes, eothinoceratids occur along with other cephalopods forming assemblages of low morphological diversity. We interpret the forms described here as demersal with a subvertical poise, but capable of making rapid buoyancy changes, living in a wide spectrum of shallow offshore to shoreface settings. Copyright © 2014 John Wiley & Sons, Ltd.  相似文献   

19.
Upper Hauterivian deposits in the Kurilovo area, Kamenica 1 section, NE of Niš, are described based of abundant and diverse orbitolinids. So far, the interval was assigned to the Barremian–Aptian on the geological map. Such a new age assignment results from the first detailed study carried out on the orbitolinid fauna contained in the Lower Cretaceous (upper Hauterivian) shallow-water limestones of eastern Serbia. The upper Hauterivian is documented on the basis of two key stratigraphic markers, specifically Valserina primitiva and Paleodictyoconus beckerae.In addition to these late Hauterivian index fossils, the studied section bears orbitolinids having a larger stratigraphic distribution: Cribellopsis neoelongata, Cribellopsis thieuloyi?, Montseciella glanensis, Orbitolinopsis debelmasi, Orbitolinopsis cf. debelmasi, Orbitolinopsis sp., Paleodictyoconus cuvillieri, Paleodictyoconus cf. cuvillieri, Paleodictyoconus cf. beckerae, Paleodictyoconus cf. actinostoma, Paleodictyoconus sp., Paracoskinolina? jourdanensis, Paracoskinolina cf. hispanica, Urgonina alpillensis, Valserina sp. The microfossil assemblage includes other foraminifers such as Charentia cuvillieri, Mayncina bulgarica, Nautiloculina cretacea, Pfenderina globosa, Pseudocyclammina cf. lituus, Pseudolituonella gavonensis, Ammobaculites sp., Bolivinopsis sp., abundant trocholinids, various miliolids, other foraminifers and sparse algae which will be presented separately.  相似文献   

20.
The subject of the study was the taxonomy of the ammonite fauna from the Upper Barremian marls and marly limestones of the Donji Milanovac Formation outcropped at the Boljetin Hill (Danubicum Unit). These sediments yielded a rich ammonite fauna which included also representatives of two superfamilies, Desmoceratoidea and Silesitoidea. The Desmoceratoidea include the family Barremitidae to which belong Plesiospitidiscus boljetinensis n. sp., Barremites balkanicus, Montanesiceras breskovskii n. sp., Barremitites strettostoma strettostoma and B. panae, Torcapella serbiensis n. sp., Pseudohaploceras tachthaliae, P. portaeferreae, Melchiorites haugi and Patruliusiceras cf. crenelatum. The Silesitoidea are represented by the family Silesitidae with Silesites trajani and S. seranonis. With the exception of the new, possibly endemic species, Plesiospitidiscus boljetinensis n. sp., Montanesiceras breskovskii n. sp., and Torcapella serbiensis n. sp., these taxa are common in the Tethyan regions. The studied deposits with ammonites belong to the lower part of the Late Barremian, in particular to the upper part of the Toxancyloceras vandenheckii ammonite Zone up to the lower part of the Imerites giraudi Zone.  相似文献   

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